A trace fossil, also ichnofossil, is a geological record of biological activity. Ichnology is the study of such traces, and is the work of ichnologists. Trace fossils may consist of impressions made on or in the substrate by an organism: for example, burrows, borings (bioerosion), urolites, footprints and feeding marks, and root cavities. The term in its broadest sense also includes the remains of other organic material produced by an organism — for example coprolites or chemical markers — or sedimentological structures produced by biological means - for example, stromatolites. Trace fossils contrast with body fossils, which are the fossilized remains of parts of organisms' bodies, usually altered by later chemical activity or mineralization.
Megalodon, meaning "big tooth", is an extinct species of shark that lived approximately 23 to 2.6 million years ago (mya), during the Early Miocene to the end of the Pliocene. It was formerly thought to be a member of the Lamnidae family, making it closely related to the great white shark. However presently there is near unanimous consensus that it belongs to the extinct family Otodontidae, which diverged from the ancestry of the great white shark during the Early Cretaceous. Its genus placement is still debated, authors placing it in either Carcharocles, Megaselachus, Otodus, or Procarcharodon.
Mosasaurs are an extinct group of large marine reptiles containing 38 genera in total. Their first fossil remains were discovered in a limestone quarry at Maastricht on the Meuse in 1764. Mosasaurs probably evolved from an extinct group of aquatic lizards known as aigialosaurs in the Early Cretaceous. During the last 20 million years of the Cretaceous period, with the extinction of the ichthyosaurs and pliosaurs, mosasaurs became the dominant marine predators. They became extinct as a result of the K-Pg event at the end of the Cretaceous period, about 66 million years ago.
The history of paleontology traces the history of the effort to understand the history of life on Earth by studying the fossil record left behind by living organisms. Since it is concerned with understanding living organisms of the past, paleontology can be considered to be a field of biology, but its historical development has been closely tied to geology and the effort to understand the history of Earth itself.
The Paleocene or Palaeocene, the "old recent", is a geological epoch that lasted from about 66 to 56 million years ago. It is the first epoch of the Paleogene Period in the modern Cenozoic Era. As with many geologic periods, the strata that define the epoch's beginning and end are well identified, but the exact ages remain uncertain.
The Smoky Hill Chalk Member of the Niobrara Chalk formation is a Cretaceous conservation Lagerstätte, or fossil rich geological formation, known primarily for its exceptionally well-preserved marine reptiles. The Smoky Hill Chalk Member is the uppermost of the two structural units of the Niobrara Chalk. It is underlain by the Fort Hays Limestone Member; and the Pierre Shale overlies the Smoky Hill Chalk. The Smoky Hill Chalk outcrops in parts of northwest Kansas, its most famous localities for fossils, and in southeastern Nebraska. Large well-known fossils excavated from the Smoky Hill Chalk include marine reptiles such as plesiosaurs, large bony fish such as Xiphactinus, mosasaurs, flying reptiles or pterosaurs, flightless marine birds such as Hesperornis, and turtles. Many of the most well-known specimens of the marine reptiles were collected by dinosaur hunter Charles H. Sternberg and his son George. The son collected a unique fossil of the giant bony fish Xiphactinus audax with the skeleton of another bony fish, Gillicus arcuatus inside the larger one. Another excellent skeleton of Xiphactinus audax was collected by Edward Drinker Cope during the late nineteenth century heyday of American paleontology and its Bone Wars.
Paleontology in Ohio refers to paleontological research occurring within or conducted by people from the U.S. state of Ohio. Ohio is well known for having a great quantity and diversity of fossils preserved in its rocks. The state's fossil record begins early in the Paleozoic era, during the Cambrian period. Ohio was generally covered by seawater from that time on through the rest of the early Paleozoic. Local invertebrates included brachiopods, cephalopods, coral, graptolites, and trilobites. Vertebrates included bony fishes and sharks. The first land plants in the state grew during the Devonian. During the Carboniferous, Ohio became a more terrestrial environment with an increased diversity of plants that formed expansive swampy deltas. Amphibians and reptiles began to inhabit the state at this time, and remained present into the ensuing Permian. A gap in the local rock record spans from this point until the start of the Pleistocene. During the Ice Age, Ohio was home to giant beavers, humans, mammoths, and mastodons. Paleo-Indians collected fossils that were later incorporated into their mounds. Ohio has been the birthplace of many world famous paleontologists, like Charles Schuchert. Many significant fossils curated by museums in Europe and the United States were found in Ohio. Major local fossil discoveries include the 1965 discovery of more than 50,000 Devonian fish fossils in Cuyahoga County. The Ordovician trilobite Isotelus maximus is the Ohio state fossil.
Paleontology in Maryland refers to paleontological research occurring within or conducted by people from the U.S. state of Maryland. The invertebrate fossils of Maryland are similar to those of neighboring Delaware. For most of the early Paleozoic era, Maryland was covered by a shallow sea, although it was above sea level for portions of the Ordovician and Devonian. The ancient marine life of Maryland included brachiopods and bryozoans while horsetails and scale trees grew on land. By the end of the era, the sea had left the state completely. In the early Mesozoic, Pangaea was splitting up. The same geologic forces that divided the supercontinent formed massive lakes. Dinosaur footprints were preserved along their shores. During the Cretaceous, the state was home to dinosaurs. During the early part of the Cenozoic era, the state was alternatingly submerged by sea water or exposed. During the Ice Age, mastodons lived in the state.
Paleontology in Delaware refers to paleontological research occurring within or conducted by people from the U.S. state of Delaware. There are no local rocks of Precambrian, Paleozoic, Triassic, or Jurassic age, so Delaware's fossil record does not begin until the Cretaceous period. As the Early Cretaceous gave way to the Late Cretaceous, Delaware was being gradually submerged by the sea. Local marine life included cephalopods like Belemnitella americana, and marine reptiles. The dwindling local terrestrial environments were home to a variety of plants, dinosaurs, and pterosaurs. Along with New Jersey, Delaware is one of the best sources of Late Cretaceous dinosaur fossils in the eastern United States. Delaware was still mostly covered by sea water through the Cenozoic era. Local marine life included manatees, porpoises, seals, and whales. Delaware was worked over by glaciers during the Ice Age. The Cretaceous belemnite Belemnitella americana is the Delaware state fossil.
Paleontology in South Carolina refers to paleontological research occurring within or conducted by people from the U.S. state of South Carolina. Evidence suggests that at least part of South Carolina was covered by a warm, shallow sea and inhabited by trilobites during the Cambrian period. Other than this, little is known about the earliest prehistory of South Carolina because the Ordovician, Silurian, Devonian, Carboniferous, Permian, Triassic, and Jurassic, are missing from the state's local rock record. The earliest fossils of South Carolina date back to the Cretaceous, when the state was partially covered by seawater. Contemporary fossils include marine invertebrates and the remains of dinosaur carcasses that washed out to sea. On land, a wide variety of trees grew. Sea levels rose and fell throughout the ensuing Cenozoic era. Local marine life included invertebrates, fish, sharks, whales. The first scientifically accurate identification of vertebrate fossils in North America occurred in South Carolina. In 1725, African slaves digging in a swamp uncovered mammoth teeth, which they recognized as originating from an elephant-like animal.
Paleontology in Vermont refers to paleontological research occurring within or conducted by people from the U.S. state of Vermont. Fossils are generally uncommon in Vermont. Nevertheless, however, significant finds have been made in the state. Very few fossils are known in Vermont east of the Green Mountains due to the type of rock underlying that area. During the early part of the Paleozoic era, Vermont was covered by a warm, shallow sea that would end up being home to creatures like brachiopods, corals, crinoids, ostracoderms, and trilobites. There are no rocks in the state from the Carboniferous, Permian, Triassic, or Jurassic periods. The few Cretaceous rocks present contain no fossils. The Paleogene and Neogene periods are also absent from the local rock record. During the Ice Age, glaciers scoured the state. At times the state was inundated by seawater, allowing marine mammals to venture in. After the seawater drained away the state was home to mastodons. Local fossils had already attracted scientific attention by the mid-19th century when mastodon remains were found in Rutland County. In 1950 a major Paleozoic invertebrate find occurred. The Pleistocene Beluga whale Delphinapterus leucas is the Vermont state fossil.
Paleontology in Louisiana refers to paleontological research occurring within or conducted by people from the U.S. state of Louisiana. Outcrops of fossil-bearing sediments and sedimentary rocks within Louisiana are quite rare. In part, this is because Louisiana’s semi-humid climate results in the rapid weathering and erosion of any exposures and the growth of thick vegetation that conceal any fossil-bearing strata. In addition, Holocene alluvial sediments left behind by rivers like the Mississippi, Red, and Ouachita, as well as marsh deposits, cover about 55% of Louisiana and deeply bury local fossiliferous strata.
Paleontology in Arkansas refers to paleontological research occurring within or conducted by people from the U.S. state of Arkansas. The fossil record of Arkansas spans from the Ordovician to the Eocene. Nearly all of the state's fossils have come from ancient invertebrate life. During the early Paleozoic, much of Arkansas was covered by seawater. This sea would come to be home to creatures including Archimedes, brachiopods, and conodonts. This sea would begin its withdrawal during the Carboniferous, and by the Permian the entire state was dry land. Terrestrial conditions continued into the Triassic, but during the Jurassic, another sea encroached into the state's southern half. During the Cretaceous the state was still covered by seawater and home to marine invertebrates such as Belemnitella. On land the state was home to long necked sauropod dinosaurs, who left behind footprints and ostrich dinosaurs such as Arkansaurus.
Paleontology in South Dakota refers to paleontological research occurring within or conducted by people from the U.S. state of South Dakota. South Dakota is an excellent source of fossils as finds have been widespread throughout the state. During the early Paleozoic era South Dakota was submerged by a shallow sea that would come to be home to creatures like brachiopods, cephalopods, corals, and ostracoderms. Local sea levels rose and fall during the Carboniferous and the sea left completely during the Permian. During the Triassic, the state became a coastal plain, but by the Jurassic it was under a sea where ammonites lived. Cretaceous South Dakota was also covered by a sea that was home to mosasaurs. The sea remained in place after the start of the Cenozoic before giving way to a terrestrial mammal fauna including the camel Poebrotherium, three-toed horses, rhinoceroses, saber teeth, and titanotheres. During the Ice Age glaciers entered the state, which was home to mammoths and mastodons. Local Native Americans interpreted fossils as the remains of the water monster Unktehi and used bits of Baculites shells in magic rituals to summon buffalo herds. Local fossils came to the attention of formally trained scientists with the Lewis and Clark expedition. The Cretaceous horned dinosaur Triceratops horridus is the South Dakota state fossil.
Paleontology in Texas refers to paleontological research occurring within or conducted by people from the U.S. state of Texas. Author Marian Murray has remarked that "Texas is as big for fossils as it is for everything else." Some of the most important fossil finds in United States history have come from Texas. Fossils can be found throughout most of the state. The fossil record of Texas spans almost the entire geologic column from Precambrian to Pleistocene. Shark teeth are probably the state's most common fossil. During the early Paleozoic era Texas was covered by a sea that would later be home to creatures like brachiopods, cephalopods, graptolites, and trilobites. Little is known about the state's Devonian and early Carboniferous life. However, evidence indicates that during the late Carboniferous the state was home to marine life, land plants and early reptiles. During the Permian, the seas largely shrank away, but nevertheless coral reefs formed in the state. The rest of Texas was a coastal plain inhabited by early relatives of mammals like Dimetrodon and Edaphosaurus. During the Triassic, a great river system formed in the state that was inhabited by crocodile-like phytosaurs. Little is known about Jurassic Texas, but there are fossil aquatic invertebrates of this age like ammonites in the state. During the Early Cretaceous local large sauropods and theropods left a great abundance of footprints. Later in the Cretaceous, the state was covered by the Western Interior Seaway and home to creatures like mosasaurs, plesiosaurs, and few icthyosaurs. Early Cenozoic Texas still contained areas covered in seawater where invertebrates and sharks lived. On land the state would come to be home to creatures like glyptodonts, mammoths, mastodons, saber-toothed cats, giant ground sloths, titanotheres, uintatheres, and dire wolves. Archaeological evidence suggests that local Native Americans knew about local fossils. Formally trained scientists were already investigating the state's fossils by the late 1800s. In 1938, a major dinosaur footprint find occurred near Glen Rose. Pleurocoelus was the Texas state dinosaur from 1997 to 2009, when it was replaced by Paluxysaurus jonesi after the Texan fossils once referred to the former species were reclassified to a new genus.
Paleontology in Colorado refers to paleontological research occurring within or conducted by people from the U.S. state of Colorado. The geologic column of Colorado spans about one third of Earth's history. Fossils can be found almost everywhere in the state but are not evenly distributed among all the ages of the state's rocks. During the early Paleozoic, Colorado was covered by a warm shallow sea that would come to be home to creatures like brachiopods, conodonts, ostracoderms, sharks and trilobites. This sea withdrew from the state between the Silurian and early Devonian leaving a gap in the local rock record. It returned during the Carboniferous. Areas of the state not submerged were richly vegetated and inhabited by amphibians that left behind footprints that would later fossilize. During the Permian, the sea withdrew and alluvial fans and sand dunes spread across the state. Many trace fossils are known from these deposits.
Paleontology in Idaho refers to paleontological research occurring within or conducted by people from the U.S. state of Idaho. The fossil record of Idaho spans much of the geologic column from the Precambrian onward. During the Precambrian, bacteria formed stromatolites while worms left behind trace fossils. The state was mostly covered by a shallow sea during the majority of the Paleozoic era. This sea became home to creatures like brachiopods, corals and trilobites. Idaho continued to be a largely marine environment through the Triassic and Jurassic periods of the Mesozoic era, when brachiopods, bryozoans, corals, ichthyosaurs and sharks inhabited the local waters. The eastern part of the state was dry land during the ensuing Cretaceous period when dinosaurs roamed the area and trees grew which would later form petrified wood.
Paleontology in Utah refers to paleontological research occurring within or conducted by people from the U.S. state of Utah. Utah has a rich fossil record spanning almost all of the geologic column. During the Precambrian, the area of northeastern Utah now occupied by the Uinta Mountains was a shallow sea which was home to simple microorganisms. During the early Paleozoic Utah was still largely covered in seawater. The state's Paleozoic seas would come to be home to creatures like brachiopods, fishes, and trilobites. During the Permian the state came to resemble the Sahara desert and was home to amphibians, early relatives of mammals, and reptiles. During the Triassic about half of the state was covered by a sea home to creatures like the cephalopod Meekoceras, while dinosaurs whose footprints would later fossilize roamed the forests on land. Sand dunes returned during the Early Jurassic. During the Cretaceous the state was covered by the sea for the last time. The sea gave way to a complex of lakes during the Cenozoic era. Later, these lakes dissipated and the state was home to short-faced bears, bison, musk oxen, saber teeth, and giant ground sloths. Local Native Americans devised myths to explain fossils. Formally trained scientists have been aware of local fossils since at least the late 19th century. Major local finds include the bonebeds of Dinosaur National Monument. The Jurassic dinosaur Allosaurus fragilis is the Utah state fossil.
Paleontology in California refers to paleontologist research occurring within or conducted by people from the U.S. state of California. California contains rocks of almost every age from the Precambrian to the Recent. Precambrian fossils are present but rare in California.
