This article needs additional citations for verification .(June 2013) |
Kermes oak | |
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Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Eudicots |
Clade: | Rosids |
Order: | Fagales |
Family: | Fagaceae |
Genus: | Quercus |
Subgenus: | Quercus subg. Cerris |
Section: | Quercus sect. Ilex |
Species: | Q. coccifera |
Binomial name | |
Quercus coccifera | |
Distribution map | |
Synonyms [2] | |
List
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Quercus coccifera, the kermes oak or holly oak, [3] is an oak shrub or tree in section Ilex of the genus. [4] It has many synonyms, including Quercus calliprinos. [2] It is native to the Mediterranean region and Northern African Maghreb, south to north from Morocco to France and west to east from Portugal to Cyprus and Turkey, crossing Spain, Italy, Libya, the Balkans, and Greece, including Crete. The Kermes oak was historically important as the food plant of Kermes scale insects, from which a red dye called crimson was obtained. [5] The etymology of the specific name coccifera is related to the production of red cochineal (crimson) dye and derived from Latin coccum which was from Greek κόκκος, the kermes insect. The Latin -fera means 'bearer'. [6]
Quercus coccifera is usually a shrub less than 2 metres (6+1⁄2 feet) high, rarely a small tree, reaching 1–6 metres (3+1⁄2–19+1⁄2 ft) tall (with 10 m or 33 ft specimens recorded in Kouf, Libya) [7] and 50 centimetres (20 inches) in trunk diameter. It is evergreen, with spiny-serrated coriaceous leaves 1.5–4 cm long and 1–3 cm broad. The acorns are 2–3 cm long and 1.5–2 cm in diameter when mature about 18 months after pollination. They are held in a cup covered in dense, elongated, reflexed scales.
It blooms from March to May in weather still wet. It is easily propagated by seed, an acorn that lies dormant until germinated by wet weather. This might occur anywhere from late summer to late autumn or early winter (October, November or December) of the following year. The acorns are very bitter, varying greatly in size and shape from one specimen to another and tasting bad. Acorns can germinate even before falling from the plant, but Q. coccifera is also multiplied by root suckers and layering.
Quercus coccifera was first described by Carl Linnaeus in 1753. [2] It is called "chêne des garrigues" (garrigue oak) in French. The term "garrigue" comes from Catalan or Occitan "garric" (meaning "twisted") the name for Q. coccifera in those languages. The common Spanish name of Q. coccifera is chaparro , which refers to its small size, a feature it shares with other oak species in similar habitats in other parts of the world, such as the chaparral communities from various parts of the Americas. The word chaparro comes from the Basque txapar meaning "little thicket". [8] Quercus coccifera is placed in section Ilex. [9]
Quercus calliprinos Webb, syn. Quercus coccifera subsp. calliprinos (Webb) Holmboe, the 'Palestinian oak' of the eastern Mediterranean, has been distinguished from the kermes oak by its larger size (more often a tree, up to 18 m) and larger acorns over 2 cm diameter.[ citation needed ] In Israel it is called the 'common oak' (Hebrew : אלון מצוי, IPA: [a'lonma'tsuj] ) or the 'Palestine oak'.[ citation needed ]As of February 2023 [update] , Plants of the World Online regards it as a synonym of Quercus coccifera, [2] but this is widely disputed, with many authors, particularly in the eastern Mediterranean, considering it distinct at least at subspecies rank, if not as a species. [3]
Quercus coccifera is an important Mediterranean bush or dwarf vegetation, where the biome it dominates often bears its name ( maquis , coscojar, garrigue , carrascal, chaparral, etc.). Q. coccifera forms monospecific communities or communities integrated with Pinus , mediterranean buckthorns, Myrtus , Arecaceae, junipers, Pistacia , Rosmarinus , Thymus , etc.
It is located throughout the region around the Mediterranean Sea, especially in central southern and eastern halves. It is similarly found on islands in the Mediterranean, from the Balearic Islands to Cyprus. It is common in Crete and can survive heavy sheep and goat grazing for long periods as a ground cover a few centimeters high. The same is true in Mallorca, Ibiza and the Iberian Peninsula.
The species grows on dry, sunny slopes. It supports both subhumid climates with dry summers or semi-desert climates with rainfall between 400 and 600 millimetres (16 and 24 in). A very hardy species, it grows well in all types of soils as a shrub, withstanding overgrazing. It is indifferent to soil chemistry, living on calcareous, pebbly, stony and poor soils. A lover of warm weather, it starts to fail from 1,000 m (3,300 ft) above sea level. It is capable of supporting the continental Mediterranean climate with more extreme temperatures and low rainfall, replacing Quercus ilex (holm oak) in drier areas where it excels in drought resistance. It also grows on sea cliffs and in windy areas where other species of Quercus or Pinus cannot resist the harsh weather conditions.
Populations typically occur in desert regions without any inhabited nucleus because crops are not economically profitable and the climate becomes progressively more continental and drier and therefore end in extreme temperatures accompanied by slow-growing dwarf juniper species. It is the last species of genus Quercus to disappear when rainfall is lacking. Their ecological importance is as a habitat and food source in these areas (they have edible acorns, although with a very bitter taste) for nesting birds, foxes, rodents and wild boars. It forms thickets, thorny and dense, some recorded as tall as 5 m. It is sometimes accompanied by other plant species of the same size and climber plants such as asparagus or Mediterranean smilax.
Q. coccifera is associated with several asparagus species, Crataegus monogyna , Mediterranean dwarf palm, ephedra , myrtle, several species of Junipers (Juniperus, sabinas...), Pistacia terebinthus , mastic, wild Olea europea , sarsaparilla, Rhamnus atlantica , Rhamnus lycioides , Rhamnus oleoides , Rhamnus catharticus etc. The communities receiving several characteristic names.
Kermes oaks have become scarce due to their replacement in wet zones by larger species such as Holm oak. It has also suffered from extensive culling for use as charcoal. It is the only food and shelter for wildlife in some areas, such as the Ebro valley and other dry areas where chaparral replaces oaks due to low rainfall.
It can survive heavy sheep and goat grazing for a long time as a ground carpet a few cm high, and will grow higher as a bush or a tree according to how much the grazing pressure is slackened.
It is included as an endangered species in the Red Book of Bulgaria. [10]
In the village of Kalopanagiotis, Cyprus, the tallest tree of this species has grown to a height of 17 meters. The tree is approximately 700 years old.[ citation needed ]
Quercus ilex, the holm oak, also evergreen oak, is a large evergreen oak native to the Mediterranean region. It is a member of the section Ilex of the genus, with acorns that mature in a single summer.
The Eastern Mediterranean conifer-sclerophyllous-forests, also known as the Eastern Mediterranean conifer-forests, is an ecoregion in the eastern Mediterranean Basin. It covers portions of Turkey, Syria, Iraq, Lebanon, Israel, Palestinian territories, Jordan, and Saudi Arabia.
Quercus aucheri, known as Aucher's oak or the Boz-Pirnal oak is a species of oak tree in the family Fagaceae. It is found in limited portions of the Aegean islands of Greece and parts of Anatolian Turkey. It is placed in section Ilex.
Quercus alnifolia, commonly known as the golden oak, is an evergreen oak species of Cyprus. Its common English name refers to the golden coloured lower surface of its leaves. Quercus alnifolia belongs to the endemic flora of the island and it is confined to the igneous geological complex of the Troodos Mountains. In February 2006, the parliament of Cyprus selected the golden oak to be the country's national tree.
Quercus vacciniifolia, the huckleberry oak, is a member of the Protobalanus section of genus Quercus. It has evergreen foliage, short styles, very bitter acorns that mature in 18 months, and a woolly acorn shell interior.
The Mediterranean woodlands and forests is an ecoregion in the coastal plains, hills, and mountains bordering the Mediterranean Sea and Atlantic Ocean in North Africa. It has a Mediterranean climate, and is in the Mediterranean forests, woodlands, and scrub biome.
The Illyrian deciduous forests is a terrestrial ecoregion in southern Europe, which extends along the eastern coast of the Adriatic Sea. It belongs to the Mediterranean forests, woodlands, and scrub biome, and is in the Palearctic realm.
Morocco provides a refuge for a rich and diverse flora with about 4,200 taxa, of which 22% are endemic. The phytogeographic zones of Morocco comprise 8 zones: the Mediterranean zone, the Cedar zone (1000-2000m), the sub-Alpine zone (2,000-2,500m), the Alpine zone (2,500m+), the semi-desert scrub zone, the Reg, the sandy desert zone and the oases.
The Italian sclerophyllous and deciduous forests ecoregion, part of the Mediterranean forests, woodlands, and scrub biome, is in Italy. The ecoregion covers most of the Italian Peninsula and includes both evergreen and deciduous forests.
The Tyrrhenian-Adriatic sclerophyllous and mixed forests is an ecoregion in southern Italy, Sicily, Sardinia, Corsica, the Dalmatian Islands of Croatia, and Malta.
Quercus rotundifolia, the holm oak or ballota oak, is an evergreen oak native to the western Mediterranean region, with the majority of the population in the Iberian Peninsula and minor populations in Northwest Africa. The species was first described by Jean-Baptiste Lamarck in 1785. It is the typical species of the Iberian dehesa or montado, where its sweet-astringent acorns are a source of food for livestock, particularly the Iberian pig. Its acorns have also been used for human nourishment since the Neolithic era. It is placed in section Ilex. Some authors described it as a subspecies of Quercus ilex.
Rhamnus lycioides, the black hawthorn, European buckthorn, or Mediterranean buckthorn, is a shrub up to about 1.5-4 metres tall in the buckthorn family, Rhamnaceae. It is found in the Mediterranean region, in southern Europe and northern Africa. Its scientific name lycioides refers to its resemblance to the botanical genus Lycium.
Quercus franchetii, commonly known as the zhui lian li evergreen oak, is a species of oak in the Ilex section of the genus, native to a wide area of eastern Asia. It is an oak native to China, northern Thailand and Vietnam, growing at altitudes between 800 and 2,600 metres.
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The Southwest Iberian Mediterranean sclerophyllous and mixed forests is a Mediterranean forests, woodlands, and scrub ecoregion in southwestern Europe. It occupies the southwestern Iberian Peninsula, encompassing coastal lowlands and mountains in portions of Portugal and Spain.
The Iberian sclerophyllous and semi-deciduous forests is a Mediterranean forests, woodlands, and scrub ecoregion in southwestern Europe. It occupies the interior valleys and plateaus of the Iberian Peninsula. The ecoregion lies mostly in Spain, and includes some portions of eastern Portugal.
The Northeastern Spain and Southern France Mediterranean forests is a Mediterranean forests, woodlands, and scrub ecoregion in southwestern Europe. It occupies the Mediterranean coastal region of northeastern Spain, Southern France, the Balearic Islands and a small part of Italy.
The Southeastern Iberian shrubs and woodlands is a Mediterranean forests, woodlands, and scrub ecoregion in southwestern Europe. It lies in southeastern coastal Spain, along the Mediterranean Sea.
Quercus subgenus Quercus is one of the two subgenera into which the genus Quercus was divided in a 2017 classification. It contains about 190 species divided among five sections. It may be called the New World clade or the high-latitude clade; most species are native to the Americas, the others being found in Eurasia and northernmost North Africa.
Quercus subgenus Cerris is one of the two subgenera into which the genus Quercus was divided in a 2017 classification. It contains about 140 species divided among three sections. It may be called the Old World clade or the mid-latitude clade; all species are native to Eurasia and North Africa.