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The Radiolaria, also called Radiozoa, are protozoa of diameter 0.1–0.2 mm that produce intricate mineral skeletons, typically with a central capsule dividing the cell into the inner and outer portions of endoplasm and ectoplasm. The elaborate mineral skeleton is usually made of silica. They are found as zooplankton throughout the global ocean. As zooplankton, radiolarians are primarily heterotrophic, but many have photosynthetic endosymbionts and are, therefore, considered mixotrophs. The skeletal remains of some types of radiolarians make up a large part of the cover of the ocean floor as siliceous ooze. Due to their rapid change as species and intricate skeletons, radiolarians represent an important diagnostic fossil found from the Cambrian onwards.
The polycystines are a group of radiolarians. They include the vast majority of the fossil radiolaria, as their skeletons are abundant in marine sediments, making them one of the most common groups of microfossils. These skeletons are composed of opaline silica. In some it takes the form of relatively simple spicules, but in others it forms more elaborate lattices, such as concentric spheres with radial spines or sequences of conical chambers. Two of the orders belonging to this group are the radially-symmetrical Spumellaria, dating back to the late Cambrian period, and the bilaterally-symmetrical Nasselaria, whose origin is placed within the lower Devonian.
Sticholonche is a genus of radiolarians with a single species, Sticholonche zanclea, found in open oceans at depths of 99–510 metres. It is generally considered a heliozoan, placed in its own order, called the Taxopodida. However it has also been classified as an unusual radiolarian, and this has gained support from genetic studies, which place it near the Acantharea.
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Siliceous ooze is a type of biogenic pelagic sediment located on the deep ocean floor. Siliceous oozes are the least common of the deep sea sediments, and make up approximately 15% of the ocean floor. Oozes are defined as sediments which contain at least 30% skeletal remains of pelagic microorganisms. Siliceous oozes are largely composed of the silica based skeletons of microscopic marine organisms such as diatoms and radiolarians. Other components of siliceous oozes near continental margins may include terrestrially derived silica particles and sponge spicules. Siliceous oozes are composed of skeletons made from opal silica SiO2·nH2O, as opposed to calcareous oozes, which are made from skeletons of calcium carbonate (CaCO3·nH2O) organisms (i.e. coccolithophores). Silica (Si) is a bioessential element and is efficiently recycled in the marine environment through the silica cycle. Distance from land masses, water depth and ocean fertility are all factors that affect the opal silica content in seawater and the presence of siliceous oozes.
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References
- ↑ Lipman R.K. 1950. Radiolyarii eotsena Kyzyl-Kumov (The radiolarians of the Eocene of Kyzyl-Kumi). Trudy Vsesoyuznogo Nauchno-Issledovatelskogo Geologicheskogo Instituta (VSEGEI) 1: 51-65
- ↑ Filogeneticheske issledovania kak osnova pri rasrabotke zonalnoi shkali borealnogo paleogena po radiolariam. GE Kozlova, Radiolaria in Biostratigraphy. Nauka, 70-81, 1990
