Ground tissue

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Flax stem cross-section:
Pith
Protoxylem
Xylem I
Phloem I
Sclerenchyma (bast fibre)
Cortex
Epidermis Stem-histology-cross-section-tag.svg
Flax stem cross-section:

The ground tissue of plants includes all tissues that are neither dermal nor vascular. It can be divided into three types based on the nature of the cell walls. This tissue system is present between the dermal tissue and forms the main bulk of the plant body.

Contents

  1. Parenchyma cells have thin primary walls and usually remain alive after they become mature. Parenchyma forms the "filler" tissue in the soft parts of plants, and is usually present in cortex, pericycle, pith, and medullary rays in primary stem and root.
  2. Collenchyma cells have thin primary walls with some areas of secondary thickening. Collenchyma provides extra mechanical and structural support, particularly in regions of new growth.
  3. Sclerenchyma cells have thick lignified secondary walls and often die when mature. Sclerenchyma provides the main structural support to the plant. [1]

Parenchyma

Parenchyma is a versatile ground tissue that generally constitutes the "filler" tissue in soft parts of plants. It forms, among other things, the cortex (outer region) and pith (central region) of stems, the cortex of roots, the mesophyll of leaves, the pulp of fruits, and the endosperm of seeds. Parenchyma cells are often living cells and may remain meristematic, meaning that they are capable of cell division if stimulated. They have thin and flexible cellulose cell walls and are generally polyhedral when close-packed, but can be roughly spherical when isolated from their neighbors. Parenchyma cells are generally large. They have large central vacuoles, which allow the cells to store and regulate ions, waste products, and water. Tissue specialised for food storage is commonly formed of parenchyma cells.

Plant cell types.svg
Cross section of a leaf showing various ground tissue types

Parenchyma cells have a variety of functions:

The shape of parenchyma cells varies with their function. In the spongy mesophyll of a leaf, parenchyma cells range from near-spherical and loosely arranged with large intercellular spaces, [2] to branched or stellate, mutually interconnected with their neighbours at the ends of their arms to form a three-dimensional network, like in the red kidney bean Phaseolus vulgaris and other mesophytes. [3] These cells, along with the epidermal guard cells of the stoma, form a system of air spaces and chambers that regulate the exchange of gases. In some works, the cells of the leaf epidermis are regarded as specialised parenchymal cells, [4] but the modern preference has long been to classify the epidermis as plant dermal tissue, and parenchyma as ground tissue. [5]

Shapes of parenchyma:

Collenchyma

Cross section of collenchyma cells Plant cell type collenchyma.png
Cross section of collenchyma cells

Collenchyma tissue is composed of elongated cells with irregularly thickened walls. They provide structural support, particularly in growing shoots and leaves (as seen, for example, the resilient strands in stalks of celery). Collenchyma cells are usually living, and have only a thick primary cell wall [6] made up of cellulose and pectin. Cell wall thickness is strongly affected by mechanical stress upon the plant. The walls of collenchyma in shaken plants (to mimic the effects of wind etc.), may be 40–100% thicker than those not shaken.

There are four main types of collenchyma:

Collenchyma cells are most often found adjacent to outer growing tissues such as the vascular cambium and are known for increasing structural support and integrity.

The first use of "collenchyma" ( /kəˈlɛŋkɪmə,kɒ-/ [7] [8] ) was by Link (1837) who used it to describe the sticky substance on Bletia (Orchidaceae) pollen. Complaining about Link's excessive nomenclature, Schleiden (1839) stated mockingly that the term "collenchyma" could have more easily been used to describe elongated sub-epidermal cells with unevenly thickened cell walls. [9]

Sclerenchyma

Sclerenchyma is the tissue which makes the plant hard and stiff. Sclerenchyma is the supporting tissue in plants. Two types of sclerenchyma cells exist: fibers cellular and sclereids. Their cell walls consist of cellulose, hemicellulose, and lignin. Sclerenchyma cells are the principal supporting cells in plant tissues that have ceased elongation. Sclerenchyma fibers are of great economic importance, since they constitute the source material for many fabrics (e.g. flax, hemp, jute, and ramie).

Unlike the collenchyma, mature sclerenchyma is composed of dead cells with extremely thick cell walls (secondary walls) that make up to 90% of the whole cell volume. The term sclerenchyma is derived from the Greek σκληρός (sklērós), meaning "hard." It is the hard, thick walls that make sclerenchyma cells important strengthening and supporting elements in plant parts that have ceased elongation. The difference between sclereids is not always clear: transitions do exist, sometimes even within the same plant.

Fibers

Cross section of sclerenchyma fibers Plant cell type sclerenchyma fibers.png
Cross section of sclerenchyma fibers

Fibers or bast are generally long, slender, so-called prosenchymatous cells, usually occurring in strands or bundles. Such bundles or the totality of a stem's bundles are colloquially called fibers. Their high load-bearing capacity and the ease with which they can be processed has since antiquity made them the source material for a number of things, like ropes, fabrics and mattresses. The fibers of flax (Linum usitatissimum) have been known in Europe and Egypt for more than 3,000 years, those of hemp (Cannabis sativa) in China for just as long. These fibers, and those of jute (Corchorus capsularis) and ramie (Boehmeria nivea, a nettle), are extremely soft and elastic and are especially well suited for the processing to textiles. Their principal cell wall material is cellulose.

Contrasting are hard fibers that are mostly found in monocots. Typical examples are the fiber of many grasses, Agave sisalana (sisal), Yucca or Phormium tenax , Musa textilis and others. Their cell walls contain, besides cellulose, a high proportion of lignin. The load-bearing capacity of Phormium tenax is as high as 20–25 kg/mm², the same as that of good steel wire (25 kg/ mm²), but the fibre tears as soon as too great a strain is placed upon it, while the wire distorts and does not tear before a strain of 80 kg/mm². The thickening of a cell wall has been studied in Linum .[ citation needed ] Starting at the centre of the fiber, the thickening layers of the secondary wall are deposited one after the other. Growth at both tips of the cell leads to simultaneous elongation. During development the layers of secondary material seem like tubes, of which the outer one is always longer and older than the next. After completion of growth, the missing parts are supplemented, so that the wall is evenly thickened up to the tips of the fibers.

Fibers usually originate from meristematic tissues. Cambium and procambium are their main centers of production. They are usually associated with the xylem and phloem of the vascular bundles. The fibers of the xylem are always lignified, while those of the phloem are cellulosic. Reliable evidence for the fibre cells' evolutionary origin from tracheids exists.[ citation needed ] During evolution the strength of the tracheid cell walls was enhanced, the ability to conduct water was lost and the size of the pits was reduced. Fibers that do not belong to the xylem are bast (outside the ring of cambium) and such fibers that are arranged in characteristic patterns at different sites of the shoot. The term "sclerenchyma" (originally Sclerenchyma) was introduced by Mettenius in 1865. [10]

Sclereids

Fresh mount of a sclereid Plant cell type sclerenchyma sclereid.png
Fresh mount of a sclereid
Long, tapered sclereids supporting a leaf edge in Dionysia kossinskyi Dionysia kossinskyi sclereids.TIF
Long, tapered sclereids supporting a leaf edge in Dionysia kossinskyi

Sclereids are the reduced form of sclerenchyma cells with highly thickened, lignified walls.

They are small bundles of sclerenchyma tissue in plants that form durable layers, such as the cores of apples and the gritty texture of pears (Pyrus communis). Sclereids are variable in shape. The cells can be isodiametric, prosenchymatic, forked or elaborately branched. They can be grouped into bundles, can form complete tubes located at the periphery or can occur as single cells or small groups of cells within parenchyma tissues. But compared with most fibres, sclereids are relatively short. Characteristic examples are brachysclereids or the stone cells (called stone cells because of their hardness) of pears and quinces (Cydonia oblonga) and those of the shoot of the wax plant (Hoya carnosa). The cell walls fill nearly all the cell's volume. A layering of the walls and the existence of branched pits is clearly visible. Branched pits such as these are called ramiform pits. The shell of many seeds like those of nuts as well as the stones of drupes like cherries and plums are made up from sclereids.

These structures are used to protect other cells.

Related Research Articles

<span class="mw-page-title-main">Plant cell</span> Type of eukaryotic cell present in green plants

Plant cells are the cells present in green plants, photosynthetic eukaryotes of the kingdom Plantae. Their distinctive features include primary cell walls containing cellulose, hemicelluloses and pectin, the presence of plastids with the capability to perform photosynthesis and store starch, a large vacuole that regulates turgor pressure, the absence of flagella or centrioles, except in the gametes, and a unique method of cell division involving the formation of a cell plate or phragmoplast that separates the new daughter cells.

<span class="mw-page-title-main">Xylem</span> Water transport tissue in vascular plants

Xylem is one of the two types of transport tissue in vascular plants, the other being phloem. The basic function of the xylem is to transport water from roots to stems and leaves, but it also transports nutrients. The word xylem is derived from the Ancient Greek word ξύλον (xylon), meaning "wood"; the best-known xylem tissue is wood, though it is found throughout a plant. The term was introduced by Carl Nägeli in 1858.

<span class="mw-page-title-main">Phloem</span> Sugar transport tissue in vascular plants

Phloem is the living tissue in vascular plants that transports the soluble organic compounds made during photosynthesis and known as photosynthates, in particular the sugar sucrose, to the rest of the plant. This transport process is called translocation. In trees, the phloem is the innermost layer of the bark, hence the name, derived from the Ancient Greek word φλοιός (phloiós), meaning "bark". The term was introduced by Carl Nägeli in 1858. Different types of phloem can be distinguished. The early phloem formed in the growth apices is called protophloem. Protophloem eventually becomes obliterated once it connects to the durable phloem in mature organs, the metaphloem. Further, secondary phloem is formed during the thickening of stem structures.

<span class="mw-page-title-main">Tissue (biology)</span> Group of similar cells performing a specific function

In biology, tissue is an assembly of similar cells and their extracellular matrix from the same embryonic origin that together carry out a specific function. Tissues occupy a biological organizational level between cells and a complete organ. Accordingly, organs are formed by the functional grouping together of multiple tissues.

<span class="mw-page-title-main">Bark (botany)</span> Outermost layers of stems and roots of woody plants

Bark is the outermost layer of stems and roots of woody plants. Plants with bark include trees, woody vines, and shrubs. Bark refers to all the tissues outside the vascular cambium and is a nontechnical term. It overlays the wood and consists of the inner bark and the outer bark. The inner bark, which in older stems is living tissue, includes the innermost layer of the periderm. The outer bark on older stems includes the dead tissue on the surface of the stems, along with parts of the outermost periderm and all the tissues on the outer side of the periderm. The outer bark on trees which lies external to the living periderm is also called the rhytidome.

<span class="mw-page-title-main">Vascular cambium</span> Main growth tissue in the stems, roots of plants

The vascular cambium is the main growth tissue in the stems and roots of many plants, specifically in dicots such as buttercups and oak trees, gymnosperms such as pine trees, as well as in certain other vascular plants. It produces secondary xylem inwards, towards the pith, and secondary phloem outwards, towards the bark.

<span class="mw-page-title-main">Cork cambium</span> Part of a plant

Cork cambium is a tissue found in many vascular plants as a part of the epidermis. It is one of the many layers of bark, between the cork and primary phloem. The cork cambium is a lateral meristem and is responsible for secondary growth that replaces the epidermis in roots and stems. It is found in woody and many herbaceous dicots, gymnosperms and some monocots. It is one of the plant's meristems – the series of tissues consisting of embryonic disk cells from which the plant grows. The function of cork cambium is to produce the cork, a tough protective material.

<span class="mw-page-title-main">Meristem</span> Type of plant tissue involved in cell proliferation

The meristem is a type of tissue found in plants. It consists of undifferentiated cells capable of cell division. Cells in the meristem can develop into all the other tissues and organs that occur in plants. These cells continue to divide until a time when they get differentiated and then lose the ability to divide.

<span class="mw-page-title-main">Sclereid</span> Plant tissue type

Sclereids are a reduced form of sclerenchyma cells with highly thickened, lignified cellular walls that form small bundles of durable layers of tissue in most plants. The presence of numerous sclereids form the cores of apples and produce the gritty texture of guavas.

The pericycle is a cylinder of parenchyma or sclerenchyma cells that lies just inside the endodermis and is the outer most part of the stele of plants.

<span class="mw-page-title-main">Cortex (botany)</span> Outer layer of a stem or root in a vascular plant

In botany, a cortex is an outer layer of a stem or root in a vascular plant, lying below the epidermis but outside of the vascular bundles. The cortex is composed mostly of large thin-walled parenchyma cells of the ground tissue system and shows little to no structural differentiation. The outer cortical cells often acquire irregularly thickened cell walls, and are called collenchyma cells.

<span class="mw-page-title-main">Palisade cell</span> Type of cell found in plant leaves

Palisade cells are plant cells located on the leaves, right below the epidermis and cuticle that is the outermost layer of the leaf. In simpler terms, they are known as leaf cells. Palisade means "stake" in Latin, they are vertically elongated and are stacked side by side, a different shape from the spongy mesophyll cells beneath them. The chloroplasts in these cells absorb a major portion of the light energy used by the leaf. Palisade cells occur in dicotyledonous plants, and also in the net-veined Monocots, the Araceae and Dioscoreaceae.

<span class="mw-page-title-main">Vascular bundle</span> Longitudinal strand of vascular tissue in the roots, stems and leaves of higher plants

A vascular bundle is a part of the transport system in vascular plants. The transport itself happens in the stem, which exists in two forms: xylem and phloem. Both these tissues are present in a vascular bundle, which in addition will include supporting and protective tissues. In addition, there is also a tissue between xylem and phloem which is the cambium.

<span class="mw-page-title-main">Epidermis (botany)</span> Layer of cells that covers leaves, flowers, roots of plants

The epidermis is a single layer of cells that covers the leaves, flowers, roots and stems of plants. It forms a boundary between the plant and the external environment. The epidermis serves several functions: it protects against water loss, regulates gas exchange, secretes metabolic compounds, and absorbs water and mineral nutrients. The epidermis of most leaves shows dorsoventral anatomy: the upper (adaxial) and lower (abaxial) surfaces have somewhat different construction and may serve different functions. Woody stems and some other stem structures such as potato tubers produce a secondary covering called the periderm that replaces the epidermis as the protective covering.

<span class="mw-page-title-main">Vascular tissue</span> Conducting tissue in vascular plants

Vascular tissue is a complex conducting tissue, formed of more than one cell type, found in vascular plants. The primary components of vascular tissue are the xylem and phloem. These two tissues transport fluid and nutrients internally. There are also two meristems associated with vascular tissue: the vascular cambium and the cork cambium. All the vascular tissues within a particular plant together constitute the vascular tissue system of that plant.

<span class="mw-page-title-main">Secondary growth</span> Type of growth in plants

In botany, secondary growth is the growth that results from cell division in the cambia or lateral meristems and that causes the stems and roots to thicken, while primary growth is growth that occurs as a result of cell division at the tips of stems and roots, causing them to elongate, and gives rise to primary tissue. Secondary growth occurs in most seed plants, but monocots usually lack secondary growth. If they do have secondary growth, it differs from the typical pattern of other seed plants.

<span class="mw-page-title-main">Lepidodendrales</span> Extinct order of vascular tree-like plants

Lepidodendrales or arborescent lycophytes are an extinct order of primitive, vascular, heterosporous, arborescent (tree-like) plants belonging to Lycopodiopsida. Members of Lepidodendrales are the best understood of the fossil lycopsids due to the vast diversity of Lepidodendrales specimens and the diversity in which they were preserved; the extensive distribution of Lepidodendrales specimens as well as their well-preservedness lends paleobotanists exceptionally detailed knowledge of the coal-swamp giants’ reproductive biology, vegetative development, and role in their paleoecosystem. The defining characteristics of the Lepidodendrales are their secondary xylem, extensive periderm development, three-zoned cortex, rootlike appendages known as stigmarian rootlets arranged in a spiralling pattern, and megasporangium each containing a single functional megaspore that germinates inside the sporangium. Many of these different plant organs have been assigned both generic and specific names as relatively few have been found organically attached to each other. Some specimens have been discovered which indicate heights of 40 and even 50 meters and diameters of over 2 meters at the base. The massive trunks of some species branched profusely, producing large crowns of leafy twigs; though some leaves were up to 1 meter long, most were much shorter, and when leaves dropped from branches their conspicuous leaf bases remained on the surface of branches. Strobili could be found at the tips of distal branches or in an area at the top of the main trunk. The underground organs of Lepidodendrales typically consisted of dichotomizing axes bearing helically arranged, lateral appendages serving an equivalent function to roots. Sometimes called "giant club mosses", they are believed to be more closely related to extant quillworts based on xylem, although fossil specimens of extinct Selaginellales from the Late Carboniferous also had secondary xylem.

<span class="mw-page-title-main">Woody plant</span> Plant that produces wood and has a hard stem

A woody plant is a plant that produces wood as its structural tissue and thus has a hard stem. In cold climates, woody plants further survive winter or dry season above ground, as opposed to herbaceous plants that die back to the ground until spring.

<span class="mw-page-title-main">Plant stem</span> Structural axis of a vascular plant

A stem is one of two main structural axes of a vascular plant, the other being the root. It supports leaves, flowers and fruits, transports water and dissolved substances between the roots and the shoots in the xylem and phloem, photosynthesis takes place here, stores nutrients, and produces new living tissue. The stem can also be called halm or haulm or culms.

<span class="mw-page-title-main">Leaf</span> Photosynthetic part of a vascular plant

A leaf is a principal appendage of the stem of a vascular plant, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, flower, and fruit collectively form the shoot system. In most leaves, the primary photosynthetic tissue is the palisade mesophyll and is located on the upper side of the blade or lamina of the leaf but in some species, including the mature foliage of Eucalyptus, palisade mesophyll is present on both sides and the leaves are said to be isobilateral. Most leaves are flattened and have distinct upper (adaxial) and lower (abaxial) surfaces that differ in color, hairiness, the number of stomata, the amount and structure of epicuticular wax and other features. Leaves are mostly green in color due to the presence of a compound called chlorophyll which is essential for photosynthesis as it absorbs light energy from the sun. A leaf with lighter-colored or white patches or edges is called a variegated leaf.

References

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Further reading