|Norway spruce (Picea abies)|
|Subfamily:|| Piceoideae |
About 35; see text.
A spruce is a tree of the genus Picea // , a genus of about 35 species of coniferous evergreen trees in the family Pinaceae, found in the northern temperate and boreal (taiga) regions of the Earth. Spruces are large trees, from about 20–60 m (about 60–200 ft) tall when mature, and have whorled branches and conical form. They can be distinguished from other members of the pine family by their needles (leaves), which are four-sided and attached singly to small persistent peg-like structures (pulvini or sterigmata ) on the branches, and by their cones (without any protruding bracts), which hang downwards after they are pollinated. The needles are shed when 4–10 years old, leaving the branches rough with the retained pegs. In other similar genera, the branches are fairly smooth.
In botany, a tree is a perennial plant with an elongated stem, or trunk, supporting branches and leaves in most species. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are usable as lumber or plants above a specified height. Trees are not a taxonomic group but include a variety of plant species that have independently evolved a woody trunk and branches as a way to tower above other plants to compete for sunlight. Trees tend to be long-lived, some reaching several thousand years old. In wider definitions, the taller palms, tree ferns, bananas, and bamboos are also trees. Trees have been in existence for 370 million years. It is estimated that there are just over 3 trillion mature trees in the world.
In botany, an evergreen is a plant that has leaves throughout the year that are always green. This is true even if the plant retains its foliage only in warm climates, and contrasts with deciduous plants, which completely lose their foliage during the winter or dry season. There are many different kinds of evergreen plants, both trees and shrubs. Evergreens include:
The Pinaceae are trees or shrubs, including many of the well-known conifers of commercial importance such as cedars, firs, hemlocks, larches, pines and spruces. The family is included in the order Pinales, formerly known as Coniferales. Pinaceae are supported as monophyletic by their protein-type sieve cell plastids, pattern of proembryogeny, and lack of bioflavonoids. They are the largest extant conifer family in species diversity, with between 220 and 250 species in 11 genera, and the second-largest in geographical range, found in most of the Northern Hemisphere, with the majority of the species in temperate climates, but ranging from subarctic to tropical. The family often forms the dominant component of boreal, coastal, and montane forests. One species, Pinus merkusii, grows just south of the equator in Southeast Asia. Major centres of diversity are found in the mountains of southwest China, Mexico, central Japan, and California.
Spruce are used as food plants by the larvae of some Lepidoptera (moth and butterfly) species, such as the eastern spruce budworm. They are also used by the larvae of gall adelgids (Adelges species).
A larva is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle.
Lepidoptera is an order of insects that includes butterflies and moths. About 180,000 species of the Lepidoptera are described, in 126 families and 46 superfamilies, 10 per cent of the total described species of living organisms. It is one of the most widespread and widely recognizable insect orders in the world. The Lepidoptera show many variations of the basic body structure that have evolved to gain advantages in lifestyle and distribution. Recent estimates suggest the order may have more species than earlier thought, and is among the four most speciose orders, along with the Hymenoptera, Diptera, and Coleoptera.
Choristoneura fumiferana, the eastern spruce budworm, is a species of moth of the family Tortricidae. It is also commonly referred to as the spruce budworm. It is one of the most destructive native insects in the northern spruce and fir forests of the eastern United States and Canada. Its range is also the widest of all the budworm species. Eastern spruce budworm populations can experience significant oscillations. During outbreaks, populations grow to extremely high densities, compared to the relatively low levels that occur in between. These outbreaks are highly destructive and can cause both economic and ecological damage. As a result, methods of control are utilized. Several theories exist regarding these cyclical outbreaks: association with balsam fir maturation, catastrophe theory, dispersal from an epicenter, and oscillations synchronized by entrainment.
In the mountains of western Sweden, scientists have found a Norway spruce, nicknamed Old Tjikko, which by reproducing through layering, has reached an age of 9,550 years and is claimed to be the world's oldest known living tree.
Sweden, formal name: the Kingdom of Sweden, is a Scandinavian Nordic country in Northern Europe. It borders Norway to the west and north and Finland to the east, and is connected to Denmark in the southwest by a bridge-tunnel across the Öresund, a strait at the Swedish-Danish border. At 450,295 square kilometres (173,860 sq mi), Sweden is the largest country in Northern Europe, the third-largest country in the European Union and the fifth largest country in Europe by area. Sweden has a total population of 10.2 million of which 2.5 million have a foreign background. It has a low population density of 22 inhabitants per square kilometre (57/sq mi). The highest concentration is in the southern half of the country.
Old Tjikko is a 9,561-year-old Norway Spruce, located on Fulufjället Mountain of Dalarna province in Sweden. Old Tjikko originally gained fame as the "world's oldest tree." Old Tjikko is, however, a clonal tree that has regenerated new trunks, branches and roots over millennia rather than an individual tree of great age. Old Tjikko is recognized as the oldest living Picea abies and the third oldest known clonal tree.
Layering has evolved as a common means of vegetative propagation of numerous species in natural environments. Layering is also utilized by horticulturists to propagate desirable plants.
The word spruce comes from a Middle English adjective spruse which literally meant from Prussia. The adjective comes from an unknown alteration of an Old French form of Prussia - Pruce, which itself comes from New Latin, which adapted it from Old Prussian.
Middle English was a form of the English language, spoken after the Norman conquest (1066) until the late 15th century. English underwent distinct variations and developments following the Old English period. Scholarly opinion varies, but the Oxford English Dictionary specifies the period when Middle English was spoken as being from 1150 to 1500. This stage of the development of the English language roughly followed the High to the Late Middle Ages.
Old French was the language spoken in Northern France from the 8th century to the 14th century. In the 14th century, these dialects came to be collectively known as the langue d'oïl, contrasting with the langue d'oc or Occitan language in the south of France. The mid-14th century is taken as the transitional period to Middle French, the language of the French Renaissance, specifically based on the dialect of the Île-de-France region.
Prussia was a historically prominent German state that originated in 1525 with a duchy centred on the region of Prussia on the southeast coast of the Baltic Sea. It was de facto dissolved by an emergency decree transferring powers of the Prussian government to German Chancellor Franz von Papen in 1932 and de jure by an Allied decree in 1947. For centuries, the House of Hohenzollern ruled Prussia, successfully expanding its size by way of an unusually well-organised and effective army. Prussia, with its capital in Königsberg and from 1701 in Berlin, decisively shaped the history of Germany.
Spruce, spruse (1412), and Sprws (1378) seem to have been generic terms for commodities brought to England by Hanseatic merchants (especially beer, boards, wooden chests and leather), and the tree thus was believed to be particular to Prussia, which for a time was figurative in England as a land of luxuries.
The Hanseatic League was a commercial and defensive confederation of merchant guilds and market towns in Northwestern and Central Europe. Growing from a few North German towns in the late 1100s, the league came to dominate Baltic maritime trade for three centuries along the coasts of Northern Europe. Hansa territories stretched from the Baltic to the North Sea and inland during the Late Middle Ages, and diminished slowly after 1450.
DNA analyseshave shown that traditional classifications based on the morphology of needle and cone are artificial. A recent study found that P. breweriana had a basal position, followed by P. sitchensis , and the other species were further divided into three clades, suggesting that Picea originated in North America. Spruce has been found in the fossil record from the early Cretaceous, 136 million years ago.
Deoxyribonucleic acid is a molecule composed of two chains that coil around each other to form a double helix carrying genetic instructions for the development, functioning, growth and reproduction of all known organisms and many viruses. DNA and ribonucleic acid (RNA) are nucleic acids; alongside proteins, lipids and complex carbohydrates (polysaccharides), nucleic acids are one of the four major types of macromolecules that are essential for all known forms of life.
Morphology is a branch of biology dealing with the study of the form and structure of organisms and their specific structural features.
A cone is an organ on plants in the division Pinophyta (conifers) that contains the reproductive structures. The familiar woody cone is the female cone, which produces seeds. The male cones, which produce pollen, are usually herbaceous and much less conspicuous even at full maturity. The name "cone" derives from the fact that the shape in some species resembles a geometric cone. The individual plates of a cone are known as scales.
Thirty-five named species of spruce exist in the world. The Plant List has 59 accepted spruce names.
Determining that a tree is a spruce is not difficult; evergreen needles that are more or less quadrangled, and especially the pulvinus, give it away. Beyond that, determination can become more difficult. Intensive sampling in the Smithers/Hazelton/Houston area of British Columbia showed Douglas (1975),according to Coates et al. (1994), that cone scale morphology was the feature most useful in differentiating species of spruce; the length, width, length: width ratio, the length of free scale (the distance from the imprint of the seed wing to the tip of the scale), and the percentage free scale (length of free scale as a percentage of the total length of the scale) were most useful in this regard. Daubenmire (1974), after range-wide sampling, had already recognized the importance of the 2 latter characters. Taylor (1959) had noted that the most obvious morphological difference between typical Picea glauca and typical P. engelmannii was the cone scale, and Horton (1956,1959) found that the most useful diagnostic features of the 2 spruces are in the cone; differences occur in the flower, shoot and needle, "but those in the cone are most easily assessed" (Horton 1959). Coupé et al. (1982) recommended that cone scale characters be based on samples taken from the midsection of each of 10 cones from each of 5 trees in the population of interest.
Without cones, morphological differentiation among spruce species and their hybrids is more difficult. Species classification for seeds collected from spruce stands in which introgressive hybridization between white and Sitka spruces (P. sitchensis) may have occurred is important for determining appropriate cultural regimens in the nursery. If, for instance, white spruce grown at container nurseries in southwestern British Columbia are not given an extended photoperiod, leader growth ceases early in the first growing season, and seedlings do not reach the minimum height specifications.But, if an extended photoperiod is provided for Sitka spruce, seedlings become unacceptably tall by the end of the first growing season . Species classification of seedlots collected in areas where hybridization of white and Sitka spruces has been reported has depended on (i) easily measured cone scale characters of seed trees, especially free scale length, (ii) visual judgements of morphological characters, e.g., growth rhythm, shoot and root weight, and needle serration, or (iii) some combination of (i) and (ii) (Yeh and Arnott 1986). Useful to a degree, these classification procedures have important limitations; genetic composition of the seeds produced by a stand is determined by both the seed trees and the pollen parents, and species classification of hybrid seedlots and estimates of their level of introgression on the basis of seed-tree characteristics can be unreliable when hybrid seedlots vary in their introgressiveness in consequence of spatial and temporal variations in contributions from the pollen parent (Yeh and Arnott 1986). Secondly, morphological characters are markedly influenced by ontogenetic and environmental influences, so that to discern spruce hybrid seedlot composition with accuracy, hybrid seedlots must differ substantially in morphology from both parent species. Yeh and Arnott (1986) pointed out the difficulties of estimating accurately the degree of introgression between white and Sitka spruces; introgression may have occurred at low levels, and/or hybrid seed lots may vary in their degree of introgression in consequence of repeated backcrossing with parental species.
Spruce seedlings are most susceptible immediately following germination, and remain highly susceptible through to the following spring. More than half of spruce seedling mortality probably occurs during the first growing season and is also very high during the first winter,when seedlings are subjected to freezing damage, frost heaving and erosion, as well as smothering by litter and snow-pressed vegetation. Seedlings that germinate late in the growing season are particularly vulnerable because they are tiny and have not had time to harden off fully.
Mortality rates generally decrease sharply thereafter, but losses often remain high for some years. "Establishment" is a subjective concept based on the idea that once a seedling has successfully reached a certain size, not much is likely to prevent its further development. Criteria vary, of course, but Noble and Ronco (1978), cm to 10 cm tall, warranted the designation "established", since only unusual factors such as snow mold, fire, trampling, or predation would then impair regeneration success. Eis (1967) suggested that in dry habitats on either mineral soil or litter seedbeds a 3-year-old seedling may be considered established; in moist habitats, seedlings may need 4 or 5 years to become established on mineral soil, possibly longer on litter seedbeds.for instance, considered that seedlings 4 to 5 years old, or 8
Growth remains very slow for several to many years. Three years after shelterwood felling in subalpine Alberta, dominant regeneration averaged 5.5 cm in height in scarified blocks, and 7.3 cm in non-scarified blocks (Day 1970), possibly reflecting diminished fertility with the removal of the A horizon.
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Spruce is useful as a building wood, commonly referred to by several different names including North American timber, SPF (spruce, pine, fir) and whitewood (the collective name for spruce wood). Spruce wood is used for many purposes, ranging from general construction work and crates to highly specialised uses in wooden aircraft. The Wright brothers' first aircraft, the Flyer , was built of spruce.
Because this species has no insect or decay resistance qualities after logging, it is generally recommended for construction purposes as indoor use only (indoor drywall framing, for example). Spruce wood, when left outside cannot be expected to last more than 12–18 months depending on the type of climate it is exposed to.
Spruce is one of the most important woods for paper uses, as it has long wood fibres which bind together to make strong paper. The fibres are thin walled and collapse to thin bands upon drying. Spruces are commonly used in mechanical pulping as they are easily bleached. Together with northern pines, northern spruces are commonly used to make NBSK. Spruces are cultivated over vast areas as pulpwood.
The fresh shoots of many spruces are a natural source of vitamin C.Captain Cook made alcoholic sugar-based spruce beer during his sea voyages in order to prevent scurvy in his crew. The leaves and branches, or the essential oils, can be used to brew spruce beer.
The tips from the needles can be used to make spruce tip syrup [ clarification needed ]. In survival situations spruce needles can be directly ingested or boiled into a tea. This replaces large amounts of vitamin C. Also, water is stored in a spruce's needles, providing an alternative means of hydration [ clarification needed ]. Spruce can be used as a preventive measure for scurvy in an environment where meat is the only prominent food source [ clarification needed ].
Spruce is the standard material used in soundboards for many musical instruments, including guitars, mandolins, cellos, violins, and the soundboard at the heart of a piano and the harp. Wood used for this purpose is referred to as tonewood.
Spruce, along with cedar, is often used for the soundboard/top of an acoustic guitar. The main types of spruce used for this purpose are Sitka, Engelmann, Adirondack and European spruces.
The resin was used in the manufacture of pitch in the past (before the use of petrochemicals); the scientific name Picea derives from Latin picea "pitch pine" (referring to Scots pine),from piceus , an adjective from pix "pitch".
Native Americans in North America use the thin, pliable roots of some species for weaving baskets and for sewing together pieces of birch bark for canoes. See also Kiidk'yaas for an unusual golden Sitka Spruce sacred to the Haida people.
Spruces are popular ornamental trees in horticulture, admired for their evergreen, symmetrical narrow-conic growth habit. For the same reason, some (particularly Picea abies and P. omorika) are also extensively used as Christmas trees, with artificial Christmas trees often being produced in their likenesses.
Spruce branches are also used at Aintree racecourse, Liverpool, to build several of the fences on the Grand National course. It is also used to make sculptures.
The closely related species Sirococcus conigenus and S. piceicola cause shoot blight and seedling mortality of conifers in North America, Europe, and North Africa. °C to 25 °C. Longer periods of wetness favour increasingly severe disease. Twig tips killed during growth the previous year shows a characteristic crook.Twig blight damage to seedlings of white and red spruces in a nursery near Asheville, North Carolina, was reported by Graves (1914). Hosts include white, black, Engelmann, Norway, and red spruces, although they are not the plants most commonly damaged. Sirococcus blight of spruces in nurseries show up randomly in seedlings to which the fungus was transmitted in infested seed. First-year seedlings are often killed, and larger plants may become too deformed for planting. Outbreaks involving < 30% of spruce seedlings in seedbeds have been traced to seed lots in which only 0.1% to 3% of seeds were infested. Seed infestation has in turn been traced to the colonization of spruce cones by S. conigenus in forests of the western interior. Infection develops readily if conidia are deposited on succulent plant parts that remain wet for at least 24 hours at 10
Rhizosphaera infects white spruce, blue spruce (Picea pungens), and Norway spruces throughout Ontario, causing severe defoliation and sometimes killing small, stressed trees. White spruce is intermediately susceptible. Dead needles show rows of black fruiting bodies. Infection usually begins on lower branches. On white spruce, infected needles are usually retained on the tree into the following summer. The fungicide Chlorthalonil is registered for controlling this needle cast (Davis 1997).
A branch and stem canker associated with the fungus Valsa kunzei Fr. var. picea was reported on white and Norway spruces in Ontario (Jorgensen and Cafley 1961)and Quebec (Ouellette and Bard 1962). In Ontario, only trees of low vigour were affected, but in Quebec vigorous trees were also infected.
Small mammals consume conifer seeds, and also eat seedlings. Cage feeding of deer mice (Peromyscus maniculatus) and red-backed vole (Myodes gapperi) showed a daily maximum seed consumption of 2000 white spruce seeds and of 1000 seeds of lodgepole pine, with the 2 species of mice consuming equal amounts of seed, but showing a preference for the pine over the spruce (Wagg 1963).The short-tailed meadow vole (Microtus pennsylvanicus Ord) voraciously ate all available white spruce and lodgepole pine seedlings, pulling them out of the ground and holding them between their front feet until the whole seedling had been consumed. Wagg (1963) attributed damage observed to the bark and cambium at ground level of small white spruce seedlings over several seasons to meadow voles.
Once shed, seeds contribute to the diet of small mammals, e.g., deer mice, red-backed voles, mountain voles (Microtus montanus), and chipmunks (Eutamias minimus). The magnitude of the loss is difficult to determine, and studies with and without seed protection have yielded conflicting results. In western Montana, for example, spruce seedling success was little better on protected than on unprotected seed spots (Schopmeyer and Helmers 1947),but in British Columbia spruce regeneration depended on protection from rodents (Smith 1955).
An important albeit indirect biotic constraint on spruce establishment is the depredation of seed by squirrels. As much as 90% of a cone crop has been harvested by red squirrels (Zasada et al. 1978).Deer mice, voles, chipmunks, and shrews can consume large quantities of seed; 1 mouse can eat 2000 seeds per night (Radvanyi 1970b). Repeated applications of half a million seeds/ha failed to produce the 750 trees/ha sought by Northwest Pulp and Power, Ltd., near Hinton, Alberta (Radvanyi 1972), but no doubt left a lot of well-fed small mammals. Foraging by squirrels for winter buds (Rowe 1952) has not been reported in relation to young plantations, but Wagg (1963) noted that at Hinton AB, red squirrels were observed cutting the lateral and terminal twigs and feeding on the vegetative and flower buds of white spruce.
Red squirrels in Alaska have harvested as much as 90% of a cone crop (Zasada et al. 1978); cm to 10 cm long on white spruce ranging "from sapling to veteran size" to squirrels foraging for winter buds, cone failure having excluded the more usual food source. The damage has not been reported in relation to small trees, outplants or otherwise.their modus operandi is to cut off great numbers of cones with great expedition early in the fall, and then "spend the rest of the fall shelling out the seeds". In Manitoba, Rowe (1952) ascribed widespread severing of branch tips 5
Porcupines (Erethizon dorsatum L.) may damage spruce (Nienstaedt 1957),but prefer red pine (McLeod 1956). Bark-stripping of white spruce by black bear (Euarctos americanus perniger) is locally important in Alaska (Lutz 1951), but the bark of white spruce is not attacked by field mice (Microtus pennsylvanicus Ord) (Littlefield et al. 1946), even in years of heavy infestation.
The eastern spruce budworm (Choristoneura fumiferana) is a major pest of spruce trees in forests throughout Canada and the eastern United States.Two of the main host plants are black spruce and white spruce. Population levels oscillate, sometimes reaching extreme outbreak levels that can cause extreme defoliation of and damage to spruce trees. To reduce destruction, there are multiple methods of control in place, including pesticides.
Horntails, or Wood Wasps, use this tree for egg laying and the larvae will live in the outer inch of the tree under the bark.
Spruce Beetles (Dendroctonus rufipennis) have destroyed swathes of spruce forest in western North America from Alaska to Wyoming.
The nuclear, mitochondrial and chloroplast genomes of British Columbia interior spruce have been sequenced. The large (20 Gbp) nuclear genome and associated gene annotations of interior spruce (genotype PG29) were published in 2013 and 2015 . For the genomes of P. glauca, please refer to https://en.wikipedia.org/wiki/Picea_glauca
The Pinophyta, also known as Coniferophyta or Coniferae, or commonly as conifers, are a division of vascular land plants containing a single extant class, Pinopsida. They are gymnosperms, cone-bearing seed plants. All extant conifers are perennial woody plants with secondary growth. The great majority are trees, though a few are shrubs. Examples include cedars, Douglas firs, cypresses, firs, junipers, kauri, larches, pines, hemlocks, redwoods, spruces, and yews. As of 1998, the division Pinophyta was estimated to contain eight families, 68 genera, and 629 living species.
Pinus albicaulis, known by the common names whitebark pine, white pine, pitch pine, scrub pine, and creeping pine, is a conifer tree native to the mountains of the western United States and Canada, specifically subalpine areas of the Sierra Nevada, Cascade Range, Pacific Coast Ranges, and Rocky Mountains from Wyoming northwards. It shares the common name "creeping pine" with several other plants.
Picea mariana, the black spruce, is a North American species of spruce tree in the pine family. It is widespread across Canada, found in all 10 provinces and all 3 Arctic territories. Its range extends into northern parts of the United States: in Alaska, the Great Lakes region, and the upper Northeast. It is a frequent part of the biome known as taiga or boreal forest.
A nursery is a place where plants are propagated and grown to usable size They include retail nurseries which sell to the general public, wholesale nurseries which sell only to businesses such as other nurseries and to commercial gardeners, and private nurseries which supply the needs of institutions or private estates.
Picea sitchensis, the Sitka spruce, is a large, coniferous, evergreen tree growing to almost 100 m (330 ft) tall, with a trunk diameter at breast height that can exceed 5 m (16 ft). It is by far the largest species of spruce and the fifth-largest conifer in the world ; and the third-tallest conifer species. The Sitka spruce is one of the few species documented to reach 91 m (299 ft) in height. Its name is derived from the community of Sitka in southeast Alaska, where it is prevalent. Its range hugs the western coast of Canada and continues into northernmost California.
Silviculture is the practice of controlling the growth, composition, health, and quality of forests to meet diverse needs and values.
Pinus edulis, the Colorado pinyon, two-needle piñon, pinyon pine, or simply piñon, is a pine in the pinyon pine group whose ancestor was a member of the Madro-Tertiary Geoflora and is native to the United States.
In agriculture and gardening transplanting or replanting is the technique of moving a plant from one location to another. Most often this takes the form of starting a plant from seed in optimal conditions, such as in a greenhouse or protected nursery bed, then replanting it in another, usually outdoor, growing location. This is common in market gardening and truck farming, where setting out or planting out are synonymous with transplanting. In the horticulture of some ornamental plants, transplants are used infrequently and carefully because they carry with them a significant risk of killing the plant.
Picea glauca, the white spruce, is a species of spruce native to the northern temperate and boreal forests in North America. Picea glauca was originally native from central Alaska all through the east, across southern/central Canada to the Avalon Peninsula in Newfoundland. It now has become naturalized southward into the far northern United States border states like Montana, Minnesota, Wisconsin, Michigan, Vermont, New Hampshire and Maine; there is also an isolated population in the Black Hills of South Dakota and Wyoming. It is also known as Canadian spruce, skunk spruce, cat spruce, Black Hills spruce, western white spruce, Alberta white spruce, and Porsild spruce.
Picea engelmannii, with common names Engelmann spruce, white spruce, mountain spruce, or silver spruce, is a species of spruce native to western North America, from central British Columbia and southwest Alberta, southwest to northern California and southeast to Arizona and New Mexico; there are also two isolated populations in northern Mexico. It is mostly a high altitude mountain tree, growing at 900 metres (3,000 ft) – 3,650 metres (11,980 ft) altitude, rarely lower in the northwest of the range; in many areas it reaches the alpine tree line.
The blue spruce, green spruce, white spruce, Colorado spruce, or Colorado blue spruce, with the Latin (scientific) name Picea pungens, is a species of spruce tree. It is native to Canada and the United States, and is found in growing zones 1 through 7. Its natural range extends from northern New Mexico through Colorado and Utah to Wyoming and into Alberta and British Columbia, but it has been widely introduced elsewhere and is used as an ornamental tree in many places far beyond its native range. The blue spruce has blue-green coloured needles and is a coniferous tree.
Picea jezoensis, the dark-bark spruce, Ezo spruce, Yezo spruce, or Jezo spruce, is a large evergreen tree growing to 30–50 m tall and with a trunk diameter of up to 2 m. It is native to northeast Asia, from the mountains of central Japan and the Changbai Mountains on the China-North Korea border, north to eastern Siberia, including the Sikhote-Alin, Kuril Islands, Sakhalin and Kamchatka. It is found in cold but humid temperate rain forests, and nowhere does its range extend more than 400 km from the Pacific Ocean. The specific epithet jezoensis derives from Ezo, an old name for Hokkaido and other islands north of the Japanese island of Honshu, where the species is found.
White spruce is a common name for several species of spruce (Picea) and may refer to:
A seedling is a young plant sporophyte developing out of a plant embryo from a seed. Seedling development starts with germination of the seed. A typical young seedling consists of three main parts: the radicle, the hypocotyl, and the cotyledons. The two classes of flowering plants (angiosperms) are distinguished by their numbers of seed leaves: monocotyledons (monocots) have one blade-shaped cotyledon, whereas dicotyledons (dicots) possess two round cotyledons. Gymnosperms are more varied. For example, pine seedlings have up to eight cotyledons. The seedlings of some flowering plants have no cotyledons at all. These are said to be acotyledons.
Protoboarmia porcelaria, the porcelain gray or dash-lined looper, is a Geometrid species of moth found throughout North America, except in the far north. The species was first described by Achille Guenée in 1857.
The Alberta Mountain forests are a temperate coniferous forests ecoregion of Canada.
Isorhapontin is a stilbenoid. It is the glucoside of isorhapontigenin. It can be found in mycorrhizal and non-mycorrhizal roots of Norway spruces, in the bark of Picea sitchensis or in white spruce.
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