This article includes a list of general references, but it lacks sufficient corresponding inline citations .(June 2013) |
Pinus cembra | |
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Swiss pine growing on the Dachstein, Austria | |
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Gymnospermae |
Division: | Pinophyta |
Class: | Pinopsida |
Order: | Pinales |
Family: | Pinaceae |
Genus: | Pinus |
Subgenus: | P. subg. Strobus |
Section: | P. sect. Quinquefoliae |
Subsection: | P. subsect. Strobus |
Species: | P. cembra |
Binomial name | |
Pinus cembra | |
Distribution |
Pinus cembra, also known as Swiss pine, Swiss stone pine, Arolla pine, Austrian stone pine, or just stone pine, is a species of pine tree in the subgenus Strobus.
The Swiss pine is a member of the white pine group, Pinus subgenus Strobus, and like all members of that group, the leaves ('needles') are in fascicles (bundles) of five, with a deciduous sheath. The mature size is typically between 25 metres (82 ft) and 35 metres (115 ft) in height, and the trunk diameter can be up to 1.5 metres (4.9 ft). However, it grows very slowly and it may take 30 years for the tree to reach 1.3 metres (4.3 ft).[ citation needed ] The cones, which contain the seeds (or nuts), of the Swiss pine are 4 centimetres (1.6 in) to 8 centimetres (3.1 in) long. Cones take 2 years (24 months) to mature. The 8 millimetres (0.31 in) to 12 millimetres (0.47 in) long seeds have only a vestigial wing and are dispersed by spotted nutcrackers. The species is long-lasting and can reach an age between 500 and 1000 years. [2]
In its natural environment, this tree usually reaches reproductive maturity at the age of 50 years (even 80 years, if the conditions are more extreme).
The very similar Siberian pine (Pinus sibirica) is treated as a variety or subspecies of Swiss pine by some botanists. The Siberian pine differs in having slightly larger cones, being more massive (taller, wider trunk), having a faster growth rate and needles with three resin canals instead of two as in the Swiss pine.
The Swiss pine grows in the Alps and Carpathian Mountains of central Europe, in Poland (Tatra Mountains), Switzerland, France, Italy, Austria, Germany, Slovenia, Slovakia (Tatra Mountains), Ukraine and Romania. It typically grows at 1,200 metres (3,900 ft) to 2,300 metres (7,500 ft) altitude. It often reaches the alpine tree line in this area.
Swiss pine associates with numerous species of mycorrhizal fungi from a young age, usually from the genus Suillus . [3] This symbiosis improves the tree's growth and survival rate.
Swiss pine is a popular ornamental tree in parks and large gardens, giving steady though not fast growth on a wide range of sites where the climate is cold. It is very tolerant of severe winter cold, hardy down to at least −50 °C (−58 °F), and also of wind exposure. The seeds are also harvested and sold as pine nuts. When cultivated, it will likely start producing cones after the age of 12 years, much faster than in the wild. This depends on the climate, soil type, mycorrhizal fungi etc. To make the tree bear cones faster, the tree can be inoculated with ectomycorrhizal fungi, such as Suillus luteus, Suillus americanus, Suillus placidus etc. These are the most effective.
Pine cones cut into slices are used to flavor schnapps, which is then sold as "Zirbenschnaps" or "Zirbeler" schnapps.
The wood is the most used for carvings in Val Gardena since the 17th century.
The cone of the Swiss pine was the field sign of the Roman legion stationed in Rhaetia in 15 BC, and hence it is used as the heraldic charge (known as Zirbelnuss in German) in the coat of arms of the city of Augsburg, the site of the Roman fort Augusta Vindelicorum.
It is also a species that is often used in bonsai.
Pinus cembra can be found in the uppermost forest belt where it helps to minimize the risk of avalanches and soil erosion. Due to this ability, the tree is valued as a stabilizing factor for afforestation projects at high elevations. [2]
Like other European and Asian white pines, Swiss pine is very resistant to white pine blister rust (Cronartium ribicola). This fungal disease was accidentally introduced from Europe into North America, where it has caused severe mortality in the American native white pines in many areas, notably, Western white pine and the closely related whitebark pine. Swiss pine is of great value for research into hybridization to develop rust resistance in these species.
A mycorrhiza is a symbiotic association between a fungus and a plant. The term mycorrhiza refers to the role of the fungus in the plant's rhizosphere, its root system. Mycorrhizae play important roles in plant nutrition, soil biology, and soil chemistry.
Western white pine, also called silver pine and California mountain pine, is a species of pine in the family Pinaceae. It occurs in mountain ranges of northwestern North America and is the state tree of Idaho.
Pinus peuce is a species of pine native to the mountains of North Macedonia, Bulgaria, Albania, Montenegro, Kosovo, the extreme southwest of Serbia, and the extreme north of Greece, growing typically at (600-) 1,000-2,200 (-2,300) m altitude. It often reaches the alpine tree line in this area. The mature size is up to 35–40 m height, and 1.5 m trunk diameter. However, the height of the tree diminishes strongly near the upper tree line and may even obtain shrub sizes.
Pinus flexilis, the limber pine, is a species of pine tree-the family Pinaceae that occurs in the mountains of the Western United States, Mexico, and Canada. It is also called Rocky Mountain white pine.
Pinus strobus, commonly called the eastern white pine, northern white pine, white pine, Weymouth pine (British), and soft pine is a large pine native to eastern North America. It occurs from Newfoundland, Canada, west through the Great Lakes region to southeastern Manitoba and Minnesota, United States, and south along the Appalachian Mountains and upper Piedmont to northernmost Georgia and perhaps very rarely in some of the higher elevations in northeastern Alabama. It is considered rare in Indiana.
The stone pine, botanical name Pinus pinea, also known as the Italian stone pine, Mediterranean stone pine, umbrella pine and parasol pine, is a tree from the pine family (Pinaceae). The tree is native to the Mediterranean region, occurring in Southern Europe and the Levant. The species was introduced into North Africa millennia ago, and is also naturalized in the Canary Islands, South Africa and New South Wales.
Pinus brutia, commonly known as the Turkish pine and Calabrian pine, is a species of pine native to the eastern Mediterranean region. The bulk of its range is in Turkey, but certain varieties are naturalized as far east as Afghanistan. It is also known as East Mediterranean pine, Afghan pine, and Brutia pine. The name "Calabrian pine" comes from an introduced grove in the Calabria region of southern Italy; historically this region was called Bruttium, which is likely where the specific epithet "brutia" comes from. Pinus brutia bears many similarities with other, closely related species such as Pinus halepensis and Pinus canariensis. Turkish pine forms a species complex with the former.
Pinus sibirica, or Siberian pine, in the family Pinaceae is a species of pine tree that occurs in Siberia from 58°E in the Ural Mountains east to 126°E in the Stanovoy Range in southern Sakha Republic, and from Igarka at 68°N in the lower Yenisei valley, south to 45°N in central Mongolia.
Suillus luteus is a bolete fungus, and the type species of the genus Suillus. A common fungus native all across Eurasia from Ireland to Korea, it has been introduced widely elsewhere, including North and South America, southern Africa, Australia and New Zealand. Commonly referred to as slippery jack or sticky bun in English-speaking countries, its names refer to the brown cap, which is characteristically slimy in wet conditions. The fungus, initially described as Boletus luteus by Carl Linnaeus in 1753, is now classified in a different fungus family as well as genus. Suillus luteus is edible, though not as highly regarded as other bolete mushrooms. It is commonly prepared and eaten in soups, stews or fried dishes. The slime coating, however, may cause indigestion if not removed before eating. It is often sold as a dried mushroom.
Rhizopogon is a genus of ectomycorrhizal basidiomycetes in the family Rhizopogonaceae. Species form hypogeous sporocarps commonly referred to as "false truffles". The general morphological characters of Rhizopogon sporocarps are a simplex or duplex peridium surrounding a loculate gleba that lacks a columnella. Basidiospores are produced upon basidia that are borne within the fungal hymenium that coats the interior surface of gleba locules. The peridium is often adorned with thick mycelial cords, also known as rhizomorphs, that attach the sporocarp to the surrounding substrate. The scientific name Rhizopogon is Greek for 'root' (Rhiz-) 'beard' (-pogon) and this name was given in reference to the rhizomorphs found on sporocarps of many species.
Suillus bovinus, also known as the Jersey cow mushroom or bovine bolete, is a pored mushroom of the genus Suillus in the family Suillaceae. A common fungus native to Europe and Asia, it has been introduced to North America and Australia. It was initially described as Boletus bovinus by Carl Linnaeus in 1753, and given its current binomial name by Henri François Anne de Roussel in 1806. It is an edible mushroom, though not highly regarded.
Suillus brevipes is a species of fungus in the family Suillaceae. First described by American mycologists in the late 19th century, it is commonly known as the stubby-stalk or the short-stemmed slippery Jack. The fruit bodies (mushrooms) produced by the fungus are characterized by a chocolate to reddish-brown cap covered with a sticky layer of slime, and a short whitish stipe that has neither a partial veil nor prominent, colored glandular dots. The cap can reach a diameter of about 10 cm, while the stipe is up to 6 cm long and 2 cm thick. Like other bolete mushrooms, S. brevipes produces spores in a vertically arranged layer of spongy tubes with openings that form a layer of small yellowish pores on the underside of the cap.
Suillus spraguei is a species of fungus in the family Suillaceae. It is known by a variety of common names, including the painted slipperycap, the painted suillus or the red and yellow suillus. Suillus spraguei has had a complex taxonomical history, and is also frequently referred to as Suillus pictus in the literature. The readily identifiable fruit bodies have caps that are dark red when fresh, dry to the touch, and covered with mats of hairs and scales that are separated by yellow cracks. On the underside of the cap are small, yellow, angular pores that become brownish as the mushroom ages. The stalk bears a grayish cottony ring, and is typically covered with soft hairs or scales.
Suillus quiescens is a pored mushroom of the genus Suillus in the family Suillaceae. First collected in 2002 on Santa Cruz Island off the coast of California, in association with Bishop Pine, the species was scientifically described and named in 2010. In addition to its distribution in coastal California, it was also found forming ectomycorrhizae with the roots of pine seedlings in the eastern Sierra Nevada, coastal Oregon, and the southern Cascade Mountains. It resembles Suillus brevipes, but can be distinguished from that species by its paler-colored immature cap and by the tiny colored glands on the stipe that darken with age.
Suillus sibiricus is a mushroom-forming fungus in the family Suillaceae. It is found in mountains of Europe, North America, and Siberia, strictly associated with several species of pine tree. Due to its specific habitat and rarity in Europe, it has been selected for inclusion in several regional Red Lists. Its fruit bodies are characterised by having slimy caps in wet weather, which can reach diameters of up to 10 cm (3.9 in). On the underside of the cap are yellow angular pores that bruise a pinkish to cinnamon colour. The stem is up to 8 cm (3.1 in) tall and 2.5 cm (1.0 in) wide and typically has a ring, a remnant of the partial veil that covers the fruit body in its early development. In North America, it is commonly called the Siberian slippery jack. Phylogenetic analysis has shown that S. sibiricus is closely related to S. umbonatus and S. americanus, and may in fact be conspecific with the latter species.
Suillus pungens, commonly known as the pungent slippery jack or the pungent suillus, is a species of fungus in the genus Suillus. The fruit bodies of the fungus have slimy convex caps up to 14 cm (5.5 in) wide. The mushroom is characterized by the very distinct color changes that occur in the cap throughout development. Typically, the young cap is whitish, later becoming grayish-olive to reddish-brown or a mottled combination of these colors. The mushroom has a dotted stem (stipe) up to 7 cm (2.8 in) long, and 2 cm (0.8 in) thick. On the underside on the cap is the spore-bearing tissue consisting of minute vertically arranged tubes that appear as a surface of angular, yellowish pores. The presence of milky droplets on the pore surface of young individuals, especially in humid environments, is a characteristic feature of this species. S. pungens can usually be distinguished from other similar Suillus species by differences in distribution, odor and taste. The mushroom is considered edible, but not highly regarded.
Suillus placidus, is a species of fungus in the genus Suillus. It is an edible pored mushroom found in European and North American coniferous forests, growing in association with several species of pine of the subgenus Strobus.
Suillus collinitus is a pored mushroom of the genus Suillus in the family Suillaceae. It is an edible mushroom found in European pine forests. The mushroom has a reddish to chestnut-brown cap that reaches up to 11 cm (4.3 in) in diameter, and a yellow stem measuring up to 7 cm (2.8 in) tall by 1 to 2 cm thick. On the underside of the cap are small angular pores, initially bright yellow before turning greenish-brown with age. A characteristic feature that helps to distinguish it from similar Suillus species, such as S. granulatus, is the pinkish mycelia at the base of the stem.
A snowbank fungus is any one of a number of diverse species of fungi that occur adjacent to or within melting snow. They are most commonly found in the mountains of western North America where a deep snowpack accumulates during the winter and slowly melts through the spring and summer, often shaded by coniferous forest. They may be saprotrophic, mycorrhizal, or in the case of Caloscypha fulgens, pathogenic.
Both naturally and artificially occurring pine species (Pinus) can hybridize, combining their genetic material and sometimes creating hybrids that can be more or less vigorous than their parent species. An example of a naturally occurring hybrid pine is Pinus × sondereggeri, a naturally occurring cross between loblolly pine and longleaf pine. An example of the many artificial hybrids is Pinus lambertiana × P. armandii.