Homininae, also called "African hominids" or "African apes", is a subfamily of Hominidae. It includes two tribes, with their extant as well as extinct species: 1) the tribe Hominini ―and 2) the tribe Gorillini (gorillas). Alternatively, the genus Pan is sometimes considered to belong to its own third tribe, Panini. Homininae comprises all hominids that arose after orangutans split from the line of great apes. The Homininae cladogram has three main branches, which lead to gorillas, and to humans and chimpanzees via the tribe Hominini and subtribes Hominina and Panina. There are two living species of Panina and two living species of gorillas, but only one extant human species. Traces of extinct Homo species, including Homo floresiensis have been found with dates as recent as 40,000 years ago. Organisms in this subfamily are described as hominine or hominines.
The bonobo, also historically called the pygmy chimpanzee, is an endangered great ape and one of the two species making up the genus Pan. While bonobos are, today, recognized as a distinct species in their own right, they were initially thought to be a subspecies of Pan troglodytes, due to the physical similarities between the two species. Taxonomically, the members of the chimpanzee/bonobo subtribe Panina—composed entirely by the genus Pan—are collectively termed panins.
Yerkish is an artificial language developed for use by non-human primates. It employs a keyboard whose keys contain lexigrams, symbols corresponding to objects or ideas.
Animal languages are forms of non-human animal communication that show similarities to human language. Animals communicate through a variety of signs, such as sounds or movements. Signing among animals may be considered complex enough to be a form of language if the inventory of signs is large. The signs are relatively arbitrary, and the animals seem to produce them with a degree of volition. In experimental tests, animal communication may also be evidenced through the use of lexigrams by chimpanzees and bonobos.
Kanzi, also known by the lexigram , is a male bonobo who has been the subject of several studies on great ape language. According to Sue Savage-Rumbaugh, a primatologist who has studied the bonobo throughout her life, Kanzi has exhibited advanced linguistic aptitude.
The origin of language, its relationship with human evolution, and its consequences have been subjects of study for centuries. Scholars wishing to study the origins of language must draw inferences from evidence such as the fossil record, archaeological evidence, contemporary language diversity, studies of language acquisition, and comparisons between human language and systems of communication existing among animals. Many argue that the origins of language probably relate closely to the origins of modern human behavior, but there is little agreement about the facts and implications of this connection.
Research into great ape language has involved teaching chimpanzees, bonobos, gorillas and orangutans to communicate with humans and each other using sign language, physical tokens, lexigrams, and imitative human speech. Some primatologists argue that the use of these communication methods indicate primate "language" ability, though this depends on one's definition of language.
Nyota, also known by the lexigram , is a bonobo. Nyota was born at the Language Research Center at Georgia State University. His mother was Panbanisha and his father was P-suke. With Panbanisha's death on November 6, 2012, Nyota became the sole surviving member of his immediate family.
Lana was a female chimpanzee, the first to be used in language research using lexigrams. She was born at the Yerkes National Primate Research Center of Emory University, and the project she was allocated to when 1 year old, the LANguage Analogue project led by Duane Rumbaugh, was named after her with the acronym LANA because the project team felt that her identity was well worth preserving.
The Mind of an Ape is a 1983 book by David Premack and his wife Ann James Premack. The authors argue that it is possible to teach language to (non-human) great apes. They write: "We now know that someone who comprehends speech must know language, even if he or she cannot produce it."
Emily Sue Savage-Rumbaugh is a psychologist and primatologist most known for her work with two bonobos, Kanzi and Panbanisha, investigating their linguistic and cognitive abilities using lexigrams and computer-based keyboards. Originally based at Georgia State University's Language Research Center in Atlanta, Georgia, she worked at the Iowa Primate Learning Sanctuary in Des Moines, Iowa from 2006 until her departure in November 2013. She currently sits on the Board of Directors of Bonobo Hope.
Ape Cognition and Conservation Initiative is a sanctuary and scientific research facility in Des Moines, Iowa. It is dedicated to understanding the origins and future of culture, language, tools and intelligence. The facility was announced in 2002 and received its first ape residents in 2004, and is currently home to a colony of seven bonobos involved in non-invasive interdisciplinary studies of their cognitive and communicative capabilities.
The evolutionary origin of religion and religious behavior is a field of study related to evolutionary psychology, the origin of language and mythology, and cross-cultural comparison of the anthropology of religion. Some subjects of interest include Neolithic religion, evidence for spirituality or cultic behavior in the Upper Paleolithic, and similarities in great ape behavior.
Nyota is a given name. It means star in the African languages Swahili and Lingala.
Panbanisha, also known by the lexigram , was a female bonobo that featured in studies on great ape language by Professor Sue Savage-Rumbaugh. Her name is Swahili for "to cleave together for the purpose of contrast."
Human Ape is a 2008 National Geographic documentary film on the genetic and evolutionary origins of human behavior, and covers the genetic and behavioural similarities and differences between humans and other great apes. The award-winning independent production company Pioneer Productions of London was commissioned by National Geographic Channels International to produce Human Ape.
Panpanzee the chimpanzee, was born at the Language Research Center at Georgia State University in Atlanta, Georgia. She lived the rest of her life at the Great Ape Trust in Iowa. Her half-brother is Kanzi, a famous bonobo. Kanzi learned 348 lexigram symbols and over 3,000 words from the English language over her lifespan.
Nicholas Patrick Toth is an American archaeologist and paleoanthropologist. He is a Professor in the Cognitive Science Program at Indiana University and is a founder and co-director of the Stone Age Institute. Toth's archaeological and experimental research has focused on the stone tool technology of Early Stone Age hominins who produced Oldowan and Acheulean artifacts which have been discovered across Africa, Asia, the Middle East, and Europe. He is best known for his experimental work, with Kathy Schick, including their work with the bonobo Kanzi who they taught to make and use simple stone tools similar to those made by our Early Stone Age ancestors.
Primate archaeology is a field of research established in 2008 that combines research interests and foci from primatology and archaeology. The main aim of primate archaeology is to study behavior of extant and extinct primates and the associated material records. The discipline attempts to move beyond archaeology's anthropocentric perspective by placing the focus on both past and present primate tool use.
, is an American qualitative investigator studying language, culture, and tools in non-human primates. He is best known for his collaboration with Sue Savage-Rumbaugh beginning in 1997 at the Language Research Center of Georgia State University. There he co-reared Nyota 
, a baby bonobo, with Panbanisha 
, Kanzi 
and Savage-Rumbaugh 
. [1] Fields and Savage-Rumbaugh are the only scientists in the world carrying out language research with bonobos. [2] 