- A few-days-old Scots pine seedling, the seed still protecting the cotyledons
- Seedling
- Dicotyledon plantlet showing roots
- Seedling of Bombax species
- Buckwheat seedling with first true leaf
A seedling is a young sporophyte developing out of a plant embryo from a seed. Seedling development starts with germination of the seed. A typical young seedling consists of three main parts: the radicle (embryonic root), the hypocotyl (embryonic shoot), and the cotyledons (seed leaves). The two classes of flowering plants (angiosperms) are distinguished by their numbers of seed leaves: monocotyledons (monocots) have one blade-shaped cotyledon, whereas dicotyledons (dicots) possess two round cotyledons. Gymnosperms are more varied. For example, pine seedlings have up to eight cotyledons. The seedlings of some flowering plants have no cotyledons at all. These are said to be acotyledons.
The plumule is the part of a seed embryo that develops into the shoot bearing the first true leaves of a plant. In most seeds, for example the sunflower, the plumule is a small conical structure without any leaf structure. Growth of the plumule does not occur until the cotyledons have grown above ground. This is epigeal germination. However, in seeds such as the broad bean, a leaf structure is visible on the plumule in the seed. These seeds develop by the plumule growing up through the soil with the cotyledons remaining below the surface. This is known as hypogeal germination.
Dicot seedlings grown in the light develop short hypocotyls and open cotyledons exposing the epicotyl. This is also referred to as photomorphogenesis. In contrast, seedlings grown in the dark develop long hypocotyls and their cotyledons remain closed around the epicotyl in an apical hook. This is referred to as skotomorphogenesis or etiolation. Etiolated seedlings are yellowish in color as chlorophyll synthesis and chloroplast development depend on light. They will open their cotyledons and turn green when treated with light.
In a natural situation, seedling development starts with skotomorphogenesis while the seedling is growing through the soil and attempting to reach the light as fast as possible. During this phase, the cotyledons are tightly closed and form the apical hook to protect the shoot apical meristem from damage while pushing through the soil. In many plants, the seed coat still covers the cotyledons for extra protection.
Upon breaking the surface and reaching the light, the seedling's developmental program is switched to photomorphogenesis. The cotyledons open upon contact with light (splitting the seed coat open, if still present) and become green, forming the first photosynthetic organs of the young plant. Until this stage, the seedling lives off the energy reserves stored in the seed. The opening of the cotyledons exposes the shoot apical meristem and the plumule consisting of the first true leaves of the young plant.
The seedlings sense light through the light receptors phytochrome (red and far-red light) and cryptochrome (blue light). Mutations in these photo receptors and their signal transduction components lead to seedling development that is at odds with light conditions, for example seedlings that show photomorphogenesis when grown in the dark..
Once the seedling starts to photosynthesize, it is no longer dependent on the seed's energy reserves. The apical meristems start growing and give rise to the root and shoot. The first "true" leaves expand and can often be distinguished from the round cotyledons through their species-dependent distinct shapes. [1] While the plant is growing and developing additional leaves, the cotyledons eventually senesce and fall off. Seedling growth is also affected by mechanical stimulation, such as by wind or other forms of physical contact, through a process called thigmomorphogenesis.
Temperature and light intensity interact as they affect seedling growth; at low light levels about 40 lumens/m2 a day/night temperature regime of 28 °C/13 °C is effective (Brix 1972). [2] A photoperiod shorter than 14 hours causes growth to stop, whereas a photoperiod extended with low light intensities to 16 h or more brings about continuous (free) growth. Little is gained by using more than 16 h of low light intensity once seedlings are in the free growth mode. Long photoperiods using high light intensities from 10,000 to 20,000 lumens/m2 increase dry matter production, and increasing the photoperiod from 15 to 24 hours may double dry matter growth (Pollard and Logan 1976, Carlson 1979). [3] [4]
The effects of carbon dioxide enrichment and nitrogen supply on the growth of white spruce and trembling aspen were investigated by Brown and Higginbotham (1986). [5] Seedlings were grown in controlled environments with ambient or enriched atmospheric CO2 (350 or 750 f1/L, respectively) and with nutrient solutions with high, medium, and low N content (15.5, 1.55, and 0.16 mM). Seedlings were harvested, weighed, and measured at intervals of less than 100 days. N supply strongly affected biomass accumulation, height, and leaf area of both species. In white spruce only, the root weight ratio (RWR) was significantly increased with the low-nitrogen regime. CO2 enrichment for 100 days significantly increased the leaf and total biomass of white spruce seedlings in the high-N regime, RWR of seedlings in the medium-N regime, and root biomass of seedlings in the low-N regime.
First-year seedlings typically have high mortality rates, drought being the principal cause, with roots having been unable to develop enough to maintain contact with soil sufficiently moist to prevent the development of lethal seedling water stress. Somewhat paradoxically, however, Eis (1967a) [6] observed that on both mineral and litter seedbeds, seedling mortality was greater in moist habitats (alluvium and Aralia–Dryopteris) than in dry habitats (Cornus–Moss). He commented that in dry habitats after the first growing season surviving seedlings appeared to have a much better chance of continued survival than those in moist or wet habitats, in which frost heave and competition from lesser vegetation became major factors in later years. The annual mortality documented by Eis (1967a) [6] is instructive.
Seedlings are particularly vulnerable to attack by pests and diseases [7] and can consequently experience high mortality rates. Diseases which are especially damaging to seedlings include damping off. Pests which are especially damaging to seedlings include cutworms, pillbugs, slugs and snails. [8]
Seedlings are generally transplanted, [9] when the first pair of true leaves appear. This is often known as pricking out in the UK. [10] [11] A shade may be provided if the area is arid or hot. A commercially available vitamin hormone concentrate may be used to avoid transplant shock which may contain thiamine hydrochloride, 1-Naphthaleneacetic acid and indole butyric acid.
In botany, a seed is a plant embryo and food reserve enclosed in a protective outer covering called a seed coat (testa). More generally, the term "seed" means anything that can be sown, which may include seed and husk or tuber. Seeds are the product of the ripened ovule, after the embryo sac is fertilized by sperm from pollen, forming a zygote. The embryo within a seed develops from the zygote and grows within the mother plant to a certain size before growth is halted.
A cotyledon is a "seed leaf" - a significant part of the embryo within the seed of a plant, and is formally defined as "the embryonic leaf in seed-bearing plants, one or more of which are the first to appear from a germinating seed." Botanists use the number of cotyledons present as one characteristic to classify the flowering plants (angiosperms): species with one cotyledon are called monocotyledonous ("monocots"); plants with two embryonic leaves are termed dicotyledonous ("dicots").
In botany, the radicle is the first part of a seedling to emerge from the seed during the process of germination. The radicle is the embryonic root of the plant, and grows downward in the soil. Above the radicle is the embryonic stem or hypocotyl, supporting the cotyledon(s).
A spruce is a tree of the genus Picea, a genus of about 40 species of coniferous evergreen trees in the family Pinaceae, found in the northern temperate and boreal (taiga) regions of the Earth. Picea is the sole genus in the subfamily Piceoideae. Spruces are large trees, from about 20 to 60 m tall when mature, and have whorled branches and conical form.
Germination is the process by which an organism grows from a seed or spore. The term is applied to the sprouting of a seedling from a seed of an angiosperm or gymnosperm, the growth of a sporeling from a spore, such as the spores of fungi, ferns, bacteria, and the growth of the pollen tube from the pollen grain of a seed plant.
The hypocotyl is the stem of a germinating seedling, found below the cotyledons and above the radicle (root).
A nursery is a place where plants are propagated and grown to a desired size. Mostly the plants concerned are for gardening, forestry, or conservation biology, rather than agriculture. They include retail nurseries, which sell to the general public; wholesale nurseries, which sell only to businesses such as other nurseries and commercial gardeners; and private nurseries, which supply the needs of institutions or private estates. Some will also work in plant breeding.
Plant physiology is a subdiscipline of botany concerned with the functioning, or physiology, of plants.
Silviculture is the practice of controlling the growth, composition/structure, as well as quality of forests to meet values and needs, specifically timber production.
In agriculture and gardening, transplanting or replanting is the technique of moving a plant from one location to another. Most often this takes the form of starting a plant from seed in optimal conditions, such as in a greenhouse or protected nursery bed, then replanting it in another, usually outdoor, growing location. The agricultural machine that does this is called a transplanter. This is common in market gardening and truck farming, where setting out or planting out are synonymous with transplanting. In the horticulture of some ornamental plants, transplants are used infrequently and carefully because they carry with them a significant risk of killing the plant.
Plant embryonic development, also plant embryogenesis, is a process that occurs after the fertilization of an ovule to produce a fully developed plant embryo. This is a pertinent stage in the plant life cycle that is followed by dormancy and germination. The zygote produced after fertilization must undergo various cellular divisions and differentiations to become a mature embryo. An end stage embryo has five major components including the shoot apical meristem, hypocotyl, root meristem, root cap, and cotyledons. Unlike the embryonic development in animals, and specifically in humans, plant embryonic development results in an immature form of the plant, lacking most structures like leaves, stems, and reproductive structures. However, both plants and animals including humans, pass through a phylotypic stage that evolved independently and that causes a developmental constraint limiting morphological diversification.
In developmental biology, photomorphogenesis is light-mediated development, where plant growth patterns respond to the light spectrum. This is a completely separate process from photosynthesis where light is used as a source of energy. Phytochromes, cryptochromes, and phototropins are photochromic sensory receptors that restrict the photomorphogenic effect of light to the UV-A, UV-B, blue, and red portions of the electromagnetic spectrum.
Shade avoidance is a set of responses that plants display when they are subjected to the shade of another plant. It often includes elongation, altered flowering time, increased apical dominance and altered partitioning of resources. This set of responses is collectively called the shade-avoidance syndrome (SAS).
Cultivation of cannabis is the production of cannabis infructescences. Cultivation techniques for other purposes differ.
An epicotyl is important for the beginning stages of a plant's life. It is the region of a seedling stem above the stalks of the seed leaves of an embryo plant. It grows rapidly, showing hypogeal germination, and extends the stem above the soil surface. A common misconception is that the epicotyl, being closer to the apex of the plant, is the first part to emerge after germination - rather, the hypocotyl, the region of the stem between the point of attachment of the cotyledons and the root - forms a hook during hypogeal germination and pushes out of the soil, allowing the more delicate tissues of the plumules and apical meristem to avoid damage from pushing through the soil. The epicotyl will expand and form the point of attachment of the shoot apex and leaf primordia or "first true leaves". Cotyledons may remain belowground or be pushed up aboveground with the growing stem depending on the plant species in question.
Etiolation is a process in flowering plants grown in partial or complete absence of light. It is characterized by long, weak stems; smaller leaves due to longer internodes; and a pale yellow color (chlorosis). The development of seedlings in the dark is known as "skotomorphogenesis" and leads to etiolated seedlings.
Important structures in plant development are buds, shoots, roots, leaves, and flowers; plants produce these tissues and structures throughout their life from meristems located at the tips of organs, or between mature tissues. Thus, a living plant always has embryonic tissues. By contrast, an animal embryo will very early produce all of the body parts that it will ever have in its life. When the animal is born, it has all its body parts and from that point will only grow larger and more mature. However, both plants and animals pass through a phylotypic stage that evolved independently and that causes a developmental constraint limiting morphological diversification.
Gap dynamics refers to the pattern of plant growth that occurs following the creation of a forest gap, a local area of natural disturbance that results in an opening in the canopy of a forest. Gap dynamics are a typical characteristic of both temperate and tropical forests and have a wide variety of causes and effects on forest life.
Hypogeal germination is a botanical term indicating that the germination of a plant takes place below the ground. An example of a plant with hypogeal germination is the pea. The opposite of hypogeal is epigeal.
Wisconsin Fast Plants is the registered trademark for a cultivar of Brassica rapa, developed as a rapid life-cycle model organism for research and teaching. Wisconsin Fast Plants are a member of the Brassicaceae family, closely related to the turnip and bok choy. Wisconsin Fast Plants were developed in accordance with an ideotype for an ideal model organism to be used in expediting plant research. Similarly, their rapid life cycle and other model organism characteristics made them easy to grow in large numbers in classrooms. For the last few decades they have been grown in classrooms and laboratories around the world.