Theraphosoidina | |
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Brachypelma klaasi , a member of the Theraphosidae | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Subphylum: | Chelicerata |
Class: | Arachnida |
Order: | Araneae |
Infraorder: | Mygalomorphae |
Clade: | Crassitarsae |
Clade: | Theraphosoidina Raven, 1985 (as a "micropicoorder") |
Families | |
Theraphosoidina is a clade of avicularioid mygalomorph spiders first proposed by Robert J. Raven in 1985, based on a morphological cladistic analysis. Raven included three families: Theraphosidae, Paratropididae and Barychelidae. [1] Subsequent molecular phylogenetic studies upheld the relationship between the Theraphosidae and Barychelidae, but found that Paratropidae fell outside the clade. [2] [3] [4] [5]
Theraphosoidina was first proposed as a taxon by Robert J. Raven in 1985, based on a morphological cladistic analysis. Raven included three families: Theraphosidae, Paratropididae and Barychelidae. The group was characterized by the similar conformation of the male tibial spur (used in mating), reduced toothing of the paired claws and many labial cuspules. [1]
Subsequent molecular phylogenetic studies upheld the relationship between the Theraphosidae and Barychelidae, but found that Paratropidae fell outside the clade. [2] [3] In 2012, it was suggested that Theraphosoidina should probably include an additional family of spiders related to Homostola . [4] A major 2020 study of the Mygalomorphae established the new family Bemmeridae (raised from the rank of tribe) containing the two genera Homostola and Spiroctenus and included it within Theraphosoidina. [5]
An outline version of the preferred cladogram from a 2020 phylogenetic study of the Mygalomorphae is shown below. (A node with lower support is marked ♦.) Although the precise position of the Theraphosoidina clade varied in some of the individual analyses, the clade itself was well supported. [5]
Avicularioidea |
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According to Opatova et al. (2020), the clade includes the following families, one of which (Bemmeridae) was first described at this rank in their study: [5]
The Mygalomorphae, or mygalomorphs, are an infraorder of spiders, and comprise one of three major groups of living spiders with over 3000 species, found on all continents except Antarctica. Many members are known as trapdoor spiders due to them forming trapdoors over their burrows. Other prominent groups include Australian funnel web spiders and tarantulas, with the latter accounting for around one third of all mygalomorphs.
The Mesothelae are a suborder of spiders that includes a single extant family, Liphistiidae, and a number of extinct families. This suborder is thought to form the sister group to all other living spiders, and to retain ancestral characters, such as a segmented abdomen with spinnerets in the middle and two pairs of book lungs. Members of Liphistiidae are medium to large spiders with eight eyes grouped on a tubercle. They are found only in China, Japan, and southeast Asia. The oldest known Mesothelae spiders are known from the Carboniferous, over 300 million years ago.
Ctenizidae is a small family of mygalomorph spiders that construct burrows with a cork-like trapdoor made of soil, vegetation, and silk. They may be called trapdoor spiders, as are other, similar species, such as those of the families Liphistiidae, Barychelidae, and Cyrtaucheniidae, and some species in the Idiopidae and Nemesiidae. The name comes from the distinctive behavior of the spiders to construct trapdoors, and ambush prey from beneath them.
Spider taxonomy is that part of taxonomy that is concerned with the science of naming, defining and classifying all spiders, members of the Araneae order of the arthropod class Arachnida with more than 48,500 described species. However, there are likely many species that have escaped the human eye to this day, and many specimens stored in collections waiting to be described and classified. It is estimated that only one third to one half of the total number of existing species have been described.
Nemesiidae, also known as funnel-web trapdoor spiders, is a family of mygalomorph spiders first described by Eugène Simon in 1889, and raised to family status in 1985. Before becoming its own family, it was considered part of "Dipluridae".
The Dionycha are a clade of spiders (Araneomorphae:Entelegynae), characterized by the possession of two tarsal claws with tufts of hairs (setae) beside them, which produce strong adhesion, enabling some species to climb glass. The circumscription of the group has varied widely; a 2021 analysis resulted in about 20 families, including Salticidae, Gnaphosidae, and Clubionidae.
The Eresoidea or eresoids are a group of araneomorph spiders that have been treated as a superfamily. As usually circumscribed, the group contains three families: Eresidae, Hersiliidae and Oecobiidae. Studies and reviews based on morphology suggested the monophyly of the group; more recent gene-based studies have found the Eresidae and Oecobiidae to fall into different clades, placing doubt on the acceptability of the taxon. Some researchers have grouped Hersiliidae and Oecobiidae into the separate superfamily Oecobioidea, a conclusion supported in a 2017 study, which does not support Eresoidea.
The Haplogynae or haplogynes are one of the two main groups into which araneomorph spiders have traditionally been divided, the other being the Entelegynae. Morphological phylogenetic studies suggested that the Haplogynae formed a clade; more recent molecular phylogenetic studies refute this, although many of the ecribellate haplogynes do appear to form a clade, Synspermiata.
The Euctenizidae are a family of mygalomorph spiders. They are now considered to be more closely related to Idiopidae.
Trichopelma is a genus of South American and Caribbean tarantulas first described by Eugène Simon in 1888.
Rhianodes is a monotypic genus of Southeast Asian brushed trapdoor spiders containing the single species, Rhianodes atratus. It was first described by Robert Raven in 1985, and was later renamed to "Rhianodes" because the name was already attributed to a genus of weevils in the subfamily Apostasimerini. These spiders have only been found in Malaysia, Singapore, and Philippines.
Homostola is a genus of African mygalomorph spiders in the family Bemmeridae. It was first described by Eugène Louis Simon in 1892. Originally placed with the Ctenizidae, it was transferred to the wafer trapdoor spiders in 1985, and to the Bemmeridae in 2020. It is a senior synonym of Stictogaster and Paromostola.
Avicularioidea is a clade of mygalomorph spiders, one of the two main clades into which mygalomorphs are divided. It has been treated at the rank of superfamily.
Anamidae is a family of Australian mygalomorph spiders. It was first described as a tribe by Simon in 1889, then raised to the subfamily Anaminae of the family Nemesiidae, before being raised to a family level by Opatova et al. in 2020.
Bemmeridae is a family of African and Asian mygalomorph spiders that was first described as the tribe Bemmereae by Eugène Simon in 1903. It was elevated to a subfamily of funnel-web trapdoor spiders (Bemmerinae) in 1985, then to its own family in 2020.
Euagridae is a family of mygalomorph spiders. The group was first described as a tribe in 1979 by Robert Raven, who in 1985 elevated it to a subfamily. In 2020, Optova et al. elevated it further to a family.
Bipectina is a clade of avicularioid mygalomorph spiders first proposed by Pablo A. Goloboff in 1993, based on a morphological cladistic analysis. The clade was marked by a number of morphological features, and in particular by the presence of two rows of teeth on the superior tarsal claws of the legs of both sexes, meaning that the claws were bipectinate. The clade was supported by some subsequent analyses, although not all. A major phylogenetic study in 2020 upheld the monophyly of the clade, which contained 19 of the 25 accepted families of the Avicularioidea.
Domiothelina is a clade of avicularioid mygalomorph spiders first proposed by Robert J. Raven in 1985, based on a morphological cladistic analysis. Raven characterized the clade by a number of shared features, including the domed apical segment of the posterior lateral spinnerets. The clade has been supported to some degree by subsequent molecular analyses, although with a somewhat different composition.
Crassitarsae is a clade of avicularioid mygalomorph spiders first proposed by Robert J. Raven in 1985, based on a morphological cladistic analysis. Raven characterized the clade by a number of shared features, including the presence of some scopulae on the tarsi. The clade has been supported to some degree by subsequent molecular analyses, although with a somewhat different composition.
Nemesioidina is a clade of avicularioid mygalomorph spiders proposed in 2020, based on a molecular phylogenetic analysis.