Cissia eous

Cissia eous
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Lepidoptera
Family:	Nymphalidae
Genus:	Cissia
Species:	C. eous
Binomial name
	Cissia eous; (Butler, 1867)
Synonyms
	Paryphthimoides bahneri (Anken, 1994); Paryphthimoides kiliani (Anken, 1999);

Last updated December 27, 2025

Cissia eous, locally known as the brown butterfly (Brazilian Portuguese : borboleta marrom), is a butterfly in the family Nymphalidae described by Arthur Gardiner Butler in 1867.^[1] It is found in the countries of Brazil, Argentina, Uruguay and Paraguay.^[1]

Biology and phenology

In the moist and shady habitats of several vegetal formations, like secondary forests and coastal forest edges in Brazil, you'll frequently find adults of this species.^[1] They're attracted to fermented fruits and other decaying materials.^[2]

Larvae of this species can feed on Poa plants.^[3] Specifically in Poa annua,^[4] Digitaria sanguinalis and Stenotaphrum secundatum.^[2]^[5]

References

1 2 3 Zacca, Thamara; Casagrande, Mirna; Mielke, Olaf.; Huertas, Blanca; Barbosa, Eduardo; Freitas, André; Magaldi, Luiza; Espeland, Marianne; Nakahara, Shinichi; Willmott, Keith (2018-06-29). "Systematics of the butterfly genus Cissia Doubleday, 1848 (Lepidoptera: Nymphalidae: Satyrinae) using an integrative approach". Arthropod Systematics & Phylogeny. 76 (2): 349–376. doi: 10.3897/asp.76.e31933 . ISSN 1864-8312.
1 2 Biezanko, C.D., 1960. Satyridae, Morphidae et Brassolidae da zona missioneira do Rio Grande do Sul. Instituto Agronômico do Sul/Escola de Agronomia'Eliseu Maciel'.
↑ ACKERY, P. R. (February 1988). "Hostplants and classification: a review of nymphalid butterflies" . Biological Journal of the Linnean Society. 33 (2): 95–203. doi:10.1111/j.1095-8312.1988.tb00446.x. ISSN 0024-4066.
↑ Biezanko, Ceslao M. de, Agustín Ruffinelli Rey, and Carlos S. Carbonell Mas. "Lepidoptera del Uruguay: lista anotada de especies." Revista de la Facultad de Agronomía, 1957 (46): 153 p. (1957).
↑ Silva A.G.A., Gonçalves C.R., Galvão D.M., Gonçalves A.J.L., Gomes J., Silva M.N., Simoni L. 1968. Quarto catálogo dos insetos que vivem nas plantas do Brasil, seus parasitos e predadores. Edição ampliada do "3° Catálogo dos insetos que vivem nas plantas do Brasil" de autoria do Prof. A. M. da Costa Lima. Parte II. Insetos, hospedeiros e inimigos naturais. Índice de insetos e 376 índice de plantas. – Ministério da Agricultura, Rio de Janeiro. 622 pp.

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[:0-1] 1 2 3 Zacca, Thamara; Casagrande, Mirna; Mielke, Olaf.; Huertas, Blanca; Barbosa, Eduardo; Freitas, André; Magaldi, Luiza; Espeland, Marianne; Nakahara, Shinichi; Willmott, Keith (2018-06-29). "Systematics of the butterfly genus Cissia Doubleday, 1848 (Lepidoptera: Nymphalidae: Satyrinae) using an integrative approach". Arthropod Systematics & Phylogeny. 76 (2): 349–376. doi: 10.3897/asp.76.e31933 . ISSN 1864-8312.

[:1-2] 1 2 Biezanko, C.D., 1960. Satyridae, Morphidae et Brassolidae da zona missioneira do Rio Grande do Sul. Instituto Agronômico do Sul/Escola de Agronomia'Eliseu Maciel'.

[3] ACKERY, P. R. (February 1988). "Hostplants and classification: a review of nymphalid butterflies" . Biological Journal of the Linnean Society. 33 (2): 95–203. doi:10.1111/j.1095-8312.1988.tb00446.x. ISSN 0024-4066.

[4] Biezanko, Ceslao M. de, Agustín Ruffinelli Rey, and Carlos S. Carbonell Mas. "Lepidoptera del Uruguay: lista anotada de especies." Revista de la Facultad de Agronomía, 1957 (46): 153 p. (1957).

[5] Silva A.G.A., Gonçalves C.R., Galvão D.M., Gonçalves A.J.L., Gomes J., Silva M.N., Simoni L. 1968. Quarto catálogo dos insetos que vivem nas plantas do Brasil, seus parasitos e predadores. Edição ampliada do "3° Catálogo dos insetos que vivem nas plantas do Brasil" de autoria do Prof. A. M. da Costa Lima. Parte II. Insetos, hospedeiros e inimigos naturais. Índice de insetos e 376 índice de plantas. – Ministério da Agricultura, Rio de Janeiro. 622 pp.

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