A cultivar is a type of plant that people have bred for desired traits, which are reproduced in each new generation by a method such as grafting, tissue culture, or carefully controlled seed production. Most cultivars arise from purposeful human manipulation, but some originate from wild plants that have distinctive characteristics. Cultivar names are chosen according to rules of the International Code of Nomenclature for Cultivated Plants (ICNCP), and not all cultivated plants qualify as cultivars. Horticulturists generally believe the word cultivarwas coined as a term meaning "cultivated variety".
Popular ornamental garden plants like roses, camellias, daffodils, rhododendrons, and azaleas are commonly cultivars produced by breeding and selection or as sports, for floral colour or size, plant form, or other desirable characteristics.Similarly, the world's agricultural food crops are almost exclusively cultivars that have been selected for characters such as improved yield, flavour, and resistance to disease, and very few wild plants are now used as food sources. Trees used in forestry are also special selections grown for their enhanced quality and yield of timber.
Cultivars form a major part of Liberty Hyde Bailey's broader group, the cultigen,which is defined as a plant whose origin or selection is primarily due to intentional human activity. A cultivar is not the same as a botanical variety, which is a taxonomic rank below subspecies, and there are differences in the rules for creating and using the names of botanical varieties and cultivars. In recent times, the naming of cultivars has been complicated by the use of statutory patents for plants and recognition of plant breeders' rights.
The International Union for the Protection of New Varieties of Plants (UPOV – French : Union internationale pour la protection des obtentions végétales) offers legal protection of plant cultivars to persons or organisations that introduce new cultivars to commerce. UPOV requires that a cultivar be "distinct, uniform", and "stable". To be "distinct", it must have characters that easily distinguish it from any other known cultivar. To be "uniform" and "stable", the cultivar must retain these characters in repeated propagation.
The naming of cultivars is an important aspect of cultivated plant taxonomy, and the correct naming of a cultivar is prescribed by the Rules and Recommendations of the International Code of Nomenclature for Cultivated Plants (ICNCP, commonly denominated the Cultivated Plant Code). A cultivar is given a cultivar name, which consists of the scientific Latin botanical name followed by a cultivar epithet. The cultivar epithet is usually in a vernacular language.
The word cultivar originated from the need to distinguish between wild plants and those with characteristics that arose in cultivation, presently denominated cultigens. This distinction dates to the Greek philosopher Theophrastus (370–285 BC), the "Father of Botany", who was keenly aware of this difference. Botanical historian Alan Morton noted that Theophrastus in his Historia Plantarum (Enquiry into Plants) "had an inkling of the limits of culturally induced (phenotypic) changes and of the importance of genetic constitution" (Historia Plantarum, Book 3, 2, 2 and Causa Plantarum, Book 1, 9, 3).
The International Code of Nomenclature for algae, fungi, and plants uses as its starting point for modern botanical nomenclature the Latin names in Linnaeus' (1707–1778) Species Plantarum (tenth edition) and Genera Plantarum (fifth edition). In Species Plantarum, Linnaeus enumerated all plants known to him, either directly or from his extensive reading. He recognised the rank of varietas (botanical "variety", a rank below that of species and subspecies) and he indicated these varieties with letters of the Greek alphabet, such as α, β, and λ, before the varietal name, rather than using the abbreviation "var." as is the present convention. Most of the varieties that Linnaeus enumerated were of "garden" origin rather than being wild plants.
In time the need to distinguish between wild plants and those with variations that had been cultivated increased. In the nineteenth century many "garden-derived" plants were given horticultural names, sometimes in Latin and sometimes in a vernacular language. From circa the 1900s, cultivated plants in Europe were recognised in the Scandinavian, Germanic, and Slavic literature as stamm or sorte,but these words could not be used internationally because, by international agreement, any new denominations had to be in Latin. In the twentieth century an improved international nomenclature was proposed for cultivated plants.
Liberty Hyde Bailey of Cornell University in New York, United States created the word cultivar in 1923 when he wrote that:
The cultigen is a species, or its equivalent, that has appeared under domestication – the plant is cultigenous. I now propose another name, cultivar, for a botanical variety, or for a race subordinate to species, that has originated under cultivation; it is not necessarily, however, referable to a recognized botanical species. It is essentially the equivalent of the botanical variety except in respect to its origin.
In that essay, Bailey used only the rank of species for the cultigen, but it was obvious to him that many domesticated plants were more like botanical varieties than species, and that realization appears to have motivated the suggestion of the new category of cultivar.
Bailey created the word cultivar. It is generally assumed to be a blend of cultivated and variety but Bailey never explicitly stated the etymology and it has been suggested that the word is actually a blend of cultigen and variety.The neologism cultivar was promoted as "euphonious" and "free from ambiguity". The first Cultivated Plant Code of 1953 subsequently commended its use, and by 1960 it had achieved common international acceptance.
The words cultigen and cultivar may be confused with each other. A cultigen is any plant that is deliberately selected for or altered in cultivation, as opposed to an indigen ; the Cultivated Plant Code states that cultigens are "maintained as recognisable entities solely by continued propagation".Cultigens can have names at any of many taxonomic ranks, including those of grex, species, cultivar group, variety, form, and cultivar; and they may be plants that have been altered in cultivation, including by genetic modification, but have not been formally denominated. A cultigen or a component of a cultigen can be accepted as a cultivar if it is recognisable and has stable characters. Therefore, all cultivars are cultigens, because they are cultivated, but not all cultigens are cultivars, because some cultigens have not been formally distinguished and named as cultivars.
The Cultivated Plant Code notes that the word cultivar is used in two different senses: first, as a "classification category" the cultivar is defined in Article 2 of the International Code of Nomenclature for Cultivated Plants (2009, 8th edition) as follows: The basic category of cultivated plants whose nomenclature is governed by this Code is the cultivar.There are two other classification categories for cultigens, the grex and the group. The Code then defines a cultivar as a "taxonomic unit within the classification category of cultivar". This is the sense of cultivar that is most generally understood and which is used as a general definition.
A cultivar is an assemblage of plants that (a) has been selected for a particular character or combination of characters, (b) is distinct, uniform and stable in those characters, and (c) when propagated by appropriate means, retains those characters.
Which plants are chosen to be named as cultivars is simply a matter of convenience as the category was created to serve the practical needs of horticulture, agriculture, and forestry.
Members of a particular cultivar are not necessarily genetically identical. The Cultivated Plant Code emphasizes that different cultivated plants may be accepted as different cultivars, even if they have the same genome, while cultivated plants with different genomes may be regarded as the same cultivar. The production of cultivars generally entails considerable human involvement although in a few cases it may be as little as simply selecting variation from plants growing in the wild (whether by collecting growing tissue to propagate from or by gathering seed).
Cultivars generally occur as ornamentals and food crops: Malus 'Granny Smith' and Malus 'Red Delicious' are cultivars of apples propagated by cuttings or grafting, Lactuca 'Red Sails' and Lactuca 'Great Lakes' are lettuce cultivars propagated by seeds. Named cultivars of Hosta and Hemerocallis plants are cultivars produced by micropropagation or division.
Cultivars that are produced asexually are genetically identical and known as clones; this includes plants propagated by division, layering, cuttings, grafts, and budding. The propagating material may be taken from a particular part of the plant, such as a lateral branch, or from a particular phase of the life cycle, such as a juvenile leaf, or from aberrant growth as occurs with witch's broom. Plants whose distinctive characters are derived from the presence of an intracellular organism may also form a cultivar provided the characters are reproduced reliably from generation to generation. Plants of the same chimera (which have mutant tissues close to normal tissue) or graft-chimeras (which have vegetative tissue from different kinds of plants and which originate by grafting) may also constitute a cultivar.
Some cultivars "come true from seed", retaining their distinguishing characteristics when grown from seed. Such plants are termed a "variety", "selection" or "strain" but these are ambiguous and confusing words that are best avoided. In general, asexually propagated cultivars grown from seeds produce highly variable seedling plants, and should not be labelled with, or sold under, the parent cultivar's name.
Seed-raised cultivars may be produced by uncontrolled pollination when characteristics that are distinct, uniform and stable are passed from parents to progeny. Some are produced as "lines" that are produced by repeated self-fertilization or inbreeding or "multilines" that are made up of several closely related lines. Sometimes they are F1 hybrids which are the result of a deliberate repeatable single cross between two pure lines. A few F2 hybrid seed cultivars also exist, such as Achillea 'Summer Berries'.
Some cultivars are agamospermous plants, which retain their genetic composition and characteristics under reproduction.Occasionally cultivars are raised from seed of a specially selected provenance – for example the seed may be taken from plants that are resistant to a particular disease.
Genetically modified plants with characteristics resulting from the deliberate implantation of genetic material from a different germplasm may form a cultivar. However, the International Code of Nomenclature for Cultivated Plants notes, "In practice such an assemblage is often marketed from one or more lines or multilines that have been genetically modified. These lines or multilines often remain in a constant state of development which makes the naming of such an assemblage as a cultivar a futile exercise."However, retired transgenic varieties such as the fish tomato, which are no longer being developed, do not run into this obstacle and can be given a cultivar name.
Cultivars may be selected because of a change in the ploidy level of a plant which may produce more desirable characteristics.
Every unique cultivar has a unique name within its denomination class (which is almost always the genus). Names of cultivars are regulated by the International Code of Nomenclature for Cultivated Plants, and may be registered with an International Cultivar Registration Authority (ICRA). There are sometimes separate registration authorities for different plant types such as roses and camellias. In addition, cultivars may be associated with commercial marketing names referred to in the Cultivated Plant Code as "trade designations" (see below).
A cultivar name consists of a botanical name (of a genus, species, infraspecific taxon, interspecific hybrid or intergeneric hybrid) followed by a cultivar epithet. The cultivar epithet is enclosed by single quotes;it should not be italicized if the botanical name is italicized; and each of the words within the epithet is capitalized (with some permitted exceptions such as conjunctions). It is permissible to place a cultivar epithet after a common name provided the common name is botanically unambiguous. Cultivar epithets published before 1 January 1959 were often given a Latin form and can be readily confused with the specific epithets in botanical names; after that date, newly coined cultivar epithets must be in a modern vernacular language to distinguish them from botanical epithets.
For example, the full cultivar name of the King Edward potato is Solanum tuberosum 'King Edward'. 'King Edward' is the cultivar epithet, which, according to the Rules of the Cultivated Plant Code, is bounded by single quotation marks.For patented or trademarked plant product lines developed from a given cultivar, the commercial product name is typically indicated by the symbols "TM" or "®", or is presented in capital letters with no quotation marks, following the cultivar name, as in the following example, where "Bloomerang" is the commercial name and 'Penda' is the cultivar epithet: Syringa 'Penda' BLOOMERANG.
Although "cv." has not been permitted by the International Code of Nomenclature for Cultivated Plants since the 1995 editionit is still widely used and recommended by other authorities.
Where several very similar cultivars exist they can be associated into a Group (formerly Cultivar-group). As Group names are used with cultivar names it is necessary to understand their way of presentation. Group names are presented in normal type and the first letter of each word capitalised as for cultivars, but they are not placed in single quotes. When used in a name, the first letter of the word "Group" is itself capitalized.
Property and Property law
Since the 1990s there has been an increasing use of legal protection for newly produced cultivars. Plant breeders expect legal protection for the cultivars they produce. According to proponents of such protections, if other growers can immediately propagate and sell these cultivars as soon as they come on the market, the breeder's benefit is largely lost.Legal protection for cultivars is obtained through the use of Plant breeders’ rights and plant Patents but the specific legislation and procedures needed to take advantage of this protection vary from country to country.
The use of legal protection for cultivars can be controversial, particularly for food crops that are staples in developing countries,or for plants selected from the wild and propagated for sale without any additional breeding work; some people consider this practice unethical.
The formal scientific name of a cultivar, like Solanum tuberosum ‘King Edward’, is a way of uniquely designating a particular kind of plant. This scientific name is in the public domain and cannot be legally protected. Plant retailers wish to maximize their share of the market and one way of doing this is to replace the cumbersome Latin scientific names on plant labels in retail outlets with appealing marketing names that are easy to use, pronounce, and remember. Marketing names lie outside the scope of the Cultivated Plant Code which refers to them as "trade designations". If a retailer or wholesaler has the sole legal rights to a marketing name then that may offer a sales advantage. Plants protected by plant breeders' rights (PBR) may have a "true" cultivar name – the recognized scientific name in the public domain – and a "commercial synonym" – an additional marketing name that is legally protected. An example would be RosaFascination = 'Poulmax', in which Rosa is the genus, Fascination is the trade designation, and ‘Poulmax’ is scientific cultivar name.
Because a name that is attractive in one language may have less appeal in another country, a plant may be given different selling names from country to country. Quoting the original cultivar name allows the correct identification of cultivars around the world.
The main body coordinating plant breeders' rights is the International Union for the Protection of New Varieties of Plants (Union internationale pour la protection des obtentions végétales, UPOV) and this organization maintains a database of new cultivars protected by PBR in all countries.
An International Cultivar Registration Authority (ICRA) is a voluntary, non-statutory organization appointed by the Commission for Nomenclature and Cultivar Registration of the International Society of Horticultural Science. ICRAs are generally formed by societies and institutions specializing in particular plant genera such as Dahlia or Rhododendron and are currently located in Europe, North America, China, India, Singapore, Australia, New Zealand, South Africa and Puerto Rico.
Each ICRA produces an annual report and its reappointment is considered every four years. The main task is to maintain a register of the names within the group of interest and where possible this is published and placed in the public domain. One major aim is to prevent the duplication of cultivar and Group epithets within a genus, as well as ensuring that names are in accord with the latest edition of the Cultivated Plant Code. In this way, over the last 50 years or so, ICRAs have contributed to the stability of cultivated plant nomenclature. In recent times many ICRAs have also recorded trade designations and trademarks used in labelling plant material, to avoid confusion with established names.
New names and other relevant data are collected by and submitted to the ICRA and in most cases there is no cost. The ICRA then checks each new epithet to ensure that it has not been used before and that it conforms with the Cultivated Plant Code. Each ICRA also ensures that new names are formally established (i.e. published in hard copy, with a description in a dated publication). They record details about the plant, such as parentage, the names of those concerned with its development and introduction, and a basic description highlighting its distinctive characters. ICRAs are not responsible for assessing the distinctiveness of the plant in question.Most ICRAs can be contacted electronically and many maintain web sites for an up-to-date listing.
In botanical nomenclature, variety is a taxonomic rank below that of species and subspecies, but above that of form. As such, it gets a three-part infraspecific name. It is sometimes recommended that the subspecies rank should be used to recognize geographic distinctiveness, whereas the variety rank is appropriate if the taxon is seen throughout the geographic range of the species.
In botany and plant taxonomy, a series is a subdivision of a genus, a taxonomic rank below that of section but above that of species.
A botanical name is a formal scientific name conforming to the International Code of Nomenclature for algae, fungi, and plants (ICN) and, if it concerns a plant cultigen, the additional cultivar or Group epithets must conform to the International Code of Nomenclature for Cultivated Plants (ICNCP). The code of nomenclature covers "all organisms traditionally treated as algae, fungi, or plants, whether fossil or non-fossil, including blue-green algae (Cyanobacteria), chytrids, oomycetes, slime moulds and photosynthetic protists with their taxonomically related non-photosynthetic groups ."
In taxonomy, a nomen nudum is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be a scientific name, but fails to be one because it has not been published with an adequate description. This makes it a "bare" or "naked" name, one which cannot be accepted as it stands. A largely equivalent but much less frequently used term is nomen tantum.
Botanical nomenclature is the formal, scientific naming of plants. It is related to, but distinct from taxonomy. Plant taxonomy is concerned with grouping and classifying plants; botanical nomenclature then provides names for the results of this process. The starting point for modern botanical nomenclature is Linnaeus' Species Plantarum of 1753. Botanical nomenclature is governed by the International Code of Nomenclature for algae, fungi, and plants (ICN), which replaces the International Code of Botanical Nomenclature (ICBN). Fossil plants are also covered by the code of nomenclature.
In botany, an infraspecific name is the scientific name for any taxon below the rank of species, i.e. an infraspecific taxon or infraspecies. The scientific names of botanical taxa are regulated by the International Code of Nomenclature for algae, fungi, and plants (ICN). This specifies a 'three part name' for infraspecific taxa, plus a 'connecting term' to indicate the rank of the name. An example of such a name is Astrophytum myriostigma subvar. glabrum, the name of a subvariety of the species Astrophytum myriostigma.
The International Code of Nomenclature for Cultivated Plants (ICNCP), also known as the Cultivated Plant Code, is a guide to the rules and regulations for naming cultigens, plants whose origin or selection is primarily due to intentional human activity. Cultigens under the purview of the ICNCP include cultivars, Groups, and grexes. All organisms traditionally considered to be plants are included. Taxa that receive a name under the ICNCP will also be included within taxa named under the International Code of Nomenclature for algae, fungi, and plants, for example, a cultivar is a member of a species.
A Group is a formal category in the International Code of Nomenclature for Cultivated Plants (ICNCP) used for cultivated plants (cultivars) that share a defined characteristic. It is represented in a botanical name by the symbol Group or Gp. "Group" or "Gp" is always written with a capital G in a botanical name, or epithet. The Group is not italicized in a plant's name. The ICNCP introduced the term and symbol "Group" in 2004, as a replacement for the lengthy and hyphenated "cultivar-group", which had previously been the category's name since 1969. For the old name "cultivar-group", the non-standard abbreviation cv. group or cv. Group is also sometimes encountered. There is a slight difference in meaning, since a cultivar-group was defined to comprise cultivars, whereas a Group may include individual plants.
Plant variety is a legal term, following the International Union for the Protection of New Varieties of Plants (UPOV) Convention. Recognition of a cultivated plant as a "variety" in this particular sense provides its breeder with some legal protection, so-called plant breeders' rights, depending to some extent on the internal legislation of the UPOV signatory countries, such as the Plant Variety Protection Act in the US.
In botanical nomenclature, a hybrid may be given a hybrid name, which is a special kind of botanical name, but there is no requirement that a hybrid name should be created for plants that are believed to be of hybrid origin. The International Code of Nomenclature for algae, fungi, and plants (ICNafp) provides the following options in dealing with a hybrid:
A cultigen or cultivated plant is a plant that has been deliberately altered or selected by humans; it is the result of artificial selection. These plants, for the most part, have commercial value in horticulture, agriculture or forestry. Because cultigens are defined by their mode of origin and not by where they are growing, plants meeting this definition remain cultigens whether they are naturalised in the wild, deliberately planted in the wild, or growing in cultivation.
Isopogon anemonifolius, commonly known as broad-leaved drumsticks, is a shrub of the family Proteaceae that is native only to eastern New South Wales in Australia. It occurs naturally in woodland, open forest, and heathland on sandstone soils. I. anemonifolius usually ranges between one and two metres in height, and is generally smaller in exposed heathland. Its leaves are divided and narrow, though broader than those of the related Isopogon anethifolius, and have a purplish tinge during the cooler months. The yellow flowers appear during late spring or early summer and are displayed prominently. They are followed by round grey cones, which give the plant its common name drumsticks. The small hairy seeds are found in the old flower parts.
The Australian Cultivar Registration Authority (ACRA) is the International Cultivar Registration Authority (ICRA) for Australian plant genera, excluding those genera or groups for which other ICRAs have been appointed.
An International Cultivation Registration Authority (ICRA) is an organization responsible for ensuring that the name of plant cultivars and cultivar groups are defined and not duplicated.
Priority is a fundamental principle of modern botanical nomenclature and zoological nomenclature. Essentially, it is the principle of recognising the first valid application of a name to a plant or animal. There are two aspects to this:
Cultivated plant taxonomy is the study of the theory and practice of the science that identifies, describes, classifies, and names cultigens—those plants whose origin or selection is primarily due to intentional human activity. Cultivated plant taxonomists do, however, work with all kinds of plants in cultivation.
The term grex, derived from the Latin noun grex, gregis, meaning 'flock', has been expanded in botanical nomenclature to describe hybrids of orchids, based solely on their parentage. Grex names are one of the three categories of plant names governed by the International Code of Nomenclature for Cultivated Plants; within a grex the cultivar group category can be used to refer to plants by their shared characteristics, and individual orchid plants can be selected and named as cultivars.
Roger David Spencer is an Australia horticultural botanist who was born at Alfreton, Derbyshire. He has an honours degree in botany from the University College of Wales, Aberystwyth, a master's degree and doctorate from the University of Melbourne and a technical certificate in gardening and turf maintenance from Oakleigh Technical College, Melbourne. He is currently horticultural botanist at the Royal Botanic Gardens Melbourne where he works in the Plant Identification Service, contributing locally and internationally to the study of cultivated plant taxonomy.
Disanthus cercidifolius is a species of flowering plant in the family Hamamelidaceae. It is native to woodland habitats in China and Japan.
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