Agelenoidea

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Tegenaria duellica, Agelenidae Tegenaria gigantea.jpg
Tegenaria duellica , Agelenidae

The Agelenoidea or agelenoids are a superfamily or informal group of entelegyne araneomorph spiders. Phylogenetic studies since 2000 have not consistently recovered such a group, with more recent studies rejecting it.

Phylogeny

In 1999, a phylogenetic study found a clade called "agelenoids" consisting of members of the families Agelenidae, Amphinectidae (now included in Desidae) and Desidae. [1] A 2005 study did not confirm this grouping, instead placing these three families plus Dictynidae in a clade called the "fused cribellar clade". [2] The Desidae have also been placed in the Dictynoidea. [3] In 2014, a cladogram produced in a study of dionychan spiders placed members of the families Amaurobiidae and Cycloctenidae in a clade with members of Agelenidae, Amphinectidae (now Desidae) and Desidae, as sister to the rest of the large RTA Clade. (Amaurobiidae, represented by the genera Pimus and Macrobunus , was not monophyletic in this study.) Shading marks families once considered agelenoids. [4]

RTA clade

Amaurobiidae (Pimus)

Stiphidiidae (Stiphidion)

Agelenidae (Neoramia)

Amaurobiidae (Macrobunus)

Cycloctenidae (Toxopsiella, Cycloctenus)

Desidae (Badumna, Calacadia, Metaltella)

"Oval Calamistrum Clade"

A 2017 study also did not support the Agelenoidea, but placed the two families previously included in this group in a more widely defined "marronoid clade", comprising Amaurobiidae, Agelenidae, Cybaeidae, Cycloctenidae, Desidae, Dictynidae, Hahniidae, Stiphidiidae and Toxopidae, with Agelenidae and Desidae quite far apart. [5]

References

  1. Griswold, Charles E.; Coddington, Jonathan A.; Platnick, Norman I. & Forster, Raymond R. (1999), "Towards a Phylogeny of Entelegyne Spiders (Araneae, Araneomorphae, Entelegynae)", Journal of Arachnology, 27 (1): 53–63, JSTOR   3705965
  2. Griswold, Charles E.; Ramirez, Martin J.; Coddington, Jonathan A. & Platnick, Norman I. (2005), "Atlas of phylogenetic data for entelegyne spiders (Araneae: Araneomorphae: Entelegynae) with comments on their phylogeny" (PDF), Proceedings of the California Academy of Sciences, 56 (Supplement II): 1–324, retrieved 2015-10-11
  3. Dunlop, Jason A. & Penney, David (2011), "Order Araneae Clerck, 1757" (PDF), in Zhang, Z.-Q. (ed.), Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness, Zootaxa, Auckland, New Zealand: Magnolia Press, ISBN   978-1-86977-850-7 , retrieved 2015-10-31
  4. Ramírez, Martín J. (2014), The morphology and phylogeny of dionychan spiders (Araneae, Araneomorphae), Bulletin of the American Museum of Natural History, p. 281
  5. Wheeler, Ward C.; Coddington, Jonathan A.; Crowley, Louise M.; Dimitrov, Dimitar; Goloboff, Pablo A.; Griswold, Charles E.; Hormiga, Gustavo; Prendini, Lorenzo; Ramírez, Martín J.; Sierwald, Petra; Almeida-Silva, Lina; Alvarez-Padilla, Fernando; Arnedo, Miquel A.; Benavides Silva, Ligia R.; Benjamin, Suresh P.; Bond, Jason E.; Grismado, Cristian J.; Hasan, Emile; Hedin, Marshal; Izquierdo, Matías A.; Labarque, Facundo M.; Ledford, Joel; Lopardo, Lara; Maddison, Wayne P.; Miller, Jeremy A.; Piacentini, Luis N.; Platnick, Norman I.; Polotow, Daniele; Silva-Dávila, Diana; Scharff, Nikolaj; Szűts, Tamás; Ubick, Darrell; Vink, Cor J.; Wood, Hannah M. & Zhang, Junxia (2017) [published online 2016], "The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling", Cladistics, 33 (6): 574–616, doi: 10.1111/cla.12182