Pinus longaeva | |
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A tree in the Ancient Bristlecone Pine Forest, White Mountains, California | |
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Gymnospermae |
Division: | Pinophyta |
Class: | Pinopsida |
Order: | Pinales |
Family: | Pinaceae |
Genus: | Pinus |
Subgenus: | P. subg. Strobus |
Section: | P. sect. Parrya |
Subsection: | P. subsect. Balfourianae |
Species: | P. longaeva |
Binomial name | |
Pinus longaeva D.K.Bailey | |
Synonyms [3] | |
List
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Pinus longaeva (commonly referred to as the Great Basin bristlecone pine, intermountain bristlecone pine, or western bristlecone pine) [4] is a long-living species of bristlecone pine tree found in the higher mountains of California, Nevada, and Utah. [5] Methuselah is a bristlecone pine that is 4,856 years old and has been credited as the oldest known living non-clonal organism on Earth. [6] To protect it, the exact location of this tree is kept secret. In 1987, the bristlecone pine was designated one of Nevada's state trees. [7]
It is a medium-size tree, reaching 5 to 15 m (16 to 49 ft) tall and with a trunk diameter of up to 2.5 to 3.6 m (8 to 12 ft). The bark is bright orange-yellow, thin and scaly at the base of the trunk. The needles are in fascicles of five, stout, 2.5 to 4 cm (1 to 1+1⁄2 in) long, deep green to blue-green on the outer face, with stomata confined to a bright white band on the inner surfaces. The leaves show the longest persistence of any plant, with some remaining green for 45 years (Ewers & Schmid 1981).
The cones are ovoid-cylindrical, 5 to 10 cm (2 to 4 in) long and 3 to 4 cm (1 to 1+1⁄2 in) broad when closed, green or purple at first, ripening orange-buff when 16 months old, with numerous thin, fragile scales, each scale with a bristle-like spine 2 to 5 mm (1⁄16 to 3⁄16 in) long. The cones open to 4 to 6 cm (1+1⁄2 to 2+1⁄2 in) broad when mature, releasing the seeds immediately after opening. The seeds are 5 mm (3⁄16 in) long, with a 12 to 22 mm (1⁄2 to 7⁄8 in) wing; they are mostly dispersed by the wind, but some are also dispersed by Clark's nutcrackers.[ citation needed ]
These ancient trees have a gnarled and stunted appearance, especially those found at high altitudes, [4] and have reddish-brown bark with deep fissures. [8] As the tree ages, much of its vascular cambium layer may die. In very old specimens, often only a narrow strip of living tissue connects the roots to a handful of live branches.[ citation needed ]
The Great Basin bristlecone pine differs from the Rocky Mountain bristlecone pine in that the needles of the former always have two uninterrupted resin canals, so it lacks the characteristic small white resin flecks appearing on the needles of the latter. The Great Basin bristlecone pine differs from the foxtail pine because the cone bristles of the former are over 2 mm (0.079 in) long, and the cones have a more rounded (not conic) base. The green pine needles give the twisted branches a bottle-brush appearance. The name 'bristlecone pine' refers to the dark purple female cones that bear incurved prickles on their surface. [9]
The species occurs in Utah, Nevada and eastern California. In California, it is restricted to the White Mountains, the Inyo Mountains, and the Panamint Range, in Mono and Inyo counties. In Nevada, it is found in most of the higher ranges of the Basin and Range from the Spring Mountains near Las Vegas north to the Ruby Mountains, and in Utah, northeast to South Tent in the Wasatch Range. Due to the inaccessibility of many of the sites that this species occupies, information on their location and abundance is incomplete, and thus is needed. Environmental niche modelling has been used to better map the distribution of Great Basin bristlecone pine using topographic and spectral variables calculated from a geographic information system (GIS). [10]
The tree grows in large open stands, unlike the related foxtail pine, which sometimes form dense forests. Pinus longaeva trees generally do not form closed canopies, usually covering only 15-50%. [1] Pinus longaeva shares habitats with a number of other pine species, including the ponderosa pine, the white fir and, notably, the limber pine, a similarly long-lived high-elevation species. [4] The tree is a "vigorous" primary succession species, growing quickly on new open ground. [4] It is a "poor competitor" in good soils, however, and the species does best in harsh terrain. Pinus longaeva is often the dominant species in high-elevation dolomite soils, where few plants can grow. [4]
Bristlecone pines are protected in a number of areas owned by the United States federal government, such as the Ancient Bristlecone Pine Forest in the White Mountains of California and the Great Basin National Park in Nevada. [9] [11] These areas prohibit the cutting or gathering of wood. [11]
Clark's nutcrackers may play a role in seed distribution for P. longaeva, though direct observations of the birds foraging on these seeds have not been reported. [12] [13] The nutcrackers use conifer seeds as a food resource, storing many for later use in the ground, and some of these stored seeds are not used and are able to grow into new plants; these trees often exhibit a "multi-trunk" growth form from several seeds germinating at the same time. The prevalence of multi-trunk P. longaeva individuals in areas in which Clark's Nutcrackers are present has been used as evidence that the birds disperse P. longaeva seeds. [13]
An introduced fungal disease known as white pine blister rust ( Cronartium ribicola ) is believed to affect some individuals. The species was placed on the IUCN Red List and listed as "Vulnerable", or threatened, in 1998. [1] In 2011, however, a population survey found the population of Pinus longaeva to be stable, with no known subpopulations decreasing in size. White pine blister rust was found to have a negligible effect on the population. As a result, the species was moved to "Least Concern". [1]
The tree is extremely susceptible to fire, and is damaged by even low-intensity burns. The resinous bark is capable of igniting quickly, and a crown fire will almost certainly kill the tree. However, populations of Pinus longaeva are known to be extremely resilient, and as a primary succession species, it is believed that populations of the tree would reestablish itself quickly after a fire. That said, large-scale fires are extremely uncommon where the species grows, and are not a major factor in the species' long-term viability. [4] Historically, Pinus longaeva stands experienced low to high severity fires, and fuels structures changed considerably across elevational gradients. In low elevation, mixed species stands, fuels are often heavy and in close proximity to anthropogenic ignition sources. Yet at high elevations near treeline, Pinus longaeva typically grow on limestone outcroppings that provide little or no surface fuels to propagate a wildfire. However, warmer temperatures will likely increase the duration of fire season, and thus the frequency of fire in Pinus longeava systems at low and mid elevations could increase where stands are typically denser and surface fuel is greatest. While rare, wild fires such as The Carpenter 1 fire in southern Nevada (July 2013) and the Phillips Fire in Great Basin National Park, (September 2000) that started in lower elevation fuel types and moved through the crowns of trees with the aid of extreme fire weather, could become more likely. [14]
A specimen located in the White Mountains of California was measured by Tom Harlan, a researcher with the Laboratory of Tree-Ring Research, to be 5,062 years old as of 2010. [6] This would make it the oldest known non-clonal tree in the world. The identity of the specimen was kept secret by Harlan. [15] Harlan passed away in 2013, and neither the tree nor the core Harlan studied have been found, making the age or existence of this tree unable to be confirmed. [16]
The confirmed oldest tree of this species, "Methuselah", is also located in the Ancient Bristlecone Pine Forest of the White Mountains. Methuselah is 4,856 years old, as measured by annual ring count on a small core taken with an increment borer. Its exact location is kept secret.[ citation needed ] [17]
Among the White Mountain specimens, the oldest trees are found on north-facing slopes, with an average of 2,000 years, as compared to the 1,000 year average on the southern slopes. The climate and the durability of their wood can preserve them long after death, with dead trees as old as 7,000 years persisting next to live ones. [18]
A pine is any conifer tree or shrub in the genus Pinus of the family Pinaceae. Pinus is the sole genus in the subfamily Pinoideae.
The term bristlecone pine covers three species of pine tree. All three species are long-lived and highly resilient to harsh weather and bad soils. One of the three species, Pinus longaeva, is among the longest-lived life forms on Earth. The oldest of this species is more than 4,800 years old, making it the oldest known individual of any species. Many scientists are curious as to why this tree is able to live so long. In one study, they discovered that Pinus longaeva has higher levels of telomerase activity, which further slows or prevents the attrition rate of telomeres. This potentially contributes to the extended life of the bristlecone pine.
Clark's nutcracker, sometimes referred to as Clark's crow or woodpecker crow, is a passerine bird in the family Corvidae, native to the mountains of western North America. The nutcracker is an omnivore, but subsists mainly on pine nuts, burying seeds in the ground in the summer and then retrieving them in the winter by memory. The bird was described by the Lewis and Clark Expedition, with William Clark first observing it in 1805 along the banks of the Salmon River, a tributary of the Columbia River.
Pine nuts, also called piñón, pinoli, or pignoli, are the edible seeds of pines. According to the Food and Agriculture Organization, only 29 species provide edible nuts, while 20 are traded locally or internationally owing to their seed size being large enough to be worth harvesting; in other pines, the seeds are also edible but are too small to be of notable value as human food. The biggest producers of pine nuts are China, Russia, North Korea, Pakistan and Afghanistan.
The White Mountains of California and Nevada are a triangular fault-block mountain range facing the Sierra Nevada across the upper Owens Valley. They extend for approximately 60 mi (97 km) as a greatly elevated plateau about 20 mi (32 km) wide on the south, narrowing to a point at the north, with elevations generally increasing south to north. The range's broad southern end is near the community of Big Pine, where Westgard Pass and Deep Springs Valley separate it from the Inyo Mountains. The narrow northern end is at Montgomery Pass, where U.S. Route 6 crosses. The Fish Lake Valley lies east of the range; the southeast part of the mountains are separated from the Silver Peak Range by block faulting across the Furnace Creek Fault Zone, forming a feeder valley to Fish Lake Valley. The range lies within the eastern section of the Inyo National Forest.
The pinyon or piñon pine group grows in southwestern North America, especially in New Mexico, Colorado, Arizona, and Utah. The trees yield edible nuts, which are a staple food of Native Americans, and widely eaten as a snack and as an ingredient in New Mexican cuisine. The name comes from the Spanish pino piñonero, a name used for both the American varieties and the stone pine common in Spain, which also produces edible nuts typical of Mediterranean cuisine. Harvesting techniques of the prehistoric American Indians are still used today to collect the pinyon seeds for personal use or for commercialization. The pinyon nut or seed is high in fats and calories. In the western United States, pinyon pines are often found in pinyon–juniper woodlands.
Pinus albicaulis, known by the common names whitebark pine, white bark pine, white pine, pitch pine, scrub pine, and creeping pine, is a conifer tree native to the mountains of the western United States and Canada, specifically subalpine areas of the Sierra Nevada, Cascade Range, Pacific Coast Ranges, Rocky Mountains, and Ruby Mountains. It shares the common name "creeping pine" with several other plants.
Pinus balfouriana, the foxtail pine, is a rare high-elevation pine that is endemic to California, United States. It is closely related to the Great Basin and Rocky Mountain bristlecone pines, in the subsection Balfourianae.
Western white pine, also called silver pine and California mountain pine, is a species of pine in the family Pinaceae. It occurs in mountain ranges of northwestern North America and is the state tree of Idaho.
Pinus lambertiana is the tallest and most massive pine tree, and has the longest cones of any conifer. The species name lambertiana was given by the Scottish botanist David Douglas, who named the tree in honour of the English botanist, Aylmer Bourke Lambert. It is native to coastal and inland mountain areas along the Pacific coast of North America, as far north as Oregon and as far south as Baja California in Mexico.
Pinus flexilis, the limber pine, is a species of pine tree in the family Pinaceae that occurs in the mountains of the Western United States, Mexico, and Canada. It is also called Rocky Mountain white pine.
Pinus edulis, the Colorado pinyon, two-needle piñon, pinyon pine, or simply piñon, is a pine in the pinyon pine group native to the Southwestern United States, used for its edible pine nuts.
Pinus monophylla, the single-leaf pinyon, is a pine in the pinyon pine group, native to North America. The range is in southernmost Idaho, western Utah, Arizona, southwest New Mexico, Nevada, eastern and southern California and northern Baja California.
Pinus aristata, the Rocky Mountain bristlecone pine or Colorado bristlecone pine is a long-lived species of bristlecone pine tree native to the United States. It is found in the Rocky Mountains in Colorado and northern New Mexico, with an isolated population in the San Francisco Peaks of Arizona. It is found at very high altitudes, from 2,100 to 4,000 meters in cold, dry subalpine climate conditions, often at the tree line, although it also forms extensive closed-canopy stands at somewhat lower elevations.
Inyo National Forest is a United States National Forest covering parts of the eastern Sierra Nevada of California and the White Mountains of California and Nevada. The forest hosts several superlatives, including Mount Whitney, the highest point in the contiguous United States; Boundary Peak, the highest point in Nevada; and the Ancient Bristlecone Pine Forest, which protects the oldest living trees in the world. The forest, encompassing much of the Owens Valley, was established by Theodore Roosevelt as a way of sectioning off land to accommodate the Los Angeles Aqueduct project in 1907, making the Inyo National Forest one of the least wooded forests in the U.S. National Forest system.
The Ancient Bristlecone Pine Forest is a protected area high in the White Mountains in Inyo County in eastern California.
Prometheus was the oldest known non-clonal organism, a Great Basin bristlecone pine tree growing near the tree line on Wheeler Peak in eastern Nevada, United States. The tree, which was at least 4,862 years old and possibly more than 5,000, was cut down in 1964 by a graduate student and United States Forest Service personnel for research purposes. Those involved did not know of its world-record age before the cutting, and the circumstances and decision-making process remain controversial.
Methuselah is a 4,856-year-old Great Basin bristlecone pine tree growing high in the White Mountains of Inyo County in eastern California. It is recognized as the non-clonal tree with the greatest confirmed age in the world. The tree's name refers to the biblical patriarch Methuselah, who is said to have reached 969 years of age before his death, thus becoming synonymous with longevity or old age in many European languages including English.
The White Pine Range Wilderness is a 40,013-acre (16,193 ha) wilderness area in southwestern White Pine County, in the U.S. state of Nevada.
This article incorporates text from the ARKive fact-file "Pinus longaeva" under the Creative Commons Attribution-ShareAlike 3.0 Unported License and the GFDL .