- Skull of Babyrousa celebensis , showing long upward curving canine tusks
- Tusks of a wild boar
- Odobenocetops , an extinct whale with a long single tusk
Tusks are elongated, continuously growing front teeth that protrude well beyond the mouth of certain mammal species. They are most commonly canine teeth, as with narwhals, chevrotains, musk deer, water deer, muntjac, pigs, peccaries, hippopotamuses and walruses, or, in the case of elephants, elongated incisors. Tusks share common features such as extra-oral position, growth pattern, composition and structure, and lack of contribution to ingestion. Tusks are thought to have adapted to the extra-oral environments, like dry or aquatic or arctic. [1] In most tusked species both the males and the females have tusks although the males' are larger. Most mammals with tusks have a pair of them growing out from either side of the mouth. Tusks are generally curved and have a smooth, continuous surface. The male narwhal's straight single helical tusk, which usually grows out from the left of the mouth, is an exception to the typical features of tusks described above. Continuous growth of tusks is enabled by formative tissues in the apical openings of the roots of the teeth. [2] [3]
Other than mammals, dicynodonts are the only known vertebrates to have true tusks. [4]
Tusks have a variety of uses depending on the animal. Social displays of dominance, particularly among males, are common, as is their use in defense against attackers. Elephants use their tusks as digging and boring tools. Walruses use their tusks to grip and haul out on ice. [5] It has been suggested that tusks' structure has evolved to be compatible with extra-oral environments. [1]
Elephant tusks are sexually dimorphic, being on average larger in males than in females, and entirely absent in female Asian elephants. Elephants with large tusks each at least 45 kilograms (99 lb) in weight are known as "tuskers", sometimes also called "big tuskers" or "great tuskers". While tuskers are rare today, it is thought that they were more common in the past, prior to human impact on elephant populations. The two record holders for longest and heaviest recorded African bush elephant tusks are around 3.49 metres (11.5 ft) long measured along the outside curve, and 107 kilograms (236 lb) in weight respectively, while the longest and heaviest Asian elephant tusks are 3.26 metres (10.7 ft) long and 73 kilograms (161 lb) respectively. Even larger tusks are known from some extinct proboscideans, such as species of Stegodon, Palaeoloxodon , and mammoths, with the longest tusk ever recorded being that of a specimen of "Mammut" borsoni from Greece, which measures 5.02 metres (16.5 ft) in length, with an estimated weight of 137 kilograms (302 lb) with some mammoth tusks exceeding 4 metres (13 ft) in length and probably 200 kilograms (440 lb) in weight. [6] The largest walrus tusks can reach lengths of over 95 centimetres (3.12 ft). [7] The longest narwhal tusks reach 3 metres (9.8 ft). [8] The upward curving maxillary tusks of babirusa can reach lengths of over 20 centimetres (7.9 in). [9]
Tusks are used by humans to produce ivory, which is used in artifacts and jewellery, and formerly in other items such as piano keys. Consequently, many tusk-bearing species have been hunted commercially and several are endangered. The ivory trade has been severely restricted by the United Nations Convention on International Trade in Endangered Species of Wild Fauna and Flora (CITES).
Tusked animals in human care may undergo tusk trimming or removal for health and safety concerns. [10] Furthermore, surgical veterinary procedures to remove tusks have been explored to mitigate human-wildlife conflicts. [11]
Elephants are the largest living land animals. Three living species are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant. They are the only surviving members of the family Elephantidae and the order Proboscidea; extinct relatives include mammoths and mastodons. Distinctive features of elephants include a long proboscis called a trunk, tusks, large ear flaps, pillar-like legs, and tough but sensitive grey skin. The trunk is prehensile, bringing food and water to the mouth and grasping objects. Tusks, which are derived from the incisor teeth, serve both as weapons and as tools for moving objects and digging. The large ear flaps assist in maintaining a constant body temperature as well as in communication. African elephants have larger ears and concave backs, whereas Asian elephants have smaller ears and convex or level backs.
Ivory is a hard, white material from the tusks and teeth of animals, that consists mainly of dentine, one of the physical structures of teeth and tusks. The chemical structure of the teeth and tusks of mammals is the same, regardless of the species of origin, but ivory contains structures of mineralised collagen. The trade in certain teeth and tusks other than elephant is well established and widespread; therefore, "ivory" can correctly be used to describe any mammalian teeth or tusks of commercial interest which are large enough to be carved or scrimshawed.
A mammoth is any species of the extinct elephantid genus Mammuthus. They lived from the late Miocene epoch into the Holocene about 4,000 years ago, and various species existed in Africa, Europe, Asia, and North America. Mammoths are distinguished from living elephants by their spirally twisted tusks and in at least some later species, the development of numerous adaptions to living in cold environments, including a thick layer of fur.
Proboscidea is a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. Three species of elephant are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant.
The walrus is a large pinniped marine mammal with discontinuous distribution about the North Pole in the Arctic Ocean and subarctic seas of the Northern Hemisphere. It is the only extant species in the family Odobenidae and genus Odobenus. This species is subdivided into two subspecies: the Atlantic walrus, which lives in the Atlantic Ocean, and the Pacific walrus, which lives in the Pacific Ocean.
Walrus ivory, also known as morse, comes from two modified upper canines of a walrus. The tusks grow throughout life and may, in the Pacific walrus, attain a length of one metre. Walrus teeth are commercially carved and traded; the average walrus tooth has a rounded, irregular peg shape and is approximately 5 cm in length.
Incisors are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight. Opossums have 18, whereas armadillos have none.
The North Sulawesi babirusa is a pig-like animal native to Sulawesi and some nearby islands in Indonesia. It has two pairs of large tusks composed of enlarged canine teeth. The upper canines penetrate the top of the snout, curving back toward the forehead. The North Sulawesi babirusa is threatened from hunting and deforestation.
Mammutidae is an extinct family of proboscideans belonging to Elephantimorpha. It is best known for the mastodons, which inhabited North America from the Late Miocene until their extinction at beginning of the Holocene, around 11,000 years ago. The earliest fossils of the group are known from the Late Oligocene of Africa, around 24 million years ago, and fossils of the group have also been found across Eurasia. The name "mastodon" derives from Greek, μαστός "nipple" and ὀδούς "tooth", referring to their characteristic teeth.
Palaeoloxodon is an extinct genus of elephant. The genus originated in Africa during the Early Pleistocene, and expanded into Eurasia at the beginning of the Middle Pleistocene. The genus contains the largest known species of elephants, over 4 metres (13 ft) tall at the shoulders, including the African Palaeoloxodon recki, the European straight-tusked elephant and the South Asian Palaeoloxodon namadicus. P. namadicus has been suggested to be the largest known land mammal by some authors based on extrapolation from fragmentary remains, though these estimates are highly speculative. In contrast, the genus also contains many species of dwarf elephants that evolved via insular dwarfism on islands in the Mediterranean, some only 1 metre (3.3 ft) in height, making them the smallest elephants known. The genus has a long and complex taxonomic history, and at various times, it has been considered to belong to Loxodonta or Elephas, but today is usually considered a valid and separate genus in its own right.
Stegodon is an extinct genus of proboscidean, related to elephants. It was originally assigned to the family Elephantidae along with modern elephants but is now placed in the extinct family Stegodontidae. Like elephants, Stegodon had teeth with plate-like lophs that are different from those of more primitive proboscideans like gomphotheres and mammutids. Fossils of the genus are known from Africa and across much of Asia, as far southeast as Timor. The oldest fossils of the genus are found in Late Miocene strata in Asia, likely originating from the more archaic Stegolophodon, subsequently migrating into Africa. While the genus became extinct in Africa during the Pliocene, Stegodon persisted in South, Southeast and Eastern Asia into the Late Pleistocene.
African elephants are members of the genus Loxodonta comprising two living elephant species, the African bush elephant and the smaller African forest elephant. Both are social herbivores with grey skin. However, they differ in the size and colour of their tusks as well as the shape and size of their ears and skulls.
The babirusas, also called deer-pigs, are a genus, Babyrousa, in the swine family found in the Indonesian islands of Sulawesi, Togian, Sula and Buru. All members of this genus were considered part of a single species until 2002, the babirusa, B. babyrussa, but following that was split into several species. This scientific name is restricted to the Buru babirusa from Buru and Sula, whereas the best-known species, the North Sulawesi babirusa, is named B. celebensis. The remarkable "prehistoric" appearance of these mammals is largely due to the prominent upwards incurving canine tusks of the males, which pierce the flesh in the snout.
Deinotheriidae is a family of prehistoric elephant-like proboscideans that lived during the Cenozoic era, first appearing in Africa, then spreading across southern Asia (Indo-Pakistan) and Europe. During that time, they changed very little, apart from growing much larger in size; by the late Miocene, they had become the largest land animals of their time. Their most distinctive features were their lack of upper tusks and downward-curving tusks on the lower jaw.
The Columbian mammoth is an extinct species of mammoth that inhabited North America from southern Canada to Costa Rica during the Pleistocene epoch. The Columbian mammoth descended from Eurasian steppe mammoths that colonised North America during the Early Pleistocene around 1.5–1.3 million years ago, and later experienced hybridisation with the woolly mammoth lineage. The Columbian mammoth was among the last mammoth species, and the pygmy mammoths evolved from them on the Channel Islands of California. The closest extant relative of the Columbian and other mammoths is the Asian elephant.
The straight-tusked elephant is an extinct species of elephant that inhabited Europe and Western Asia during the Middle and Late Pleistocene. One of the largest known elephant species, mature fully grown bulls on average had a shoulder height of 4 metres (13 ft) and a weight of 13 tonnes (29,000 lb). Straight-tusked elephants likely lived very similarly to modern elephants, with herds of adult females and juveniles and solitary adult males. The species was primarily associated with temperate and Mediterranean woodland and forest habitats, flourishing during interglacial periods, when its range would extend across Europe as far north as Great Britain and eastwards into Russia. Skeletons found in association with stone tools and wooden spears suggest they were scavenged and hunted by early humans, including Neanderthals. It is the ancestor of most dwarf elephants that inhabited islands in the Mediterranean.
Palaeoloxodon namadicus is an extinct species of prehistoric elephant known from the early Middle to Late Pleistocene of the Indian subcontinent, and possibly also elsewhere in Asia. The species grew larger than any living elephant, and some authors have suggested it to have been the largest known land mammal based on extrapolation from fragmentary remains, though these estimates are speculative.
The woolly mammoth is an extinct species of mammoth that lived from the Middle Pleistocene until its extinction in the Holocene epoch. It was one of the last in a line of mammoth species, beginning with the African Mammuthus subplanifrons in the early Pliocene. The woolly mammoth began to diverge from the steppe mammoth about 800,000 years ago in Siberia. Its closest extant relative is the Asian elephant. The Columbian mammoth lived alongside the woolly mammoth in North America, and DNA studies show that the two hybridised with each other.
Stegotetrabelodon is an extinct genus of primitive elephantid from the Late Miocene to Early Pliocene of Africa, the Arabian Peninsula, and Italy.
"Mammut" borsoni is an extinct species of mammutid proboscidean known from the Late Miocene to Early Pleistocene of Eurasia, spanning from western Europe to China. It is the last known mammutid in Eurasia, and amongst the largest of all proboscideans and largest known land mammals.