These regions are further subdivided into a variety of ecoregions. Many ecosystems and the animal and plant communities that depend on them extend across a number of continents and cover large portions of the Holarctic realm. This continuity is the result of those regions’ shared glacial history.
Major ecosystems
Within the Holarctic realm, there are a variety of ecosystems. The type of ecosystem found in a given area depends on its latitude and the local geography. In the far north, a band of Arctic tundra encircles the shore of the Arctic Ocean. The ground beneath this land is permafrost (frozen year-round). In these difficult growing conditions, few plants can survive. South of the tundra, the boreal forest stretches across North America and Eurasia. This land is characterized by coniferous trees. Further south, the ecosystems become more diverse. Some areas are temperate grassland, while others are temperate forests dominated by deciduous trees. Many of the southernmost parts of the Holarctic are deserts, which are dominated by plants and animals adapted to the dry conditions.[1]
Animal species with a Holarctic distribution
A variety of animal species are distributed across continents, throughout much of the Holarctic realm. These include the brown bear, grey wolf, red fox, wolverine, moose, caribou, golden eagle and common raven.
The brown bear (Ursus arctos) is found in mountainous and semi-open areas distributed throughout the Holarctic. It once occupied much larger areas, but has been driven out by human development and the resulting habitat fragmentation. Today it is only found in remaining wilderness areas.
The grey wolf (Canis lupus) is found in a wide variety of habitats from tundra to desert, with different populations adapted for each. Its historical distribution encompasses the vast majority of the Holarctic realm, though human activities such as development and active extermination have extirpated the species from much of this range.
The red fox (Vulpes vulpes) is a highly adaptable predator. It has the widest distribution of any terrestrial carnivore, and is adapted to a wide range of habitats, including areas of intense human development. Like the wolf, it is distributed throughout the majority of the Holarctic, but it has avoided extirpation.
The wolverine (Gulo gulo) is a large member of the weasel family found primarily in the arctic and in boreal forests, ranging south in mountainous regions. It is distributed in such areas throughout Eurasia and North America.
The moose (Alces alces) is the largest member of the deer family. It is found throughout most of the boreal forest through continental Eurasia into Scandinavia, eastern North America, and boreal and montane regions of western North America. In some areas it ranges south into the deciduous forest.
The caribou, or reindeer (Rangifer tarandus) is found in boreal forest and tundra in the northern parts of the Holarctic. In Eurasia, it has been domesticated. It is divided into several subspecies, which are adapted to different habitats and geographic areas.
The golden eagle (Aquila chrysaetos) is one of the best-known birds of prey in the Northern Hemisphere. It is the most widely distributed species of eagle. Golden eagles use their agility and speed combined with powerful feet and massive, sharp talons to snatch up a variety of prey (mainly hares, rabbits, marmots and other ground squirrels).
The common raven (Corvus corax) is the most widespread of the corvids, and one of the largest. It is found in a variety of habitats, but primarily wooded northern areas. It has been known to adapt well to areas of human activity. Their distribution also makes up most of the Holarctic realm.
Leptothorax acervorum is a small red Holarctic ant widely distributed across Eurasia, ranging from central Spain and Italy to the northernmost parts of Scandinavia and Siberia.
Zygiella x-notata is a species of orb-weaving spider with a Holarctic distribution, mostly inhabiting urban and suburban regions of Europe and parts of North America.
Hemerobius humulinus is a species of brown lacewing in the family Hemerobiidae. It is found in Europe and Northern Asia (excluding China), North America, and Southern Asia.[2]
Origin
The continuity of the northern parts of the Holarctic results from their shared glacial history. During the Pleistocene (Ice Age), these areas were subjected to repeated glaciations. Icecaps expanded, scouring the land of life and reshaping its topography. During glacial periods, species survived in refugia, small areas that maintained a suitable climate due to local geography. These areas are believed to have been primarily in southern regions, but some genetic and paleontological evidence points to additional refugia in the sheltered areas of the north.[3]
Wherever these areas were found, they became source populations during interglacial periods. When the glaciers receded, plants and animals spread rapidly into the newly opened areas. Different taxa responded to these rapidly changing conditions in different ways. Tree species spread outward from refugia during interglacial periods, but in varied patterns, with different trees dominating in different periods.[4]Insects, on the other hand, shifted their ranges with the climate, maintaining consistency in species for the most part throughout the period.[5] Their high degree of mobility allowed them to move as the glaciers advanced or retreated, maintaining a constant habitat despite the climatic oscillations. Despite their apparent lack of mobility, plants managed to colonize new areas rapidly as well. Studies of fossil pollen indicate that trees recolonized these lands at an exponential rate.[6] Mammals recolonized at varying rates. Brown bears, for instance, moved quickly from refugia with the receding glaciers, becoming one of the first large mammals to recolonize the land.[7] The Last Glacial Period ended about 10,000 years ago, resulting in the present distribution of ecoregions.
Another factor contributing to the continuity of Holarctic ecosystems is the movement between continents allowed by the Bering land bridge, which was exposed by the lowering of sea level due to the expansion of the ice caps. The communities found in the Palearctic and the Nearctic are different, but have many species in common. This is the result of several faunal interchanges that took place across the Bering land bridge. However, these migrations were mostly limited to large, cold-tolerant species.[8] Today it is mainly these species which are found throughout the realm.
Threats
As the Holarctic is an enormous area, it is subject to environmental problems of international scale. The primary threats throughout the region result from global warming and habitat fragmentation.[citation needed] The former is of particular concern in the north, as these ecosystems are adapted to cold. The latter is more of a concern in the south, where development is prevalent.
Global warming is a threat to all the Earth's ecosystems, but it is a more immediate threat to those found in cold climates. The communities of species found at these latitudes are adapted to the cold, so any significant warming can upset the balance. For instance, insects struggle to survive the cold winters typical of the boreal forest. Many do not make it, especially in harsh winters. However, recently the winters have grown milder, which has had a drastic effect on the forest. Winter mortality of some insect species drastically decreased, allowing the population to build on itself in subsequent years. In some areas the effects have been severe. Spruce beetle outbreaks have wiped out up to ninety percent of the Kenai Peninsula's spruce trees; this is blamed primarily on a series of unusually warm years since 1987.[9]
In this case a native species has caused massive disturbance of habitat as a result of climate change. Warming temperatures may also allow pest species to enlarge their range, moving into habitats that were previously unsuitable. Studies of potential areas for outbreaks of bark beetles indicate that as the climate shifts, these beetles will expand to the north and to higher elevations than they have previously affected.[10] With warmer temperatures, insect infestation will become a greater problem throughout the northern parts of the Holarctic.
Another potential effect of global warming to northern ecosystems is the melting of permafrost. This can have significant effects on the plant communities that are adapted to the frozen soil, and may also have implications for further climate change. As permafrost melts, any trees growing above it may die, and the land shifts from forest to peatland. In the far north, shrubs may later take over what was formerly tundra. The precise effect depends on whether the water that was locked up is able to drain off. In either case, the habitat will undergo a shift. Melting permafrost may also accelerate climate change in the future. Within the permafrost, vast quantities of carbon are locked up. If this soil melts, the carbon may be released into the air as either carbon dioxide or methane. Both of these are greenhouse gases.[11]
Habitat fragmentation threatens a wide variety of habitats throughout the world, and the Holarctic is no exception. Fragmentation has a variety of negative effects on populations. As populations become cut off, their genetic diversity suffers and they become susceptible to sudden disasters and extinction. While the northern parts of the Holarctic represent some of the largest areas of wilderness left on Earth, the southern parts are in some places extensively developed. This realm contains most of the world's developed countries, including the United States and the nations of Western Europe. Temperate forests were the primary ecosystem in many of the most developed areas today. These lands are now used for intensive agriculture or have become urbanized. As lands have been developed for agricultural uses and human occupation, natural habitat has for the most part become limited to areas considered unsuitable for human use, such as slopes or rocky areas.[12] This pattern of development limits the ability of animals, especially large ones, to migrate from place to place.
Large carnivores are particularly affected by habitat fragmentation. These mammals, such as brown bears and wolves, require large areas of land with relatively intact habitat to survive as individuals. Much larger areas are required to maintain a sustainable population. They may also serve as keystone species, regulating the populations of the species they prey on. Thus, their conservation has direct implications for a wide range of species, and is difficult to accomplish politically due to the large size of the areas they need.[13] With increasing development, these species in particular are at risk, which could have effects that carry down throughout the ecosystem.
Conservation actions
This section needs to be updated. Please help update this article to reflect recent events or newly available information.(November 2020)
The threats to the Holarctic realm are not going unrecognized. Many efforts are being made to mitigate these threats, with the hope of preserving the biodiversity of the region. International agreements to combat global warming may help to lessen the effects of climate change on this region. Efforts are also underway to fight habitat fragmentation, both on local and regional scales.
The most comprehensive effort to combat global warming to date is the Kyoto Protocol. Developed countries who sign this protocol agree to cut their collective greenhouse gas emissions by five percent since 1990 by sometime between 2008 and 2012. The vast majority of these nations are found within the Holarctic. Each country is given a target for emission levels, and they may trade emissions credits in a market-based system that includes developing countries as well. Once this period is ended, a new agreement will be written to further mitigate the effects of climate change. The process of drafting a new agreement has already begun. In late 2007, an international meeting in Bali was held to begin planning for the successor to the Kyoto Protocol. This agreement will aim to build on the successes and failures of Kyoto to produce a more effective method of cutting greenhouse gas emissions (UNFCCC). If these efforts are successful, the biodiversity of the Holarctic and the rest of the world will see fewer effects of climate change.
Fighting habitat fragmentation is a major challenge in conserving the wide-ranging species of the Holarctic. Some efforts are limited to a local scale of protection, while others are regional in scope. Local efforts include creating reserves and establishing safe routes for animals to cross roads and other human-made barriers. Regional efforts to combat habitat fragmentation take a broader scope.
One major such effort in the Holarctic is the Yellowstone to Yukon Conservation Initiative. This organization was started in 1997 to help establish a contiguous network of protection for the northern Rocky Mountains, from mid Wyoming to the border between Alaska and Canada's Yukon. It brings together a wide variety of environmental organizations for a shared purpose. The goal of the Initiative is to create a core of protected areas, connected by corridors and surrounded by buffer zones. This will build on the many existing protected areas in this region, with a focus on integrating existing and future human activities into the conservation plan rather than seeking to exclude them (Yellowstone to Yukon). If these efforts are successful, they will be especially beneficial to wide-ranging species such as grizzly bears. If these species can survive, other members of the communities they live in will survive as well.
Related Research Articles
In physical geography, tundra is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. There are three regions and associated types of tundra: Arctic tundra, alpine tundra, and Antarctic tundra.
Taiga or tayga, also known as boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruces, and larches. The taiga or boreal forest is the world's largest land biome. In North America, it covers most of inland Canada, Alaska, and parts of the northern contiguous United States. In Eurasia, it covers most of Sweden, Finland, much of Russia from Karelia in the west to the Pacific Ocean, much of Norway and Estonia, some of the Scottish Highlands, some lowland/coastal areas of Iceland, and areas of northern Kazakhstan, northern Mongolia, and northern Japan.
The Nearctic realm is one of the eight biogeographic realms constituting the Earth's land surface.
In biology, a refugium is a location which supports an isolated or relict population of a once more widespread species. This isolation (allopatry) can be due to climatic changes, geography, or human activities such as deforestation and overhunting.
The mammoth steppe, also known as steppe-tundra, was once the Earth's most extensive biome. During glacial periods in the later Pleistocene it stretched east-to-west, from the Iberian Peninsula in the west of Europe, across Eurasia to North America, through Beringia and northwest Canada; from north-to-south, the steppe reached from the Arctic southward to southern Europe, Central Asia and northern China. The mammoth steppe was cold and dry, and relatively featureless, though climate, topography, and geography varied considerably throughout. Certain areas of the biome—such as coastal areas—had wetter and milder climates than others. Some areas featured rivers which, through erosion, naturally created gorges, gulleys, or small glens. The continual glacial recession and advancement over millennia contributed more to the formation of larger valleys and different geographical features. Overall, however, the steppe is known to be flat and expansive grassland. The vegetation was dominated by palatable, high-productivity grasses, herbs and willow shrubs.
In paleoclimatology of the Holocene, the Boreal was the first of the Blytt–Sernander sequence of north European climatic phases that were originally based on the study of Danish peat bogs, named for Axel Blytt and Rutger Sernander, who first established the sequence. In peat bog sediments, the Boreal is also recognized by its characteristic pollen zone. It was preceded by the Younger Dryas, the last cold snap of the Pleistocene, and followed by the Atlantic, a warmer and moister period than our most recent climate. The Boreal, transitional between the two periods, varied a great deal, at times having within it climates like today's.
A boreal ecosystem is an ecosystem with a subarctic climate located in the Northern Hemisphere, approximately between 50° and 70°N latitude. These ecosystems are commonly known as taiga and are located in parts of North America, Europe, and Asia. The ecosystems that lie immediately to the south of boreal zones are often called hemiboreal. There are a variety of processes and species that occur in these areas as well.
Last Glacial Maximum refugia were places (refugia) in which humans and other species survived during the Last Glacial Period, around 25,000 to 18,000 years ago. Glacial refugia are areas that climate changes were not as severe, and where species could recolonize after deglaciation.
Arctic ecology is the scientific study of the relationships between biotic and abiotic factors in the arctic, the region north of the Arctic Circle. This region is characterized by two biomes: taiga and tundra. While the taiga has a more moderate climate and permits a diversity of both non-vascular and vascular plants, the tundra has a limited growing season and stressful growing conditions due to intense cold, low precipitation, and a lack of sunlight throughout the winter. Sensitive ecosystems exist throughout the Arctic region, which are being impacted dramatically by global warming.
The fauna of Europe is all the animals living in Europe and its surrounding seas and islands. Europe is the western part of the Palearctic realm. Lying within the temperate region, the wildlife is not as rich as in the hottest regions, but is nevertheless diverse due to the variety of habitats and the faunal richness of Eurasia as a whole.
Ecotourism in the United States is commonly practiced in protected areas such as national parks and nature reserves. The principles and behaviors of ecotourism are slowly becoming more widespread in the United States; for example, hotels in some regions strive to be more sustainable.
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The boreal woodland caribou, also known as Eastern woodland caribou, boreal forest caribou and forest-dwelling caribou, is a North American subspecies of reindeer found primarily in Canada with small populations in the United States. Unlike the Porcupine caribou and barren-ground caribou, boreal woodland caribou are primarily sedentary.
The West Siberian taiga ecoregion covers the West Siberian Plain in Russia, from the Ural Mountains in the west to the Yenisei River in the east, and roughly from 56° N to 66° N latitude. It is a vast, flat lowland region of boreal forests (taiga), and wetlands, covering an area about 1,800 km west–east, by 1,000 km north–south.
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Changing climate conditions are amplified in polar regions and northern high-latitude areas are projected to warm at twice the rate of the global average. These modifications result in ecosystem interactions and feedbacks that can augment or mitigate climatic changes. These interactions may have been important through the large climate fluctuations since the glacial period. Therefore it is useful to review the past dynamics of vegetation and climate to place recent observed changes in the Arctic into context. This article focuses on northern Alaska where there has been much research on this theme.
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Sphagnum wulfianum, commonly known as Wulf's peatmoss, is a species of moss belonging to the family Sphagnaceae. It has a circumboreal distribution, occurring primarily in moist boreal forest environments across Eurasia and North America, with rare occurrences in Arctic tundra regions. The species is morphologically distinctive among peat mosses, characterised by having six to twelve branches per fascicle, a unique feature in the genus. First described in 1860 from Estonia, it typically grows in boggy mineral-rich spruce forests and at the borders of mires, forming small carpets and hummocks. While the species has a large geographic range and stable overall population trends in Europe, it faces regional conservation challenges, particularly at its range edges, where it is considered endangered in several countries due to habitat degradation from peatland drainage, forestry, and agriculture. Genetic studies indicate remarkably low genetic diversity throughout its range, suggesting high rates of gene flow across large distances despite infrequent spore production.
The Taiga of North America is a Level I ecoregion of North America designated by the Commission for Environmental Cooperation (CEC) in its North American Environmental Atlas.
The Tundra of North America is a Level I ecoregion of North America designated by the Commission for Environmental Cooperation (CEC) in its North American Environmental Atlas.
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