Mediterranean forests, woodlands, and scrub

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Extent of Mediterranean forests, woodlands, and scrub Biome map 12.svg
Extent of Mediterranean forests, woodlands, and scrub

Mediterranean forests, woodlands and scrub is a biome defined by the World Wide Fund for Nature. [1] The biome is generally characterized by dry summers and rainy winters, although in some areas rainfall may be uniform. Summers are typically hot in low-lying inland locations but can be cool near colder seas. Winters are typically mild to cool in low-lying locations but can be cold in inland and higher locations. All these ecoregions are highly distinctive, collectively harboring 10% of the Earth's plant species. [2]

Contents

Jabal Moussa Biosphere Reserve in Lebanon Jabal Moussa Biosphere Reserve 03.jpg
Jabal Moussa Biosphere Reserve in Lebanon

Distribution

A Mediterranean forest, in the Region of Murcia (Spain). Sierra de Burete.JPG
A Mediterranean forest, in the Region of Murcia (Spain).

The Mediterranean forests, woodlands, and scrub biome mostly occurs in, but not limited to, the Mediterranean climate zones, in the mid-latitudes: [1]

The biome is not limited to the Mediterranean climate zone. It can also be present in other climate zones (which typically border the Mediterranean climate zone), such as the drier regions of the oceanic and humid subtropical climates, and as well as the lusher areas of the semi-arid climate zone. Non-Mediterranean climate regions that would feature Mediterranean vegetation include the Nile River Valley in Egypt (extending upstream along the riverbanks), parts of the Eastern Cape in South Africa, southeastern Australia, southeastern Azerbaijan, southeastern Turkey, far northern Iraq, the Mazandaran Province in Iran, Central Italy, parts of the Balkans (including Northern Greece), as well as Northern and Western Jordan.

Vegetation

Close up of Ephesos ancient Greek amphitheatre with a maquis shrubland Ephesos amphitheatre.jpg
Close up of Ephesos ancient Greek amphitheatre with a maquis shrubland

Vegetation types range from forests to woodlands, savannas, shrublands, and grasslands; "mosaic habitat" landscapes are common, where differing vegetation types are interleaved with one another in complex patterns created by variations in soil, topography, exposure to wind and sun, and fire history. Much of the woody vegetation in Mediterranean-climate regions is sclerophyll, which means 'hard-leaved' in Greek. Sclerophyllous vegetation generally has small, dark leaves covered with a waxy outer layer to retain moisture in the dry summer months.

Phytogeographers consider the fynbos (South Africa) as a separate floral kingdom because 68% of the 8,600 vascular plant species crowded into its 90,000 square kilometers (35,000 sq mi) are endemic and highly distinctive at several taxonomic levels. [1] [3] This is equivalent to about 40% of the plant species of the United States and Canada combined, found within an area the size of the state of Maine. The fynbos and Southwest Australia shrublands have flora that are significantly more diverse than the other ecoregions, although any Mediterranean shrubland is still rich in species and endemics relative to other non-forest ecoregions. [1] [3]

Biome plant groups

Springtime in Chilean Matorral a few kilometers north of Santiago along the Pan-American Highway Matorral Cerro Aconcagua.JPG
Springtime in Chilean Matorral a few kilometers north of Santiago along the Pan-American Highway

Major plant communities in this biome include:

Fire as a medium of change

Scrubland of the Sierra Pelona Mountains, Southern California Liebre Mountains 001.jpg
Scrubland of the Sierra Pelona Mountains, Southern California

Fire, both natural and human-caused, has played a large role in shaping the ecology of Mediterranean ecoregions. [4] The hot, dry summers make much of the region prone to fires, and lightning-caused fires occur with some frequency. Many of the plants are pyrophytes, or fire-loving, adapted or even depending on fire for reproduction, recycling of nutrients, and the removal of dead or senescent vegetation. In both the Australian and Californian Mediterranean-climate eco-regions, native peoples used fire extensively to clear brush and trees, making way for the grasses and herbaceous vegetation that supported game animals and useful plants.

The plant communities in these areas adapted to the frequent human-caused fires, and pyrophyte species grew more common and more fire-loving, while plants that were poorly adapted to fire retreated. After European colonization of these regions, fires were suppressed, which has caused some unintended consequences in these ecoregions; fuel builds up, so that when fires do come they are much more devastating, and some species dependent on fire for their reproduction are now threatened. The European shrublands have also been shaped by anthropogenic fire, [5] historically associated with transhumance herding of sheep and goats.

Though adapted to infrequent fires, chaparral plant communities can be eliminated by frequent fires. A high frequency of fire (less than ten years) will result in the loss of obligate seeding shrub species such as Manzanita spp. This high frequency disallows seeder plants to reach their reproductive size before the next fire and the community shifts to a sprouter-dominance. If high frequency fires continue over time, obligate resprouting shrub species can also be eliminated by exhausting their energy reserves below-ground. Today, frequent accidental ignitions can convert chaparral from a native shrubland to non-native annual grassland and drastically reduce species diversity, especially under drought brought about by climate change. [6] [7]

On 25 July 2023, devastating wildfires were burning in at least nine countries across the Mediterranean, including Croatia, Italy, and Portugal, with thousands of firefighters in Europe and North Africa working to contain flames stoked by high temperatures, dry conditions, and strong winds. The wildfires led to casualties, evacuations of thousands of people, and widespread destruction of homes and forests. [8]

Degradation

Mediterranean cork oak trees, in Monchique, Algarve, Portugal Sobreiros - Monchique - 04.11.2018.jpg
Mediterranean cork oak trees, in Monchique, Algarve, Portugal
Sclerophyll woodland, Sydney Royal National Park NSW 2233, Australia - panoramio (23).jpg
Sclerophyll woodland, Sydney
Festival of degradation VRUJA established by activists and artists in 2021 in Dalmatia, Croatia VRUJA FESTIVAL 20 10 28 064000.jpeg
Festival of degradation VRUJA established by activists and artists in 2021 in Dalmatia, Croatia

Mediterranean ecoregions are some of the most endangered and vulnerable on the planet. Many have suffered tremendous degradation and habitat loss through logging, overgrazing, conversion to agriculture, urbanization, fire suppression, and introduction of exotic and invasive species. The ecoregions around the Mediterranean basin and in California have been particularly affected by degradation due to human activity, suffering extensive loss of forests and soil erosion, and many native plants and animals have become extinct or endangered.

See also

Related Research Articles

<span class="mw-page-title-main">Chaparral</span> Shrubland plant community in western North America

Chaparral is a shrubland plant community found primarily in California, in southern Oregon and in the northern portion of the Baja California Peninsula in Mexico. It is shaped by a Mediterranean climate and infrequent, high-intensity crown fires.

<span class="mw-page-title-main">Sclerophyll</span> Type of plant

Sclerophyll is a type of vegetation that is adapted to long periods of dryness and heat. The plants feature hard leaves, short internodes and leaf orientation which is parallel or oblique to direct sunlight. The word comes from the Greek sklēros (hard) and phyllon (leaf). The term was coined by A.F.W. Schimper in 1898, originally as a synonym of xeromorph, but the two words were later differentiated.

The Global 200 is the list of ecoregions identified by the World Wide Fund for Nature (WWF), the global conservation organization, as priorities for conservation. According to WWF, an ecoregion is defined as a "relatively large unit of land or water containing a characteristic set of natural communities that share a large majority of their species dynamics, and environmental conditions". For example, based on their levels of endemism, Madagascar gets multiple listings, ancient Lake Baikal gets one, and the North American Great Lakes get none.

<span class="mw-page-title-main">Savanna</span> Mixed woodland-grassland ecosystem

A savanna or savannah is a mixed woodland-grassland ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to support an unbroken herbaceous layer consisting primarily of grasses. According to Britannica, there exists four savanna forms; savanna woodland where trees and shrubs form a light canopy, tree savanna with scattered trees and shrubs, shrub savanna with distributed shrubs, and grass savanna where trees and shrubs are mostly nonexistent.

<span class="mw-page-title-main">California chaparral and woodlands</span> Ecoregion in the western United States and Mexico

The California chaparral and woodlands is a terrestrial ecoregion of southwestern Oregon, northern, central, and southern California and northwestern Baja California (Mexico), located on the west coast of North America. It is an ecoregion of the Mediterranean forests, woodlands, and scrub biome, and part of the Nearctic realm.

<span class="mw-page-title-main">Ecology of California</span> Environments and natural history of California

The ecology of California can be understood by dividing the state into a number of ecoregions, which contain distinct ecological communities of plants and animals in a contiguous region. The ecoregions of California can be grouped into four major groups: desert ecoregions, Mediterranean ecoregions, forested mountains, and coastal forests.

<span class="mw-page-title-main">Southwest Australia</span> Biogeographic region of Western Australia

Southwest Australia is a biogeographic region in Western Australia. It includes the Mediterranean-climate area of southwestern Australia, which is home to a diverse and distinctive flora and fauna.

<span class="mw-page-title-main">Northern California coastal forests</span> Temperate coniferous forests ecoregion in northern California and southwestern Oregon

The Northern California coastal forests are a temperate coniferous forests ecoregion of coastal Northern California and southwestern Oregon.

<span class="mw-page-title-main">Shrubland</span> Vegetation dominated by shrubs

Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominated by shrubs, often also including grasses, herbs, and geophytes. Shrubland may either occur naturally or be the result of human activity. It may be the mature vegetation type in a particular region and remain stable over time, or a transitional community that occurs temporarily as the result of a disturbance, such as fire. A stable state may be maintained by regular natural disturbance such as fire or browsing. Shrubland may be unsuitable for human habitation because of the danger of fire. The term was coined in 1903.

<span class="mw-page-title-main">Mediterranean Basin</span> Region of lands around the Mediterranean Sea that have a Mediterranean climate

In biogeography, the Mediterranean Basin, also known as the Mediterranean Region or sometimes Mediterranea, is the region of lands around the Mediterranean Sea that have mostly a Mediterranean climate, with mild to cool, rainy winters and warm to hot, dry summers, which supports characteristic Mediterranean forests, woodlands, and scrub vegetation. It was a very important part of Mediterranean civilizations.

<span class="mw-page-title-main">Garrigue</span> Shrubland plant community in the Mediterranean

Garrigue or garigue, also known as phrygana, is a type of low scrubland ecoregion and plant community in the Mediterranean forests, woodlands, and scrub biome.

<span class="mw-page-title-main">Coastal sage scrub</span> Shrubland plant community of California

Coastal sage scrub, also known as coastal scrub, CSS, or soft chaparral, is a low scrubland plant community of the California coastal sage and chaparral subecoregion, found in coastal California and northwestern coastal Baja California. It is within the California chaparral and woodlands ecoregion, of the Mediterranean forests, woodlands, and scrub biome.

<span class="mw-page-title-main">Klamath Mountains (ecoregion)</span> Temperate coniferous forests ecoregion in northern California and southwestern Oregon

The Klamath Mountains ecoregion of Oregon and California lies inland and north of the Coast Range ecoregion, extending from the Umpqua River in the north to the Sacramento Valley in the south. It encompasses the highly dissected ridges, foothills, and valleys of the Klamath and Siskiyou Mountains. It corresponds to the Level III ecoregion designated by the Environmental Protection Agency and to the Klamath-Siskiyou forests ecoregion designated by the World Wide Fund for Nature.

<span class="mw-page-title-main">Chilean Matorral</span> Mediterranean forests, woodlands, and scrub ecoregion in central Chile

The Chilean Matorral (NT1201) is a terrestrial ecoregion of central Chile, located on the west coast of South America. It is in the Mediterranean forests, woodlands, and scrub biome, part of the Neotropical realm.

<span class="mw-page-title-main">California coastal sage and chaparral</span> Mediterranean forests, woodlands, and scrub ecoregion in Mexico and the United States

The California coastal sage and chaparral is a Mediterranean forests, woodlands, and scrub ecoregion located in southwestern California and northwestern Baja California (Mexico). It is part of the larger California chaparral and woodlands ecoregion.

<span class="mw-page-title-main">California montane chaparral and woodlands</span> Mediterranean forests, woodlands, and scrub ecoregion in California, United States

The California montane chaparral and woodlands is an ecoregion defined by the World Wildlife Fund, spanning 7,900 square miles (20,000 km2) of mountains in the Transverse Ranges, Peninsular Ranges, and Coast Ranges of southern and central California. The ecoregion is part of the larger California chaparral and woodlands ecoregion, and belongs to the Mediterranean forests, woodlands, and scrub biome.

<span class="mw-page-title-main">California interior chaparral and woodlands</span> Mediterranean forests, woodlands, and scrub ecoregion in California, United States

The California interior chaparral and woodlands ecoregion covers 24,900 square miles (64,000 km2) in an elliptical ring around the California Central Valley. It occurs on hills and mountains ranging from 300 feet (91 m) to 3,000 feet (910 m). It is part of the Mediterranean forests, woodlands, and scrub biome, with cool, wet winters and hot, dry summers. Temperatures within the coast can range from 53° to 65 °F and 32° to 60 °F within the mountains. Many plant and animal species in this ecoregion are adapted to periodic fire.

<span class="mw-page-title-main">Montane ecosystems</span> Ecosystems found in mountains

Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures fall as elevation increases, causing the ecosystem to stratify. This stratification is a crucial factor in shaping plant community, biodiversity, metabolic processes and ecosystem dynamics for montane ecosystems. Dense montane forests are common at moderate elevations, due to moderate temperatures and high rainfall. At higher elevations, the climate is harsher, with lower temperatures and higher winds, preventing the growth of trees and causing the plant community to transition to montane grasslands and shrublands or alpine tundra. Due to the unique climate conditions of montane ecosystems, they contain increased numbers of endemic species. Montane ecosystems also exhibit variation in ecosystem services, which include carbon storage and water supply.

<span class="mw-page-title-main">Mediterranean California</span> Ecoregion of North America

Mediterranean California is a Level I ecoregion of North America designated by the Commission for Environmental Cooperation (CEC) in its North American Environmental Atlas. The region is present only in California and Baja California.

References

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  2. Cody, M.L. (1986). "Diversity, rarity, and conservation in Mediterranean-climate regions". In Soulé, M.E. (ed.). Conservation biology. Massachusetts, US. pp. 122–152.{{cite book}}: CS1 maint: location missing publisher (link)
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  7. Pratt, R. Brandon; Jacobsen, Anna L.; Ramirez, Aaron R.; Helms, Anjel M.; Traugh, Courtney A.; Tobin, Michael F.; Heffner, Marcus S.; Davis, Stephen D. (1 March 2014). "Mortality of resprouting chaparral shrubs after a fire and during a record drought: physiological mechanisms and demographic consequences". Global Change Biology. 20 (3): 893–907. Bibcode:2014GCBio..20..893P. doi:10.1111/gcb.12477. ISSN   1365-2486. PMID   24375846. S2CID   19688559.
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