Squalicorax Temporal range: Albian-Maastrichtian, | |
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Squalicorax pristodontus tooth from the late Maastrichtian of Khouribga, Morocco | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Chondrichthyes |
Subclass: | Elasmobranchii |
Subdivision: | Selachimorpha |
Order: | Lamniformes |
Family: | † Anacoracidae |
Genus: | † Squalicorax Whitley, 1939 |
Type species | |
†Squalicorax pristodontus Agassiz, 1835 | |
Species | |
List of species
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Squalicorax, commonly known as the crow shark, is a genus of extinct lamniform shark known to have lived during the Cretaceous period. The genus had a global distribution in the Late Cretaceous epoch. Multiple species within this genus are considered to be wastebasket taxon due to morphological similarities in the teeth.
The name Squalicorax is derived from the Latin squalus for shark and the Greek κόραξ, "korax" for raven.
Squalicorax was a medium-sized shark, typically measuring about 1.8–3 metres (5.9–9.8 ft) long. The largest specimen of S. pristodontus, SDSM 47683, was signigicantly larger, measuring up to 4.8 metres (16 ft) long. [3]
Their bodies were similar to the modern gray reef sharks, but the shape of the teeth is strikingly similar to that of a tiger shark. The teeth are numerous, relatively small, with a curved crown and serrated, up to 2.5 – 3 cm in height. Large numbers of fossil teeth have been found in Europe, North Africa, and North America. [4] Squalicorax is one of three Cretaceous lamniformes to garner serrations along with Pseudocorax and Galeocorax . [4]
Squalicorax was a coastal predator, but also scavenged as evidenced by a Squalicorax tooth found embedded in the metatarsal (foot) bone of a terrestrial hadrosaurid dinosaur that most likely died on land and ended up in the water. [5] Other food sources included turtles, mosasaurs, ichthyodectids, and other bony fishes and sea creatures. Tooth marks from this shark have also been found on the bones of Pteranodon , but whether the shark actively snatched such large pterosaurs out of the air, attacked them as they dove after prey, or was simply scavenging is not known. [6]
The following are the best studied American species for which relatively complete skeletons are described:
The world's largest and most complete semiarticulated fossil of Squalicorax was found in 2014 in stores of the Canadian Fossil Discovery Centre in Morden, Manitoba, in Canada, where it is now displayed. It measures more than 3 m in length. [8]
The Lamniformes are an order of sharks commonly known as mackerel sharks. It includes some of the most familiar species of sharks, such as the great white, as well as more unusual representatives, such as the goblin shark and megamouth shark.
Cow sharks are a shark family, the Hexanchidae, characterized by an additional pair or pairs of gill slits. Its 37 species are placed within the 10 genera: Gladioserratus, Heptranchias, Hexanchus, Notidanodon, Notorynchus, Pachyhexanchus, Paraheptranchias, Pseudonotidanus, Welcommia, and Weltonia.
Cretoxyrhina is an extinct genus of large mackerel shark that lived about 107 to 73 million years ago during the late Albian to late Campanian of the Late Cretaceous. The type species, C. mantelli, is more commonly referred to as the Ginsu shark, first popularized in reference to the Ginsu knife, as its theoretical feeding mechanism is often compared with the "slicing and dicing" when one uses the knife. Cretoxyrhina is traditionally classified as the likely sole member of the family Cretoxyrhinidae but other taxonomic placements have been proposed, such as within the Alopiidae and Lamnidae.
Xiphactinus, colloquially referred to as the X-fish, is an extinct genus of large predatory marine bony fish that lived during the late Albian to the late Maastrichtian. The genus grew up to 5–6 metres (16–20 ft) in length, and superficially resembled a gargantuan, fanged tarpon.
Prognathodon is an extinct genus of marine lizard belonging to the mosasaur family. It is classified as part of the Mosasaurinae subfamily, alongside genera like Mosasaurus and Clidastes. Prognathodon has been recovered from deposits ranging in age from the Campanian to the Maastrichtian in the Middle East, Europe, New Zealand, and North America.
Otodus is an extinct, cosmopolitan genus of mackerel shark which lived from the Paleocene to the Pliocene epoch. The name Otodus comes from Ancient Greek ὠτ- and ὀδούς – thus, "ear-shaped tooth".
Otodus angustidens is a species of prehistoric megatoothed sharks in the genus Otodus, which lived during the Oligocene and Miocene epochs about 33 to 22 million years ago. The largest individuals were about 11–12 metres (36–39 ft) long. This shark is related to another extinct megatoothed shark, Otodus megalodon.
Axelrodichthys is an extinct genus of mawsoniid coelacanth from the Cretaceous of Africa, North and South America, and Europe. Several species are known, the remains of which were discovered in the Lower Cretaceous (Aptian-Albian) of Brazil, North Africa, and possibly Mexico, as well as in the Upper Cretaceous of Morocco (Cenomanian), Madagascar and France. The Axelrodichthys of the Lower Cretaceous frequented both brackish and coastal marine waters while the most recent species lived exclusively in fresh waters. The French specimens are the last known fresh water coelacanths. Most of the species of this genus reached 1 metre to 2 metres in length. Axelrodichthys was named in 1986 by John G. Maisey in honor of the American ichthyologist Herbert R. Axelrod.
Ptychodus is a genus of extinct durophagous (shell-crushing) sharks from the Late Cretaceous. Fossils of Ptychodus teeth are found in many Late Cretaceous marine sediments worldwide. Numerous species have been described. The youngest remains date to around 85 million years ago. A large number of remains have been found in the former Western Interior Seaway. A 2016 publication found that Ptychodus are likely true sharks belonging to Selachimorpha, rather than hybodonts or batoids as previously thought. Their life history coincides with the typical life of many other large sharks: they lived relatively long lives and were slow growing and produced large offspring and small litters.
Toxochelys is an extinct genus of marine turtle from the Late Cretaceous period. It is the most commonly found fossilized turtle species in the Smoky Hill Chalk, in western Kansas.
Anomoeodus is an extinct genus of prehistoric marine ray-finned fish belonging to the family Pycnodontidae. This genus lived during the Late Cretaceous period, ranging from the Albian to Maastrichtian ages, and had a wide geographic distribution, with fossils found in France, Belgium, the Netherlands, Spain, Egypt, Uzbekistan, and the United States. Some studies have recovered it as a wastebasket taxon. The first fossils of Anomoeodus were described by Louis Agassiz in 1833, although they were described under Pycnodus.
Hemipristis serra is an extinct species of weasel shark which existed during the Miocene epoch. It was described by Louis Agassiz in 1843. While today's snaggletooth shark is not very large or dangerous, Hemipristis serra, which lived in the Atlantic Ocean during the Oligocene and Miocene, was considerably larger than its modern-day relative and had much larger teeth. Its total length is estimated to be 6 metres (20 ft). Marks made by the teeth of H. serra are often found on the bones of the dugong Metaxytherium leading some scientists to hypothesize that H. serra specialized in preying on these sirenians. In the Gatun Formation of Panama, H. serra was contemporary with pups of the large lamniform shark Otodus megalodon, and both it and the great hammerhead are theorized to have preyed on the pups of this larger shark due to their presence within the formation.
The Menuha Formation is the name given to an Upper Cretaceous chalk, marly chalk and conglomeratic chalk unit exposed throughout the Makhtesh Ramon region of southern Israel and parts of northern Israel.
Otodus auriculatus is an extinct species of large sharks in the genus Otodus of the family Otodontidae, closely related to the sharks of the genus Otodus, and also closely related to the later species megalodon. The largest individuals were about 9.5 metres (31 ft) long. Its teeth were large, having coarse serrations on the cutting edge, and also with two large cusplets. The teeth can reach up to 130 millimetres (5.1 in), and belonged to a large "megatoothed" shark.
Meristodonoides is an extinct genus of hybodont. The type species is M. rajkovichi, which was originally a species in the genus Hybodus. The species, along with other Hybodus species such as H. butleri and H. montanensis, was reassigned to Meristodonoides by Charlie J. Underwood and Stephen L. Cumbaa in 2010. The species is primarily known from remains from the Cretaceous of North America, spanning from the Aptian/Albian to Maastrichtian, making it one of the last surviving hybodont genera, though records of the genus likely extend back as far as the Late Jurassic, based on an undescribed skeleton from the Tithonian of England, and fragmentary teeth from the Kimmeridgian of Poland, England and Switzerland. Other remains of the genus are known from the Coniacian of England, the Aptian-Albian of France, and the Campanian of European Russia. The morphology of the teeth suggests an adaptation to tearing prey. Fossils from the Western Interior Seaway suggest that it preferred nearshore marine environments, being absent from deeper-water areas, with it likely also being able to tolerate brackish and freshwater conditions.
Cretalamna is a genus of extinct otodontid shark that lived from the latest Early Cretaceous to Eocene epoch. It is considered by many to be the ancestor of the largest sharks to have ever lived, such as Otodus angustidens, Otodus chubutensis, and Otodus megalodon.
Pseudocorax is an extinct genus of mackerel sharks that lived during the Late Cretaceous. It contains six valid species that have been found in Europe, the Middle East, North Africa, and North America. It was formerly assigned to the family Anacoracidae, but is now placed in its own family Pseudocoracidae along with Galeocorax. The former species "P." australis and "P." primulus have been reidentified as species of Echinorhinus and Squalicorax, respectively.
The Kristianstad Basin is a Cretaceous-age structural basin and geological formation in northeastern Skåne, the southernmost province of Sweden. The basin extends from Hanöbukten, a bay in the Baltic Sea, in the east to the town of Hässleholm in the west and ends with the two horsts Linderödsåsen and Nävlingeåsen in the south. The basin's northern boundary is more diffuse and there are several outlying portions of Cretaceous-age sediments. During the Cretaceous, the region was a shallow subtropical to temperate inland sea and archipelago.
Paraisurus is an extinct genus of mackerel sharks that lived during the Cretaceous. It contains four valid species, which have been found in Europe, Asia, North America, and Australia. A fifth species, P. amudarjensis, is now considered a synonym of P. compressus. While this genus is mostly known from isolated teeth, an associated dentition of P. compressus was found in the Weno Formation of Texas. It went extinct around the Albian-Cenomanian boundary, as a supposed Coniacian occurrence of "P. sp." is likely a misidentified pseudoscapanorhynchid.
Ptychocorax is an extinct genus of mackerel sharks that lived during the Late Cretaceous. It contains three valid species that have been found in Europe and Asia. It was originally identified as a hybodontiform, but was later reidentified as an anacoracid. It has also been considered to belong to its own family, Ptychocoracidae. Ptychocorax is characterized by its unique dentition, combining Squalicorax-like, cutting anterior teeth with Ptychodus-like, crushing posterior teeth.