Trogloxenes or subtroglophiles, also called cave guests, are animal species which periodically live in underground habitats such as caves or at the very entrance, but cannot live exclusively in such habitats. [1] [2] Among many scientists, trogloxenes and subtroglophile have slightly different but closely related meanings, with the former covering species that are occasional visitors to underground habitat and the latter species that live more permanently there, but have to go outside (for example, to find food). [2] Both these are in contrast to troglobites, which strictly live in underground habitats. [2]
Examples of trogloxene/subtroglophile species are bats, rats, raccoons and some opiliones (this last group also has fully troglobitic species). Several extinct trogloxenes are known like cave bears, cave lions, cave leopards, and cave hyenas. Indications trusted by geologists and archaeologists combine to show that these animals lived there in the latter part, at least, of the third interglacial epoch, and on through the fourth and last glacial advance, when, although central Europe was free from an ice cap, an almost Arctic climate prevailed, with much rain. This is what is known as the Upper Paleolithic when humanity in Europe was represented by the Neanderthals. [3] During the Upper Paleolithic, many carnivores gradually adapted by increased fur and resorting far more than previously to the shelter of caves. The cave bear (Ursus spelaeus) was the most habitual in its use of caves, and occupied caves before humans began to do so. [3]
The name Trogloxene comes from Greek, Troglos meaning cave and Xenos guest. [4]
A cave or cavern is a natural void in the ground, specifically a space large enough for a human to enter. Caves often form by the weathering of rock and often extend deep underground. The word cave can refer to smaller openings such as sea caves, rock shelters, and grottos, that extend a relatively short distance into the rock and they are called exogene caves. Caves which extend further underground than the opening is wide are called endogene caves.
Lascaux is a network of caves near the village of Montignac, in the department of Dordogne in southwestern France. Over 600 parietal wall paintings cover the interior walls and ceilings of the cave. The paintings represent primarily large animals, typical local contemporary fauna that correspond with the fossil record of the Upper Paleolithic in the area. They are the combined effort of many generations. With continued debate, the age of the paintings is now usually estimated at around 17,000 years. Because of the outstanding prehistoric art in the cave, Lascaux was inducted into the UNESCO World Heritage List in 1979, as an element of the Prehistoric Sites and Decorated Caves of the Vézère Valley.
The Mesolithic or Middle Stone Age is the Old World archaeological period between the Upper Paleolithic and the Neolithic. The term Epipaleolithic is often used synonymously, especially for outside northern Europe, and for the corresponding period in the Levant and Caucasus. The Mesolithic has different time spans in different parts of Eurasia. It refers to the final period of hunter-gatherer cultures in Europe and the Middle East, between the end of the Last Glacial Maximum and the Neolithic Revolution. In Europe it spans roughly 15,000 to 5,000 BP; in the Middle East roughly 20,000 to 10,000 BP. The term is less used of areas farther east, and not at all beyond Eurasia and North Africa.
The Paleolithic or Palaeolithic, also called the Old Stone Age, is a period in human prehistory that is distinguished by the original development of stone tools, and which represents almost the entire period of human prehistoric technology. It extends from the earliest known use of stone tools by hominins, c. 3.3 million years ago, to the end of the Pleistocene, c. 11,650 cal BP.
Speleology is the scientific study of caves and other karst features, as well as their composition, structure, physical properties, history, ecology, and the processes by which they form (speleogenesis) and change over time (speleomorphology). The term speleology is also sometimes applied to the recreational activity of exploring caves, but this is more properly known as caving, potholing, or spelunking. Speleology and caving are often connected, as the physical skills required for in situ study are the same.
In archaeology, cave paintings are a type of parietal art, found on the wall or ceilings of caves. The term usually implies prehistoric origin. These paintings were often created by Homo sapiens, but also Denisovans and Neanderthals; other species in the same Homo genus. Discussion around prehistoric art is important in understanding the history of the Homo sapiens species and how Homo sapiens have come to have unique abstract thoughts. Some point to these prehistoric paintings as possible examples of creativity, spirituality, and sentimental thinking in prehistoric humans.
The Chauvet-Pont-d'Arc Cave in the Ardèche department of southeastern France is a cave that contains some of the best-preserved figurative cave paintings in the world, as well as other evidence of Upper Paleolithic life. It is located near the commune of Vallon-Pont-d'Arc on a limestone cliff above the former bed of the river Ardèche, in the Gorges de l'Ardèche.
The Upper Paleolithic is the third and last subdivision of the Paleolithic or Old Stone Age. Very broadly, it dates to between 50,000 and 12,000 years ago, according to some theories coinciding with the appearance of behavioral modernity in early modern humans, until the advent of the Neolithic Revolution and agriculture.
Troglofauna are small cave-dwelling animals that have adapted to their dark surroundings. Troglofauna and stygofauna are the two types of subterranean fauna. Both are associated with subterranean environments – troglofauna are associated with caves and spaces above the water table and stygofauna with water. Troglofaunal species include spiders, insects, myriapods and others. Some troglofauna live permanently underground and cannot survive outside the cave environment. Troglofauna adaptations and characteristics include a heightened sense of hearing, touch and smell. Loss of under-used senses is apparent in the lack of pigmentation as well as eyesight in most troglofauna. Troglofauna insects may exhibit a lack of wings and longer appendages.
Risovača Cave, is situated at the very entrance of the town of Aranđelovac in central Serbia around 17 m (56 ft) above the Kubršnica river valley. It is one of the most important archaeological sites of the Paleolithic in Serbia besides the Gradac Cave near Kragujevac. Its discovery confirmed the assumed existence of the Paleolithic culture south of the Sava-Danube line and provided new information on the life of prehistoric humans in Europe.
The group of over 700 sites of prehistoric Rock art of the Iberian Mediterranean Basin, also known as Levantine art, were collectively declared a World Heritage Site by UNESCO in 1998. The sites are in the eastern part of Spain and contain rock art dating to the Upper Paleolithic or Mesolithic periods of the Stone Age. The art consists of small painted figures of humans and animals, which are the most advanced and widespread surviving from this period, certainly in Europe, and arguably in the world, at least in the earlier works. It is notable for the number of places included, the largest concentration of such art in Europe. Its name refers to the Mediterranean Basin; however, while some sites are located near the sea, many of them are inland in Aragon and Castilla–La Mancha; it is also often referred to as Levantine Art.
The art of the Upper Paleolithic represents the oldest form of prehistoric art. Figurative art is present in Europe and Southeast Asia, beginning between about 40,000 to 35,000 years ago. Non-figurative cave paintings, consisting of hand stencils and simple geometric shapes, are somewhat older, at least 40,000 years old, and possibly as old as 64,000 years. This latter estimate is due to a controversial 2018 study based on uranium-thorium dating, which would imply Neanderthal authorship and qualify as art of the Middle Paleolithic.
The prehistory in the Valencian Community refers to the period from the Paleolithic, including the appearance of the first populations, until the appearance of colonizing peoples, in the territory of the Valencian Community.
Cro-Magnons or European early modern humans (EEMH) were the first early modern humans to settle in Europe, migrating from western Asia, continuously occupying the continent possibly from as early as 56,800 years ago. They interacted and interbred with the indigenous Neanderthals of Europe and Western Asia, who went extinct 40,000 to 35,000 years ago. The first wave of modern humans in Europe left no genetic legacy to modern Europeans; however, from 37,000 years ago a second wave succeeded in forming a single founder population, from which all subsequent Cro-Magnons descended and which contributes ancestry to present-day Europeans. Cro-Magnons produced Upper Palaeolithic cultures, the first major one being the Aurignacian, which was succeeded by the Gravettian by 30,000 years ago. The Gravettian split into the Epi-Gravettian in the east and Solutrean in the west, due to major climatic degradation during the Last Glacial Maximum (LGM), peaking 21,000 years ago. As Europe warmed, the Solutrean evolved into the Magdalenian by 20,000 years ago, and these peoples recolonised Europe. The Magdalenian and Epi-Gravettian gave way to Mesolithic cultures as big game animals were dying out and the Last Glacial Period drew to a close.
Subterranean fauna refers to animal species that are adapted to live in an underground environment. Troglofauna and stygofauna are the two types of subterranean fauna. Both are associated with hypogeal habitats – troglofauna is associated with terrestrial subterranean environment, and stygofauna with all kind of subterranean waters.
Biospeleology, also known as cave biology, is a branch of biology dedicated to the study of organisms that live in caves and are collectively referred to as troglofauna.
Shulgan-Tash Cave, also known as Kapova Cave, is a limestone karst cave in the Burzyansky District of Bashkortostan, Russia. It is located in the southern Ural Mountains, approximately 200 km (120 mi) south-east of Ufa. Located on the Belaya River in the natural reserve Shulgan-Tash, the cave is best known for the 16,000 years old Upper Paleolithic rock paintings and drawings. It contains the northernmost known ancient paintings.
Pešturina is a cave in the municipality of Niška Banja in southeast Serbia. It is located southwest of Jelašnica and 20 km (12 mi) southeast of Niš. Artifacts from the Middle and Upper Paleolithic periods were discovered since the archaeological excavations began in 2006. The remains, identified as the Mousterian culture, were dated from 111,000 BP+ 5,000 to 39,000 BP + 3,000, which makes Pešturina one of the latest surviving Neanderthal habitats. The cave has been nicknamed the "Serbian Atapuerca".
