| Zoophycos | |
|---|---|
 | |
| Zoophycos trace fossil from the Mississippian of northern Kentucky. Scale bar is 5.0 cm (2 inches). | |
Trace fossil classification  | |
| Ichnogenus: | † Zoophycos Massalongo, 1855 |
Zoophycos is a somewhat cosmopolitan ichnogenus thought to be produced by moving and feeding polychaete worms. [1]
Zoophycos occurs in two forms, one planar, and one which resembles a corkscrew. [2] In the latter helicoidal form, successive turns have larger or smaller radii. A marginal tube surrounds the perimeter of the corkscrew, linked to the vertical shaft that connects the burrow to the surface. Spreiten occur between the marginal tube and the corkscrew axis. The burrows can exceed a metre in vertical and horizontal dimension. [3]
One hypothesis proposes that Zoophycos represents gardening behaviour, in a similar fashion to Palaeodictyon . According to this view, there should be a fractionation of carbon isotopes between the burrow infills and the matrix – but such differentiation was not observed in Quaternary instances from deep-water cores off the Portuguese coast. [3] (Variation in morphology between different zoophycos 'species' means that these may not be representative of all zoophycos instances.) Instead, increased concentration of organic carbon within burrows suggests that the burrows may have been used to cache food resources. [3] Alternatively, the burrows may simply be deposit feeding traces. [4]
Zoophycos gives its name to the Zoophycos ichnofacies, an assemblage of trace fossils which is associated with the shelf break. [5] Nevertheless, it is now appreciated that it has a somewhat more cosmopolitan occurrence that has changed through time. [5] It is typically associated with deep marine muds and sands, often between turbidite beds. It is known from both the fossil record and deep sea sediment cores. [6] However, it can also occur in shallow-marine storm deposits, [7] and indeed in Moroccan deposits it seems only to occur in beds that have some form of storm-supplied sediment input. [8]
It occurs from the early Ordovician until the present. [2] It is restricted to deep waters in the Silurian, but occurs in nearshore settings from the Devonian until the end of the Permian. [2]
The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago (Mya), to the beginning of the Devonian Period, 419.2 Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out.
A trace fossil, also known as an ichnofossil, is a fossil record of biological activity by lifeforms but not the preserved remains of the organism itself. Trace fossils contrast with body fossils, which are the fossilized remains of parts of organisms' bodies, usually altered by later chemical activity or mineralization. The study of such trace fossils is ichnology and is the work of ichnologists.
Bioturbation is defined as the reworking of soils and sediments by animals or plants. It includes burrowing, ingestion, and defecation of sediment grains. Bioturbating activities have a profound effect on the environment and are thought to be a primary driver of biodiversity. The formal study of bioturbation began in the 1800s by Charles Darwin experimenting in his garden. The disruption of aquatic sediments and terrestrial soils through bioturbating activities provides significant ecosystem services. These include the alteration of nutrients in aquatic sediment and overlying water, shelter to other species in the form of burrows in terrestrial and water ecosystems, and soil production on land.
Tempestites are storm deposits that can be recognized throughout the geologic record. They are studied in the scientific disciplines of sedimentary geology and paleotempestology. The deposits derive their meaning from the word tempest, a violent storm. Tempestites are preserved within a multitude of sedimentary environments including delta systems, estuarian systems, coastal environments, deep sea environments, and fresh water lacustrine environments. Tempesites most often form in wave-dominated delta systems and preserve, within the sedimentary record, evidence of events and processes below fair weather wave base and above storm weather wave base. They are commonly characterized by hummocky cross-stratified beds that have an erosive base, and can form under combined flow regimes. This erosive base is often seen in the form of gutter casts.
Trace fossils are classified in various ways for different purposes. Traces can be classified taxonomically, ethologically, and toponomically, that is, according to their relationship to the surrounding sedimentary layers. Except in the rare cases where the original maker of a trace fossil can be identified with confidence, phylogenetic classification of trace fossils is an unreasonable proposition.
Marine sediment, or ocean sediment, or seafloor sediment, are deposits of insoluble particles that have accumulated on the seafloor. These particles either have their origins in soil and rocks and have been transported from the land to the sea, mainly by rivers but also by dust carried by wind and by the flow of glaciers into the sea, or they are biogenic deposits from marine organisms or from chemical precipitation in seawater, as well as from underwater volcanoes and meteorite debris.
Paleodictyon is a trace fossil, usually interpreted to be a burrow, which appears in the geologic marine record beginning in the Precambrian/Early Cambrian and in modern ocean environments. Paleodictyon were first described by Giuseppe Meneghini in 1850. The origin of the trace fossil is enigmatic and numerous candidates have been proposed.
Cruziana is a trace fossil consisting of elongate, bilobed, approximately bilaterally symmetrical burrows, usually preserved along bedding planes, with a sculpture of repeated striations that are mostly oblique to the long dimension. It is found in marine and freshwater sediments. It first appears in upper Fortunian rocks of northern Iran and northern Norway. Cruziana has been extensively studied because it has uses in biostratigraphy, and because the traces can reveal many aspects of their makers' behavior.
The Tumblagooda Sandstone is a geological formation deposited during the Silurian or Ordovician periods, between four and five hundred million years ago, and is now exposed on the west coast of Australia in river and coastal gorges near the tourist town of Kalbarri, Kalbarri National Park and the Murchison River gorge, straddling the boundary of the Carnarvon and Perth basins. Visible trackways are interpreted by some to be the earliest evidence of fully terrestrial animals.
Sedimentary structures include all kinds of features in sediments and sedimentary rocks, formed at the time of deposition.
Skolithos is a common trace fossil ichnogenus that is, or was originally, an approximately vertical cylindrical burrow with a distinct lining. It was produced globally by a variety of organisms, mostly in shallow marine environments, and appears as linear features in sedimentary rocks.
An ichnofacies is an assemblage of trace fossils that provides an indication of the conditions that their formative organisms inhabited.
Rusophycus is a trace fossil ichnogenus allied to Cruziana. Rusophycus is the resting trace, recording the outline of the tracemaker; Cruziana is made when the organism moved. The sculpture of Rusophycus may reveal the approximate number of legs that the tracemaker had, although striations (scratchmarks) from a single leg may overlap or be repeated.
The Burgess Shale of British Columbia is famous for its exceptional preservation of mid-Cambrian organisms. Around 69 other sites have been discovered of a similar age, with soft tissues preserved in a similar, though not identical, fashion. Additional sites with a similar form of preservation are known from the Ediacaran and Ordovician periods.
The "Cambrian substrate revolution" or "Agronomic revolution", evidenced in trace fossils, is a sudden diversification of animal burrowing during the early Cambrian period.
Diplocraterion is an ichnogenus describing vertical U-shaped burrows having a spreite between the two limbs of the U. The spreite of an individual Diplocraterion trace can be either protrusive or retrusive. Some ichnospecies have both types. The presence/absence of funnel-shaped openings should not be used as an ichnotaxobase due to the high probability that the upper portions of the trace may have been eroded away. Observation of the orientation of Diplocraterion in the field is frequently used to determine the way up of rock strata at outcrop.
Ophiomorpha is an ichnotaxon, usually interpreted as a burrow of an organism living in the near-shore environment. The burrow lining is more or less smooth on the inside, and densely to strongly mammalated or nodose on the outside, due to the packing of nodules for support of the burrow. Branching is irregular but Y-shaped where present. It is often considered part of the Skolithos ichnofacies, where it has occurred since the early Permian, though it has also occurred in deep water settings since the Late Jurassic, such as well-oxygenated turbidites.
Shallow water marine environment refers to the area between the shore and deeper water, such as a reef wall or a shelf break. This environment is characterized by oceanic, geological and biological conditions, as described below. The water in this environment is shallow and clear, allowing the formation of different sedimentary structures, carbonate rocks, coral reefs, and allowing certain organisms to survive and become fossils.
Chondrites is a trace fossil ichnogenus, preserved as small branching burrows of the same diameter that superficially resemble the roots of a plant. The origin of these structures is currently unknown. Chondrites is found in marine sediments from the Cambrian period of the Paleozoic onwards. It is especially common in sediments that were deposited in reduced-oxygen environments.
Nereites is a genus of trace fossil. Modern tracemakers of incipient Nereites include worm-like organisms, horseshoe crabs and hermit crabs. Traditionally, two models have been proposed for Nereites:
