Vorona is a monotypic genus of prehistoric birds. It was described from fossils found in a Maevarano Formation quarry near the village of Berivotra, Mahajanga Province, Madagascar. The age of the fossilised specimen is Late Cretaceous, probably Maastrichtian. V. berivotrensis is known from scattered remains, possibly from a single individual.
Rapetosaurus is a genus of titanosaurian sauropod dinosaur that lived in Madagascar from 70 to 66 million years ago, at the end of the Cretaceous Period. Only one species, Rapetosaurus krausei, has been identified.
Rahonavis is a genus of bird-like theropods from the Late Cretaceous of what is now northwestern Madagascar. It is known from a partial skeleton found by Catherine Forster and colleagues in Maevarano Formation rocks at a quarry near Berivotra, Mahajanga Province. Rahonavis was a small predator, at about 70 centimetres (2.3 ft) long and 0.45-2.27 kg, with the typical Velociraptor-like raised sickle claw on the second toe. It was originally the first African coelurosaur until the Nqwebasaurus was discovered in 2000.
Simosuchus is an extinct genus of notosuchian crocodylomorphs from the Late Cretaceous of Madagascar. It is named for its unusually short skull. Fully grown individuals were about 0.75 metres (2.5 ft) in length. The type species is Simosuchus clarki, found from the Maevarano Formation in Mahajanga Province, although some fossils have been found in India.
Madtsoiidae is an extinct family of mostly Gondwanan snakes with a fossil record extending from early Cenomanian to late Pleistocene strata located in South America, Africa, India, Australia and Southern Europe. Madtsoiidae include very primitive snakes, which like extant boas and pythons would likely dispatch their prey by constriction. Genera include Madtsoia, one of the longest snakes known, at an estimated 10 metres (33 ft), and the Australian Wonambi and Yurlunggur. As a grouping of basal forms the composition and even the validity of Madtsoiidae is in a state of flux as new pertinent finds are described, with more recent evidence suggesting that it is paraphyletic as previously defined.
The Maevarano Formation is a Late Cretaceous sedimentary rock formation found in the Mahajanga Province of northwestern Madagascar. It is most likely Maastrichtian in age, and records a seasonal, semiarid environment with rivers that had greatly varying discharges. Notable animal fossils recovered include the theropod dinosaur Majungasaurus, the early bird Vorona, the paravian Rahonavis, the titanosaurian sauropod Rapetosaurus, and the giant frog Beelzebufo.
Mahajangasuchus is an extinct genus of crocodyliform which had blunt, conical teeth. The type species, M. insignis, lived during the Late Cretaceous; its fossils have been found in the Maevarano Formation in northern Madagascar. It was a fairly large predator, measuring up to 4 metres (13 ft) long.
Beelzebufo ampinga was a particularly large species of prehistoric frog described in 2008. Common names assigned by the popular media include devil frog, devil toad, and the frog from hell.
Madtsoia is an extinct genus of madtsoiid snakes. It is known from the Eocene of Argentina, the Paleocene of Brazil, the Late Cretaceous (Maastrichtian) of India, and the Late Cretaceous (Maastrichtian) of Madagascar. The type species was the largest with an estimated length of 9–10 m (30–33 ft), and the other three species were smaller. A 5.1 m (17 ft) long M. madagascariensis would have weighed 50 kg (110 lb), but an isolated specimen suggests that this species reached 8 m (26 ft) in maximum length.
Pitekunsaurus is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Anacleto Formation of Neuquén, Argentina. It was described by L. Filippi and A. Garrido in 2008. The type species is P. macayai. The generic name is derived from Mapudungun pitekun, meaning "to discover", the epitheton honours the discoverer, oil company explorer Luis Macaya, who found the fossil in April 2004.
The Ankarafantsika Formation is a Late Cretaceous (Cenomanian) geologic formation of the Mahajanga Basin in the Boeny region of Madagascar, Africa. The fine-grained sandstones of the formation were deposited in a fluvial to lacustrine environment.
The Ankazomihaboka Formation is a Coniacian geologic formation in the Mahajanga Basin of northwestern Madagascar. The formation comprises claystones and sandstones deposited in a fluvial to lacustrine environment. The formation is overlain by the Marovoay Beds and overlies basalt.
Kinkonychelys is an extinct genus of side-necked turtle which existed in Madagascar during the Late Cretaceous period. It contains the single species Kinkonychelys rogersi, named in honor of its discoverer, Raymond R. Rogers. The genus and species are based on UA 9748, a nearly complete skull, which represents the first turtle skull described from the pre-Holocene era in Madagascar. A number of isolated skull and jaw bones have also been assigned to K. rogersi. These specimens were found in rocks of the Maastrichtian-age Maevarano Formation in the Mahajanga Basin of northwestern Madagascar. Another specimen, FMNH PR 2446, is speculated to represent another species, currently known as Kinkonychelys sp., but consensus on its distinction from K. rogersi remains unclear.
Menarana is an extinct genus of madtsoiid snake which existed in Madagascar during the Late Cretaceous. The type species is Menarana nosymena. Several vertebrae and rib fragments as well as part of the basicranium have been found from the Maastrichtian-age Maevarano Formation in the Mahajanga Basin.
Kelyophis is an extinct genus of nigerophiid snake which existed in Madagascar during the Late Cretaceous. The type species is Kelyophis hechti. Trunk vertebrae have been found from the Maastrichtian-age Maevarano Formation in the Mahajanga Basin. Kelyophis is similar to other nigerophiids in its small size, long centra with posterior surfaces that deflect slightly downward, tubercular-shaped neural spines that are directed toward the back of the neural arches, and several other features of the vertebrae.
UA 8699 is a fossil mammalian tooth from the Cretaceous of Madagascar. A broken lower molar about 3.5 mm (0.14 in) long, it is from the Maastrichtian of the Maevarano Formation in northwestern Madagascar. Details of its crown morphology indicate that it is a boreosphenidan, a member of the group that includes living marsupials and placental mammals. David W. Krause, who first described the tooth in 2001, interpreted it as a marsupial on the basis of five shared characters, but in 2003 Averianov and others noted that all those are shared by zhelestid placentals and favored a close relationship between UA 8699 and the Spanish zhelestid Lainodon. Krause used the tooth as evidence that marsupials were present on the southern continents (Gondwana) as early as the late Cretaceous and Averianov and colleagues proposed that the tooth represented another example of faunal exchange between Africa and Europe at the time.
Susan E. Evans is British palaeontologist and herpetologist. She is the author or co-author of over 100 peer-reviewed papers and book chapters.
The geology of Madagascar comprises a variety of rocks of Precambrian age which make up the larger part of the east and centre of the island. They are intruded by basalts and rhyolites of Mesozoic to Cenozoic age. In contrast, the western part of the island is formed from sedimentary rocks of Carboniferous to Quaternary age. Archean rocks occur from the northeast portion of the island down to the south in the Ranotsara shear zone. Rocks in the northern portion of Madagascar are greenstone belts, from the Archean or Paleoproterozoic age.
Sahonachelys is an extinct genus of pelomedusoid turtle from the Late Cretaceous (Maastrichtian) Maevarano Formation of Madagascar. The genus contains a single species, Sahonachelys mailakavava.
Madagascar 
