Evergreen

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A silver fir shoot showing three successive years of retained leaves. Abies alba 02.jpg
A silver fir shoot showing three successive years of retained leaves.

In botany, an evergreen is a plant which has foliage that remains green and functional throughout the year. This contrasts with deciduous plants, which lose their foliage completely during the winter or dry season.

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Evergreen species

There are many different kinds of evergreen plants, both trees and shrubs. Evergreens include:

The Latin binomial term sempervirens , meaning "always green", refers to the evergreen nature of the plant, for instance:

Cupressus sempervirens (a cypress)
Lonicera sempervirens (a honeysuckle)
Sequoia sempervirens (a sequoia)

The longevity of individual leaves in evergreen plants varies from a few months to several decades (over 30 years in the Great Basin bristlecone pine [1] ).

Evergreen families

Family nameExample
Araucariaceae Kauri
Cupressaceae Sequoia
Pinaceae Pine
Podocarpaceae Real yellowwood
Taxaceae Yew
Cyatheaceae Australian tree fern
Aquifoliaceae Holly
Fagaceae Live oak
Oleaceae Shamel ash
Myrtaceae Eucalyptus
Arecaceae Coconut
Lauraceae Bay
Magnoliaceae Southern magnolia
Cycadaceae Queen sago

Japanese umbrella pine is unique in that it has its own family of which it is the only species.

Differences between evergreen and deciduous species

Evergreen and deciduous species vary in a range of morphological and physiological characters. Generally, broad-leaved evergreen species have thicker leaves than deciduous species, with a larger volume of parenchyma and air spaces per unit leaf area. [2] They have larger leaf biomass per unit leaf area, and hence a lower specific leaf area. Construction costs don’t differ between the groups.[ citation needed ] Evergreens have generally a larger fraction of total plant biomass present as leaves (LMF), [3] but they often have a lower rate of photosynthesis.

Reasons for being evergreen or deciduous

A southern live oak in South Carolina during winter Live oak Georgetown.jpg
A southern live oak in South Carolina during winter

Deciduous trees shed their leaves usually as an adaptation to a cold or dry/wet season. Evergreen trees also lose leaves, but each tree loses its leaves gradually and not all at once. Most tropical rainforest plants are considered to be evergreens, replacing their leaves gradually throughout the year as the leaves age and fall, whereas species growing in seasonally arid climates may be either evergreen or deciduous. Most warm temperate climate plants are also evergreen.[ citation needed ] In cool temperate climates, fewer plants are evergreen. In such climates, there is a predominance of conifers because few evergreen broadleaf plants can tolerate severe cold below about −26 °C (−15 °F).[ clarification needed ][ citation needed ]

In areas where there is a reason for being deciduous, e.g. a cold season or dry season, evergreen plants are usually an adaptation of low nutrient levels. Additionally, they usually have hard leaves and have an excellent water economy due to scarce resources in the area in which they reside. [4] The excellent water economy within the evergreen species is due to high abundance when compared to deciduous species. [4] Whereas deciduous trees lose nutrients whenever they lose their leaves. In warmer areas, species such as some pines and cypresses grow on poor soils and disturbed ground.[ citation needed ] In Rhododendron , a genus with many broadleaf evergreens, several species grow in mature forests but are usually found on highly acidic soil where the nutrients are less available to plants.[ citation needed ] In taiga or boreal forests, it is too cold for the organic matter in the soil to decay rapidly, so the nutrients in the soil are less easily available to plants, thus favoring evergreens.[ citation needed ]

In temperate climates, evergreens can reinforce their own survival; evergreen leaf and needle litter has a higher carbon-nitrogen ratio than deciduous leaf litter, contributing to a higher soil acidity and lower soil nitrogen content. This is the case with Mediterranean evergreen seedlings, which have unique C and N storages that allow stored resources to determine fast growth within the species, limiting competition and bolstering survival. [5] These conditions favor the growth of more evergreens and make it more difficult for deciduous plants to persist. In addition, the shelter provided by existing evergreen plants can make it easier for younger evergreen plants to survive cold and/or drought. [6] [7] [8]

See also


Related Research Articles

<span class="mw-page-title-main">Taiga</span> Biome characterized by coniferous forests

Taiga, generally referred to in North America as a boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruces, and larches.

<span class="mw-page-title-main">Deciduous</span> Plants that shed leaves seasonally

In the fields of horticulture and botany, the term deciduous means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, after flowering; and to the shedding of ripe fruit. The antonym of deciduous in the botanical sense is evergreen.

<span class="mw-page-title-main">Conifer</span> Group of cone-bearing seed plants

Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta, also known as Coniferophyta or Coniferae. The division contains a single extant class, Pinopsida. All extant conifers are perennial woody plants with secondary growth. The great majority are trees, though a few are shrubs. Examples include cedars, Douglas-firs, cypresses, firs, junipers, kauri, larches, pines, hemlocks, redwoods, spruces, and yews. The division Pinophyta contains seven families, 60 to 65 genera, and more than 600 living species.

<span class="mw-page-title-main">Sclerophyll</span> Type of plant

Sclerophyll is a type of vegetation that is adapted to long periods of dryness and heat. The plants feature hard leaves, short internodes and leaf orientation which is parallel or oblique to direct sunlight. The word comes from the Greek sklēros (hard) and phyllon (leaf). The term was coined by A.F.W. Schimper in 1898, originally as a synonym of xeromorph, but the two words were later differentiated.

<span class="mw-page-title-main">Temperate coniferous forest</span> Forests found in areas with warm summers and cool winters

Temperate coniferous forest is a terrestrial biome defined by the World Wide Fund for Nature. Temperate coniferous forests are found predominantly in areas with warm summers and cool winters, and vary in their kinds of plant life. In some, needleleaf trees dominate, while others are home primarily to broadleaf evergreen trees or a mix of both tree types. A separate habitat type, the tropical coniferous forests, occurs in more tropical climates.

<span class="mw-page-title-main">Perennial</span> Plant that lives for more than two years

In botany, a perennial plant or simply perennial is a plant that lives more than two years. The term is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also widely used to distinguish plants with little or no woody growth from trees and shrubs, which are also technically perennials. Notably, it is estimated that 94% of plant species fall under the category of perennials, underscoring the prevalence of plants with lifespans exceeding two years in the botanical world.

<span class="mw-page-title-main">Coarse woody debris</span>

Coarse woody debris (CWD) or coarse woody habitat (CWH) refers to fallen dead trees and the remains of large branches on the ground in forests and in rivers or wetlands. A dead standing tree – known as a snag – provides many of the same functions as coarse woody debris. The minimum size required for woody debris to be defined as "coarse" varies by author, ranging from 2.5–20 cm (1–8 in) in diameter.

<i>Cupressus sempervirens</i> Species of conifer

Cupressus sempervirens, the Mediterranean cypress, is a species of cypress native to the eastern Mediterranean region and Iran. Cupressus sempervirens is a seeded vascular plant, meaning that it contains both xylem and phloem. Since it is a seeded plant, it uses seeds to reproduce. It has many uses that can contribute to medicine and other things that are beneficial, whether that be the seeds that it produces or the oil that comes from the leaves. It is well adapted to the conditions and the environment that it lives in. Cupressus sempervirens is widely present in culture, most notably in Iran, where it is both a sacred tree and is a metaphor for "the graceful figure of the beloved".

A temperate forest is a forest found between the tropical and boreal regions, located in the temperate zone. It is the second largest biome on our planet, covering 25% of the world's forest area, only behind the boreal forest, which covers about 33%. These forests cover both hemispheres at latitudes ranging from 25 to 50 degrees, wrapping the planet in a belt similar to that of the boreal forest. Due to its large size spanning several continents, there are several main types: deciduous, coniferous, mixed forest, and rainforest.

<span class="mw-page-title-main">Ecosystem ecology</span> Study of living and non-living components of ecosystems and their interactions

Ecosystem ecology is the integrated study of living (biotic) and non-living (abiotic) components of ecosystems and their interactions within an ecosystem framework. This science examines how ecosystems work and relates this to their components such as chemicals, bedrock, soil, plants, and animals.

<span class="mw-page-title-main">Abscission</span> Shedding of various parts of an organism

Abscission is the shedding of various parts of an organism, such as a plant dropping a leaf, fruit, flower, or seed. In zoology, abscission is the intentional shedding of a body part, such as the shedding of a claw, husk, or the autotomy of a tail to evade a predator. In mycology, it is the liberation of a fungal spore. In cell biology, abscission refers to the separation of two daughter cells at the completion of cytokinesis.

<span class="mw-page-title-main">California mixed evergreen forest</span>

California mixed evergreen forest is a plant community found in the mountain ranges of California and southwestern Oregon.

<span class="mw-page-title-main">Oasification</span> Antonym to desertification by soil erosion

In hydrology, oasification is the antonym to desertification by soil erosion. This technique has limited application and is normally considered for much smaller areas than those threatened by desertification.

<span class="mw-page-title-main">Temperate deciduous forest</span> Deciduous forest in the temperate regions

Temperate deciduous or temperate broad-leaf forests are a variety of temperate forest 'dominated' by deciduous trees that lose their leaves each winter. They represent one of Earth's major biomes, making up 9.69% of global land area. These forests are found in areas with distinct seasonal variation that cycle through warm, moist summers, cold winters, and moderate fall and spring seasons. They are most commonly found in the Northern Hemisphere, with particularly large regions in eastern North America, East Asia, and a large portion of Europe, though smaller regions of temperate deciduous forests are also located in South America. Examples of trees typically growing in the Northern Hemisphere's deciduous forests include oak, maple, basswood, beech and elm, while in the Southern Hemisphere, trees of the genus Nothofagus dominate this type of forest. Temperate deciduous forests provide several unique ecosystem services, including habitats for diverse wildlife, and they face a set of natural and human-induced disturbances that regularly alter their structure.

<span class="mw-page-title-main">Woody plant</span> Plant that produces wood and has a hard stem

A woody plant is a plant that produces wood as its structural tissue and thus has a hard stem. In cold climates, woody plants further survive winter or dry season above ground, as opposed to herbaceous plants that die back to the ground until spring.

<span class="mw-page-title-main">Plant litter</span> Dead plant material that has fallen to the ground

Plant litter is dead plant material that have fallen to the ground. This detritus or dead organic material and its constituent nutrients are added to the top layer of soil, commonly known as the litter layer or O horizon. Litter is an important factor in ecosystem dynamics, as it is indicative of ecological productivity and may be useful in predicting regional nutrient cycling and soil fertility.

<span class="mw-page-title-main">Drought deciduous</span> Plants that shed their leaves during periods of drought or generally in a dry season

Drought deciduous, or drought semi-deciduous plants refers to plants that shed their leaves during periods of drought or in the dry season. This phenomenon is a natural process of plants and is caused due to the limitation of water around the environment where the plant is situated. In the spectrum of botany, deciduous is defined as a certain plant species that carry out abscission, the shedding of leaves of a plant or tree either due to age or other factors that causes the plant to regard these leaves as useless or not worth keeping over the course of a year. Deciduous plants can also be categorised differently than their adaptation to drought or dry seasons, which can be temperate deciduous during cold seasons, and in contrast to evergreen plants which do not shed leaves annually, possessing green leaves throughout the year.

Biomass partitioning is the process by which plants divide their energy among their leaves, stems, roots, and reproductive parts. These four main components of the plant have important morphological roles: leaves take in CO2 and energy from the sun to create carbon compounds, stems grow above competitors to reach sunlight, roots absorb water and mineral nutrients from the soil while anchoring the plant, and reproductive parts facilitate the continuation of species. Plants partition biomass in response to limits or excesses in resources like sunlight, carbon dioxide, mineral nutrients, and water and growth is regulated by a constant balance between the partitioning of biomass between plant parts. An equilibrium between root and shoot growth occurs because roots need carbon compounds from photosynthesis in the shoot and shoots need nitrogen absorbed from the soil by roots. Allocation of biomass is put towards the limit to growth; a limit below ground will focus biomass to the roots and a limit above ground will favor more growth in the shoot.

Pit and mounds are small, persistent microtopographical features that present themselves after a disturbance event occurs and uproots trees via windthrow. The uprooted tree falls, and a pit forms in the forest floor where the root mass and associated soil matrix used to be. Eventually after a period of time in which the roots decay, the associated soil matrix that was pulled out of the ground with the roots falls back to the ground, creating a corresponding mound.

Biomass allocation is a concept in plant biology which indicates the relative proportion of plant biomass present in the different organs of a plant. It can also be used for whole plant communities.

References

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  2. Villar, Rafael; Ruiz-Robleto, Jeannete; Ubera, José Luis; Poorter, Hendrik (October 2013). "Exploring variation in leaf mass per area (LMA) from leaf to cell: An anatomical analysis of 26 woody species". American Journal of Botany. 100 (10): 1969–1980. doi:10.3732/ajb.1200562. PMID   24107583.
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  4. 1 2 Álvarez-Yépiz, Juan C.; Búrquez, Alberto; Martínez-Yrízar, Angelina; Teece, Mark; Yépez, Enrico A.; Dovciak, Martin (2017-02-01). "Resource partitioning by evergreen and deciduous species in a tropical dry forest". Oecologia. 183 (2): 607–618. Bibcode:2017Oecol.183..607A. doi:10.1007/s00442-016-3790-3. ISSN   1432-1939. PMID   27915413. S2CID   3798020. Archived from the original on 2022-09-13. Retrieved 2021-02-16.
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  6. Aerts, R. (1995). "The advantages of being evergreen" Archived 2015-09-24 at the Wayback Machine . Trends in Ecology & Evolution 10 (10): 402–407.
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