The examples and perspective in this article deal primarily with North America and do not represent a worldwide view of the subject.(April 2023) |
Krummholz ( ‹See Tfd› German : krumm, "crooked, bent, twisted" and Holz, "wood") — also called knieholz ("knee timber") — is a type of stunted, deformed vegetation encountered in the subarctic and subalpine tree line landscapes, shaped by continual exposure to fierce, freezing winds. Under these conditions, trees can only survive where they are sheltered by rock formations or snow cover. As the lower portion of these trees continues to grow, the coverage becomes extremely dense near the ground. [1] In Newfoundland and Labrador, the formation is known as tuckamore. [2] [3] [4] Krummholz trees are also found on beaches such as the Oregon coast, where trees can become much taller than their subalpine cousins.
The labeling of diverse sets of tree species in different ecological contexts may be problematic. The ecological requirements of krummholz trees in the Alps, for example, are different from those in the Rockies. The terms scrub or shrubland may be more appropriate for some communities with krummholz trees. [5]
Krummholz trees can cover nearly all of the area in which they inhabit, with only patches of moss and flowers in between. Frequent fog and cloudy conditions, along with cool weather, create a rather moist microclimate around the shrubs. [6] Krummholz might depend on less acidic soil to survive. This means that they are threatened by acid rain. The thin soils that cover mountaintops have low buffering capacity, that is the capacity to resist changes in acidity. [7] These trees are also endangered by the use of them as timber for fires, and other human activity. [5]
Common trees showing krummholz formation include European spruce, mountain pine, balsam fir, red spruce, black spruce, subalpine fir, subalpine larch, Engelmann spruce, whitebark pine, limber pine, Bristlecone pine, and lodgepole pine. Instances of the krummholz form of black spruce, Picea mariana , are found in the northern Canadian boreal forests. [8] Krummholz-form black spruce and balsam fir are abundant in the alpine transition zone of the White Mountains of Maine and New Hampshire and of the same zone in the Green Mountains of Vermont. [6]
Subalpine fir is the most common associate of spruce in krummholz vegetation. Other associated coniferous species include alpine larch ( Larix lyallii ), whitebark pine ( Pinus albicaulis ), limber pine ( Pinus flexilis ), and western white pine ( Pinus monticola ) in southern British Columbia, and mountain hemlock ( Tsuga mertensiana ) on the eastern slopes of the Coast Range and in the Revelstoke area of British Columbia. Lodgepole pine (Pinus contorta var. contorta) is a minor associate in most of the British Columbia interior, except in dry alpine areas of the southwest Cariboo/Chilcotin district where it is abundant (Pojar 1985). [9] Ericaceous species (Vaccinium scoparium, V. membranaceum, V. caespitosum, Cassiope mertensiana, Phyllodoce empetriformis) are common in the snow accumulation zone around the base of krummholz colonies.[ citation needed ]
In the Alps in Europe, a scrubland of Pinus mugo is described as occupying the area above the tree line. This is formed by variants of the spruce, beech, and rarely the green alder. These European species were first labeled as a "krummholz belt" by scientists. In the Rocky Mountains, several tree species appear in a similar stunted form, such as specific North American variants of spruce, fir, and pine. These formations were sometimes called "elfin-wood" or "wind-timber". However, English-speaking scientists began to refer to these formations as krummholz as well. [5]
A variation of krummholz formation is a flag tree or banner tree. Branches on the windward side are killed or deformed by the almost constant strong winds, giving the tree a characteristic flag-like appearance. Where the lower portion of the tree is protected by snow cover or rocks, only the exposed upper portion may have this appearance. This is a rather common occurrence in red spruce trees of the highest peaks of the central and even the southern Appalachian Mountains, and is most often seen in the windswept high peaks and plateaus of the Allegheny Mountains.[ citation needed ] This formation most notably occurs with high frequency in the Dolly Sods and Roaring Plains West Wilderness areas along the Allegheny Front in eastern West Virginia, typically occurring at elevations of 3,800 feet (1,200 m) and higher.[ citation needed ] Trade winds in tropical regions near the equator can also shape trees in a similar manner. [1] The southernmost reaches of the Magellanic subpolar forests of Chile contain many flag trees also.[ citation needed ]
Pinus albicaulis, known by the common names whitebark pine, white bark pine, white pine, pitch pine, scrub pine, and creeping pine, is a conifer tree native to the mountains of the western United States and Canada, specifically subalpine areas of the Sierra Nevada, Cascade Range, Pacific Coast Ranges, Rocky Mountains, and Ruby Mountains. It shares the common name "creeping pine" with several other plants.
The tree line is the edge of a habitat at which trees are capable of growing and beyond which they are not. It is found at high elevations and high latitudes. Beyond the tree line, trees cannot tolerate the environmental conditions. The tree line is sometimes distinguished from a lower timberline, which is the line below which trees form a forest with a closed canopy.
Pinus flexilis, the limber pine, is a species of pine tree in the family Pinaceae that occurs in the mountains of the Western United States, Mexico, and Canada. It is also called Rocky Mountain white pine.
Pinus contorta, with the common names lodgepole pine and shore pine, and also known as twisted pine, and contorta pine, is a common tree in western North America. It is common near the ocean shore and in dry montane forests to the subalpine, but is rare in lowland rain forests. Like all pines, it is an evergreen conifer.
The Rocky Mountains subalpine zone is the biotic zone immediately below tree line in the Rocky Mountains of North America. In northern New Mexico, the subalpine zone occupies elevations approximately from 9,000 to 12,000 feet ; while in northern Alberta, the subalpine zone extends from 1,350 to 2,300 metres.
The Sierra Nevada subalpine zone refers to a biotic zone below treeline in the Sierra Nevada mountain range of California, United States. This subalpine zone is positioned between the upper montane zone at its lower limit, and tree line at its upper limit.
Spruce-pine-fir (SPF) is a classification of lumber that can be traded on commodities exchanges.
The biogeoclimatic zones of British Columbia are units of a classification system used by the British Columbia Ministry of Forests for the Canadian province's fourteen different broad, climatic ecosystems. The classification system, termed Biogeoclimatic Ecosystem Classification, exists independently of other ecoregion systems, one created by the World Wildlife Fund and the other in use by Environment Canada, which is based on one created by the Commission for Environmental Cooperation (CEC) and also in use by the US Environmental Protection Agency (EPA). The system of biogeoclimatic ecosystem classification was partly created for the purpose of managing forestry resources, but is also in use by the British Columbia Ministry of Environment and Climate Change Strategy and other provincial agencies. A biogeoclimatic zone is defined as "a geographic area having similar patterns of energy flow, vegetation and soils as a result of a broadly homogenous macroclimate."
The Ecology of the North Cascades is heavily influenced by the high elevation and rain shadow effects of the mountain range. The North Cascades is a section of the Cascade Range from the South Fork of the Snoqualmie River in Washington, United States, to the confluence of the Thompson and Fraser Rivers in British Columbia, Canada, where the range is officially called the Cascade Mountains but is usually referred to as the Canadian Cascades. The North Cascades Ecoregion is a Level III ecoregion in the Commission for Environmental Cooperation's classification system.
The Alberta–British Columbia foothills forests are a temperate coniferous forests ecoregion of Western Canada, as defined by the World Wildlife Fund (WWF) categorization system. This ecoregion borders Canada's taiga and contains a mix of subarctic forest and temperate forest species as a result. This makes the region an ecotone region, or a region that acts as a buffer between two other biomes.
The Yukon Interior dry forests is a taiga ecoregion of Canada.
The South Central Rockies forests is a temperate coniferous forest ecoregion of the United States located mainly in Wyoming, Idaho, and Montana. It has a considerably drier climate than the North Central Rockies forest.
The North Central Rockies forests is a temperate coniferous forest ecoregion of Canada and the United States. This region overlaps in large part with the North American inland temperate rainforest and gets more rain on average than the South Central Rockies forests and is notable for containing the only inland populations of many species from the Pacific coast.
Central Europe contains several life zones, depending on location and elevation.
A snowbank fungus is any one of a number of diverse species of fungi that occur adjacent to or within melting snow. They are most commonly found in the mountains of western North America where a deep snowpack accumulates during the winter and slowly melts through the spring and summer, often shaded by coniferous forest. They may be saprotrophic, mycorrhizal, or in the case of Caloscypha fulgens, pathogenic.
Waterton Biosphere Region is a UNESCO Biosphere Reserve encompassing Waterton Lakes National Park in the extreme southwest of the Province of Alberta, Canada. The region includes a section of the east slopes of the Rocky Mountains extending from the Continental Divide to the edge of the Canadian Great Plains to the east, including the Municipality of Pincher Creek and Cardston County. The Glacier Biosphere Reserve and National Park in Montana, USA is located to the south of the area.
The Alps conifer and mixed forests is a temperate coniferous forest ecoregion in central Europe. It extends along the Alps mountains through portions of France, Italy, Switzerland, Germany, Liechtenstein, Austria, and Slovenia. The ecoregion extends from the lower slopes of the Alps to its peaks, which include Mont Blanc, at 4,809 m (15,778 ft) the highest peak in the Alps.