|Korean red pine (Pinus densiflora), North Korea|
|Range of Pinus|
A pine is any conifer in the genus Pinus ( // ) of the family Pinaceae. Pinus is the sole genus in the subfamily Pinoideae. The Plant List compiled by the Royal Botanic Gardens, Kew and Missouri Botanical Garden accepts 126 species names of pines as current, together with 35 unresolved species and many more synonyms.
The Pinophyta, also known as Coniferophyta or Coniferae, or commonly as conifers, are a division of vascular land plants containing a single extant class, Pinopsida. They are gymnosperms, cone-bearing seed plants. All extant conifers are perennial woody plants with secondary growth. The great majority are trees, though a few are shrubs. Examples include cedars, Douglas firs, cypresses, firs, junipers, kauri, larches, pines, hemlocks, redwoods, spruces, and yews. As of 1998, the division Pinophyta was estimated to contain eight families, 68 genera, and 629 living species.
A genus is a taxonomic rank used in the biological classification of living and fossil organisms, as well as viruses, in biology. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus.
Family is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy; it is classified between order and genus. A family may be divided into subfamilies, which are intermediate ranks between the ranks of family and genus. The official family names are Latin in origin; however, popular names are often used: for example, walnut trees and hickory trees belong to the family Juglandaceae, but that family is commonly referred to as being the "walnut family".
The modern English name "pine" derives from Latin pinus, which some have traced to the Indo-European base *pīt- ‘resin’ (source of English pituitary).Before the 19th century, pines were often referred to as firs (from Old Norse fura, by way of Middle English firre). In some European languages, Germanic cognates of the Old Norse name are still in use for pines—in Danish fyr, in Norwegian fura/fure/furu, Swedish fura/furu, Dutch vuren, and German Föhre—but in modern English, fir is now restricted to fir (Abies) and Douglas fir (Pseudotsuga).
Middle English was a form of the English language, spoken after the Norman conquest (1066) until the late 15th century. English underwent distinct variations and developments following the Old English period. Scholarly opinion varies, but the Oxford English Dictionary specifies the period when Middle English was spoken as being from 1150 to 1500. This stage of the development of the English language roughly followed the High to the Late Middle Ages.
Danish is a North Germanic language spoken by around six million people, principally in Denmark and in the region of Southern Schleswig in northern Germany, where it has minority language status. Also, minor Danish-speaking communities are found in Norway, Sweden, Spain, the United States, Canada, Brazil, and Argentina. Due to immigration and language shift in urban areas, around 15–20% of the population of Greenland speak Danish as their first language.
Norwegian is a North Germanic language spoken mainly in Norway, where it is the official language. Along with Swedish and Danish, Norwegian forms a dialect continuum of more or less mutually intelligible local and regional varieties, and some Norwegian and Swedish dialects, in particular, are very close. These Scandinavian languages, together with Faroese and Icelandic as well as some extinct languages, constitute the North Germanic languages. Faroese and Icelandic are hardly mutually intelligible with Norwegian in their spoken form because continental Scandinavian has diverged from them. While the two Germanic languages with the greatest numbers of speakers, English and German, have close similarities with Norwegian, neither is mutually intelligible with it. Norwegian is a descendant of Old Norse, the common language of the Germanic peoples living in Scandinavia during the Viking Era.
Pine trees are evergreen, coniferous resinous trees (or, rarely, shrubs) growing 3–80 m (10–260 ft) tall, with the majority of species reaching 15–45 m (50–150 ft) tall. The smallest are Siberian dwarf pine and Potosi pinyon, and the tallest is an 81.79 m (268.35 ft) tall ponderosa pine located in southern Oregon's Rogue River-Siskiyou National Forest.
In botany, an evergreen is a plant that has leaves throughout the year that are always green. This is true even if the plant retains its foliage only in warm climates, and contrasts with deciduous plants, which completely lose their foliage during the winter or dry season. There are many different kinds of evergreen plants, both trees and shrubs. Evergreens include:
In polymer chemistry and materials science, resin is a solid or highly viscous substance of plant or synthetic origin that is typically convertible into polymers. Resins are usually mixtures of organic compounds. This article focuses on naturally-occurring resins.
In botany, a tree is a perennial plant with an elongated stem, or trunk, supporting branches and leaves in most species. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are usable as lumber or plants above a specified height. Trees are not a taxonomic group but include a variety of plant species that have independently evolved a woody trunk and branches as a way to tower above other plants to compete for sunlight. Trees tend to be long-lived, some reaching several thousand years old. In wider definitions, the taller palms, tree ferns, bananas, and bamboos are also trees. Trees have been in existence for 370 million years. It is estimated that there are just over 3 trillion mature trees in the world.
Pines are long lived and typically reach ages of 100–1,000 years, some even more. The longest-lived is the Great Basin bristlecone pine, Pinus longaeva. One individual of this species, dubbed "Methuselah", is one of the world's oldest living organisms at around 4,600 years old. This tree can be found in the White Mountains of California. [ citation needed ]An older tree, now cut down, was dated at 4,900 years old. It was discovered in a grove beneath Wheeler Peak and it is now known as "Prometheus" after the Greek immortal.
Methuselah is a 4,850-year-old Great Basin bristlecone pine tree growing high in the White Mountains of Inyo County in eastern California. It is recognized as the tree with the greatest confirmed age in the world.
The White Mountains of California and Nevada are a triangular fault-block mountain range facing the Sierra Nevada across the upper Owens Valley. They extend for approximately 60 mi (97 km) as a greatly elevated plateau about 20 mi (32 km) wide on the south, narrowing to a point at the north, with elevations generally increasing south to north. The range's broad southern end is near the community of Big Pine, where Westgard Pass and Deep Springs Valley separate it from the Inyo Mountains. The narrow northern end is at Montgomery Pass, where U.S. Route 6 crosses. The Fish Lake Valley lies east of the range; the southeast part of the mountains are separated from the Silver Peak Range by block faulting across the Furnace Creek Fault Zone, forming a feeder valley to Fish Lake Valley. The range lies within the eastern section of the Inyo National Forest.
Wheeler Peak is the tallest mountain in the Snake Range and in White Pine County, in Nevada, United States. The summit elevation of 13,065 feet (3,982 m) makes it the second-highest peak in Nevada, just behind Boundary Peak. With a topographic prominence of 7,563 feet (2,305 m), Wheeler Peak is the most topographically prominent peak in White Pine County and the second-most prominent peak in Nevada, just behind Mount Charleston. The mountain is located in Great Basin National Park and was named for George Wheeler, leader of the Wheeler Survey of the late 19th century.
The bark of most pines is thick and scaly, but some species have thin, flaky bark. The branches are produced in regular "pseudo whorls", actually a very tight spiral but appearing like a ring of branches arising from the same point. Many pines are uninodal, producing just one such whorl of branches each year, from buds at the tip of the year's new shoot, but others are multinodal, producing two or more whorls of branches per year.
Bark is the outermost layers of stems and roots of woody plants. Plants with bark include trees, woody vines, and shrubs. Bark refers to all the tissues outside the vascular cambium and is a nontechnical term. It overlays the wood and consists of the inner bark and the outer bark. The inner bark, which in older stems is living tissue, includes the innermost area of the periderm. The outer bark in older stems includes the dead tissue on the surface of the stems, along with parts of the innermost periderm and all the tissues on the outer side of the periderm. The outer bark on trees which lies external to the last formed periderm is also called the rhytidome.
In botany, a bud is an undeveloped or embryonic shoot and normally occurs in the axil of a leaf or at the tip of a stem. Once formed, a bud may remain for some time in a dormant condition, or it may form a shoot immediately. Buds may be specialized to develop flowers or short shoots, or may have the potential for general shoot development. The term bud is also used in zoology, where it refers to an outgrowth from the body which can develop into a new individual.
A stem is one of two main structural axes of a vascular plant, the other being the root. The stem is normally divided into nodes and internodes:
The spiral growth of branches, needles, and cone scales may be arranged in Fibonacci number ratios.[ citation needed ] The new spring shoots are sometimes called "candles"; they are covered in brown or whitish bud scales and point upward at first, then later turn green and spread outward. These "candles" offer foresters a means to evaluate fertility of the soil and vigour of the trees.
In mathematics, the Fibonacci numbers, commonly denoted Fn form a sequence, called the Fibonacci sequence, such that each number is the sum of the two preceding ones, starting from 0 and 1. That is,
A forester is a person who practices forestry, the science, art, and profession of managing forests. Foresters engage in a broad range of activities including ecological restoration and management of protected areas. Foresters manage forests to provide a variety of objectives including direct extraction of raw material, outdoor recreation, conservation, hunting and aesthetics. Emerging management practices include managing forestlands for biodiversity, carbon sequestration and air quality.
Pines have four types of leaf:
Pines are mostly monoecious, having the male and female cones on the same tree, though a few species are sub-dioecious, with individuals predominantly, but not wholly, single-sex. The male cones are small, typically 1–5 cm long, and only present for a short period (usually in spring, though autumn in a few pines), falling as soon as they have shed their pollen. The female cones take 1.5–3 years (depending on species) to mature after pollination, with actual fertilization delayed one year. At maturity the female cones are 3–60 cm long. Each cone has numerous spirally arranged scales, with two seeds on each fertile scale; the scales at the base and tip of the cone are small and sterile, without seeds.
The seeds are mostly small and winged, and are anemophilous (wind-dispersed), but some are larger and have only a vestigial wing, and are bird-dispersed (see below). At maturity, the cones usually open to release the seeds, but in some of the bird-dispersed species (e.g. whitebark pine), the seeds are only released by the bird breaking the cones open. In others, the seeds are stored in closed ("serotinous") cones for many years until an environmental cue triggers the cones to open, releasing the seeds. The most common form of serotiny is pyriscence, in which a resin binds the cones shut until melted by a forest fire.
Conifers developed about 300 million years ago, and pines perhaps around 153 million years ago. The genus Pinus is thought to have diverged from other pines approximately 95 million years ago.
Pines are gymnosperms. The genus is divided into two subgenera, which can be distinguished by cone, seed, and leaf characters:
Pines are native to the Northern Hemisphere, and in a few parts of the tropics in the Southern Hemisphere. Most regions of the Northern Hemisphere (see List of pines by region) host some native species of pines. One species (Sumatran pine) crosses the equator in Sumatra to 2°S. In North America, various species occur in regions at latitudes from as far north as 66°N to as far south as 12°N.[ citation needed ]
Pines may be found in a very large variety of environments, ranging from semi-arid desert to rainforests, from sea level up to 5,200 metres (17,100 ft), from the coldest to the hottest environments on Earth. They often occur in mountainous areas with favorable soils and at least some water.[ citation needed ]
Various species have been introduced to temperate and subtropical regions of both hemispheres, where they are grown as timber or cultivated as ornamental plants in parks and gardens. A number of such introduced species have become naturalized, and some species are considered invasive in some areasand threaten native ecosystems.
Pines grow well in acid soils, some also on calcareous soils; most require good soil drainage, preferring sandy soils, but a few (e.g. lodgepole pine) can tolerate poorly drained wet soils. A few are able to sprout after forest fires (e.g. Canary Island pine). Some species of pines (e.g. bishop pine) need fire to regenerate, and their populations slowly decline under fire suppression regimens.
Several species are adapted to extreme conditions imposed by elevation and latitude (e.g. Siberian dwarf pine, mountain pine, whitebark pine, and the bristlecone pines). The pinyon pines and a number of others, notably Turkish pine and gray pine, are particularly well adapted to growth in hot, dry semidesert climates.[ citation needed ]
The seeds are commonly eaten by birds, such as grouse, crossbills, jays, nuthatches, siskins, and woodpeckers, and by squirrels. Some birds, notably the spotted nutcracker, Clark's nutcracker, and pinyon jay, are of importance in distributing pine seeds to new areas. Pine needles are sometimes eaten by some Lepidoptera (butterfly and moth) species (see list of Lepidoptera that feed on pines), the Symphytan species pine sawfly, and goats.[ citation needed ]
Pine pollen may play an important role in the functioning of detrital food webs.Nutrients from pollen aid detritivores in development, growth, and maturation, and may enable fungi to decompose nutritionally scarce litter. Pine pollen is also involved in moving plant matter between terrestrial and aquatic ecosystems.
Pines are among the most commercially important tree species valued for their timber and wood pulp throughout the world.In temperate and tropical regions, they are fast-growing softwoods that grow in relatively dense stands, their acidic decaying needles inhibiting the sprouting of competing hardwoods. Commercial pines are grown in plantations for timber that is denser and therefore more durable than spruce (Picea). Pine wood is widely used in high-value carpentry items such as furniture, window frames, panelling, floors, and roofing, and the resin of some species is an important source of turpentine.
Because pines have no insect- or decay-resistant qualities after logging, untreated they are generally recommended for construction purposes as indoor use only (indoor drywall framing, for example). For outside use, pine needs to be treated with copper azole, chromated copper arsenate or other suitable chemical preservative.
Many pine species make attractive ornamental plantings for parks and larger gardens with a variety of dwarf cultivars being suitable for smaller spaces. Pines are also commercially grown and harvested for Christmas trees. Pine cones, the largest and most durable of all conifer cones, are craft favorites. Pine boughs, appreciated especially in wintertime for their pleasant smell and greenery, are popularly cut for decorations.Pine needles are also used for making decorative articles such as baskets, trays, pots, etc, and during the U.S. Civil War, the needles of the longleaf pine "Georgia pine" were widely employed in this. This originally Native American skill is now being replicated across the world. Pine needle handicrafts are made in the US, Canada, Mexico, Nicaragua, and India. Pine needles are also versatile and have been used by Latvian designer Tamara Orjola to create different biodegradable products including paper, furniture, textiles and dye.
Pine needles serve as food for various Lepidoptera. See List of Lepidoptera that feed on pines. Several species are attacked by nematodes, causing pine wilt disease, which can kill some quickly.
When grown for sawing timber, pine plantations can be harvested after 30 years, with some stands being allowed to grow up to 50 (as the wood value increases more quickly as the trees age). Imperfect trees (such as those with bent trunks or forks, smaller trees, or diseased trees) are removed in a "thinning" operation every 5–10 years. Thinning allows the best trees to grow much faster, because it prevents weaker trees from competing for sunlight, water, and nutrients. Young trees removed during thinning are used for pulpwood, while most older ones are good enough for saw timber.
The final wood quality can be improved by pruning small branches at ages 5, 7, and 9. Pruning usually goes up to a height of 6 metres (20 ft). This results in smooth timber with no knots, which is considerably more valuable.
A 30-year-old commercial pine tree grown in good conditions will be about 0.3 m (1.0 ft) in diameter and about 20 m (66 ft) high. After 50 years, the same tree will be about 0.5 m (1.6 ft) in diameter and 25 m (82 ft) high, and its wood will be worth about seven times as much as the 30-year-old tree.
Trees are planted 3–4 m apart, or about 1000 per hectare (100,000 per km2).
Some species have large seeds, called pine nuts, that are harvested and sold for cooking and baking. They are an essential ingredient of pesto alla genovese.
The soft, moist, white inner bark (cambium) found clinging to the woody outer bark is edible and very high in vitamins A and C. It can be eaten raw in slices as a snack or dried and ground up into a powder for use as an ersatz flour or thickener in stews, soups, and other foods, such as bark bread. Adirondack Indians got their name from the Mohawk Indian word atirú:taks, meaning "tree eaters".
A tea made by steeping young, green pine needles in boiling water (known as tallstrunt in Sweden) is high in vitamins A and C.[ citation needed ] In eastern Asia, pine and other conifers are accepted among consumers as a beverage product, and used in teas, as well as wine.
Pine needles from Pinus densiflora were found to contain 30.54 mg/g of proanthocyanidins when extracted with hot water.Comparative to ethanol extraction resulting in 30.11 mg/g, simply extracting in hot water is preferable.
In traditional Chinese medicine (TCM), pine resin is used for burns, wounds and dermal complaints.
Pines have been a frequently mentioned tree throughout history, including in literature, paintings and other art, and in religious texts.
Writers of various nationalities and ethnicities have written of pines. Among them, John Muir,Dora Sigerson Shorter, Eugene Field, the Chinese, Theodore Winthrop, and Rev. George Allan D.D.
Pines are often featured in art, whether painting and fine art,drawing, photography, or folk art.
Pine trees, as well as other conifers, are mentioned in The Bible. In Nehemiah 8:15, the King James Version renders the following translation:
"And that they should publish and proclaim in all their cities, and in Jerusalem, saying, Go forth unto the mount, and fetch olive branches, and pine branches [emphasis added], and myrtle branches, and palm branches, and branches of thick trees, to make booths, as it is written."
Pines are also mentioned in Isaiah 41:
"17: When the poor and needy seek water, and there is none, and their tongue faileth for thirst, I the LORD will hear them, I the God of Israel will not forsake them. 18: I will open rivers in high places, and fountains in the midst of the valleys: I will make the wilderness a pool of water, and the dry land springs of water. 19: I will plant in the wilderness the cedar, the shittah tree, and the myrtle, and the oil tree; I will set in the desert the fir tree, and the pine, and the box tree together: 20: That they may see, and know, and consider, and understand together, that the hand of the LORD hath done this, and the Holy One of Israel hath created it."
And in Isaiah 60:
"13: The glory of Lebanon shall come unto thee, the fir tree, the pine tree, and the box together, to beautify the place of my sanctuary; and I will make the place of my feet glorious."
Pseudotsuga menziesii is an evergreen conifer species in the pine family, Pinaceae. It is native to western North America and is known as Douglas fir, Douglas-fir, Oregon pine, and Columbian pine. There are two varieties: coast Douglas-fir, and Rocky Mountain Douglas-fir.
The Pinaceae are trees or shrubs, including many of the well-known conifers of commercial importance such as cedars, firs, hemlocks, larches, pines and spruces. The family is included in the order Pinales, formerly known as Coniferales. Pinaceae are supported as monophyletic by their protein-type sieve cell plastids, pattern of proembryogeny, and lack of bioflavonoids. They are the largest extant conifer family in species diversity, with between 220 and 250 species in 11 genera, and the second-largest in geographical range, found in most of the Northern Hemisphere, with the majority of the species in temperate climates, but ranging from subarctic to tropical. The family often forms the dominant component of boreal, coastal, and montane forests. One species, Pinus merkusii, grows just south of the equator in Southeast Asia. Major centres of diversity are found in the mountains of southwest China, Mexico, central Japan, and California.
Pinus albicaulis, known by the common names whitebark pine, white pine, pitch pine, scrub pine, and creeping pine, is a conifer tree native to the mountains of the western United States and Canada, specifically subalpine areas of the Sierra Nevada, Cascade Range, Pacific Coast Ranges, and Rocky Mountains from Wyoming northwards. It shares the common name "creeping pine" with several other plants.
Pinus nigra, the Austrian pine or black pine, is a moderately variable species of pine, occurring across southern Mediterranean Europe from Spain to the eastern Mediterranean on Anatolian peninsula of Turkey and on Corsica/Cyprus, including Crimea, and in the high mountains of the Maghreb in North Africa.
Pinus jeffreyi also known as Jeffrey pine, Jeffrey's pine, yellow pine and black pine, is a North American pine tree. It is mainly found in California, but also in the westernmost part of Nevada, southwestern Oregon, and northern Baja California. It is named in honor of its botanist documenter John Jeffrey.
Pinus strobiformis, commonly known as southwestern white pine, Mexican white pine or Chihuahua white pine, is a medium-sized white pine tree whose native habitat is in southwestern United States and Mexico. It is typically a high-elevation pine growing mixed with other conifers.
Pinus strobus, commonly denominated the eastern white pine, northern white pine, white pine, Weymouth pine (British), and soft pine is a large pine native to eastern North America. It occurs from Newfoundland, Canada west through the Great Lakes region to southeastern Manitoba and Minnesota, United States, and south along the Appalachian Mountains and upper Piedmont to northernmost Georgia and perhaps very rarely in some of the higher elevations in northeastern Alabama.
Pinus resinosa, known as red pine or Norway pine, is a pine native to North America. It occurs from Newfoundland west to Manitoba, and south to Pennsylvania, with several smaller, disjunct populations occurring in the Appalachian Mountains in Virginia and West Virginia, as well as a few small pockets in extreme northern New Jersey and northern Illinois.
Pinus wallichiana is a coniferous evergreen tree native to the Himalaya, Karakoram and Hindu Kush mountains, from eastern Afghanistan east across northern Pakistan and north west India to Yunnan in southwest China. It grows in mountain valleys at altitudes of 1800–4300 m, between 30 m and 50 m in height. It favours a temperate climate with dry winters and wet summers. In Pashto, it is known as Nishtar.
Pinus virginiana, the Virginia pine, scrub pine, Jersey pine, is a medium-sized tree, often found on poorer soils from Long Island in southern New York south through the Appalachian Mountains to western Tennessee and Alabama. The usual size range for this pine is 9–18 m, but can grow taller under optimum conditions. The trunk can be as large as 0.5 m diameter. This tree prefers well-drained loam or clay, but will also grow on very poor, sandy soil, where it remains small and stunted. The typical life span is 65 to 90 years.
Jack pine is an eastern North American pine. Its native range in Canada is east of the Rocky Mountains from the Mackenzie River in the Northwest Territories to Cape Breton Island in Nova Scotia, and the north-central and northeast of the United States from Minnesota to Maine, with the southernmost part of the range just into northwest Indiana and northwest Pennsylvania. It is also known as grey pine and scrub pine.
Abies balsamea or balsam fir is a North American fir, native to most of eastern and central Canada and the northeastern United States.
Pinus longaeva is a long-living species of bristlecone pine tree found in the higher mountains of California, Nevada, and Utah. Methuselah is a bristlecone pine that is 4,850 years old and the oldest known living non-clonal organism on Earth. In 1987, the bristlecone pine was designated one of Nevada's state trees.
Pinus roxburghii, commonly known as chir pine or longleaf Indian pine, is a species of pine, native to the Himalayas. It was named after William Roxburgh.
Pinus kesiya is one of the most widely distributed pines in Asia. Its range extends south and east from the Khasi Hills in the northeast Indian state of Meghalaya, to northern Thailand, Philippines, Burma, Cambodia, Laos, southernmost China, and Vietnam. It is an important plantation species elsewhere in the world, including in southern Africa and South America.
Pinus quadrifolia, the Parry pinyon, is a pine in the pinyon pine group native to southernmost California in the United States and northern Baja California in Mexico, from 33° 30' N south to 30° 30' N. It occurs at moderate altitudes from 1,300 metres (4,300 ft) to 1,800 metres (5,900 ft), rarely as low as 1,200 metres (3,900 ft) and as high as 2,500 metres (8,200 ft). It is scarce and often scattered in this region, forming open woodlands, usually mixed with junipers. Other common names include nut pine and fourleaf pinyon pine.
Pinus aristata, the Rocky Mountain bristlecone pine, is a long-living species of bristlecone pine tree native to the United States. It appears in the Rocky Mountains in Colorado and northern New Mexico, with isolated populations in the San Francisco Peaks in Arizona and the Kaibab National Forest north of the Grand Canyon. It is usually found at very high altitudes, from 7,000–13,000 feet (2,100–4,000 m), in cold, dry subalpine climate conditions, often at the tree line, although it also forms extensive closed-canopy stands at somewhat lower elevations.
Pseudotsuga menziesii var. menziesii, also known as coast Douglas-fir, Pacific Douglas-fir, Oregon pine, or Douglas spruce, is an evergreen conifer native to western North America from west-central British Columbia, Canada southward to central California, United States. In Oregon and Washington its range is continuous from the Cascades crest west to the Pacific Coast Ranges and Pacific Ocean. In California, it is found in the Klamath and California Coast Ranges as far south as the Santa Lucia Mountains with a small stand as far south as the Purisima Hills, Santa Barbara County. In the Sierra Nevada it ranges as far south as the Yosemite region. It occurs from near sea level along the coast to 1,800 metres (5,900 ft) in the California Mountains. Further inland, coast Douglas-fir is replaced by Rocky Mountain or interior Douglas-fir. Interior Douglas-fir intergrades with coast Douglas-fir in the Cascades of northern Washington and southern British Columbia.
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