Corvidae

Last updated

Corvids
Temporal range: Middle Miocene to present
Cyanocitta-cristata-004.jpg
Blue jay
Cyanocitta cristata
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Passeriformes
Superfamily: Corvoidea
Family: Corvidae
Leach, 1820
Subfamilies
Corvidae range.png
Distribution map of the Corvidae.
  Native  (Re)Introduced
  Extinct (post-1500)  Extinct (pre-1500)

Corvidae is a cosmopolitan family of oscine passerine birds that contains the crows, ravens, rooks, magpies, jackdaws, jays, treepies, choughs, and nutcrackers. [1] [2] [3] In colloquial English, they are known as the crow family or corvids. Currently, 139 species are included in this family. The genus Corvus containing 50 species makes up over a third of the entire family. [4] Corvids (ravens) are the largest passerines.

Contents

Corvids display remarkable intelligence for animals of their size, and are among the most intelligent birds thus far studied. [5] Specifically, members of the family have demonstrated self-awareness in mirror tests (Eurasian magpies) and tool-making ability (e.g. crows and rooks [6] ), skills which until recently were thought to be possessed only by humans and a few other higher mammals. Their total brain-to-body mass ratio is equal to that of non-human great apes and cetaceans, and only slightly lower than that of humans. [7]

They are medium to large in size, with strong feet and bills, rictal bristles, and a single moult each year (most passerines moult twice). Corvids are found worldwide, except for the southern tip of South America and the polar ice caps. [3] The majority of the species are found in tropical South and Central America and in southern Asia, with fewer than 10 species each in Africa and Australasia. The genus Corvus has re-entered Australia in relatively recent geological prehistory, with five species and one subspecies there. Several species of raven have reached oceanic islands, and some of these species are now highly threatened with extinction, or have already become extinct.

Systematics, taxonomy, and evolution

The name Corvidae for the family was introduced by the English zoologist William Elford Leach in a guide to the contents of the British Museum published in 1820. [8] [9] Over the years, much disagreement has arisen on the exact evolutionary relationships of the corvid family and their relatives. What eventually seemed clear was that corvids are derived from Australasian ancestors, [10] and spread throughout the world from there. Other lineages derived from these ancestors evolved into ecologically diverse, but often Australasian, groups. In the late 1970s and throughout the 1980s, Sibley and Ahlquist united the corvids with other taxa in the Corvida, based on DNA–DNA hybridization. The presumed corvid relatives included: currawongs, birds of paradise, whipbirds, quail-thrushes, whistlers, monarch flycatchers and drongos, shrikes, vireos, and vangas, [2] but current research favors the theory that this grouping is partly artificial. The corvids constitute the core group of the Corvoidea, together with their closest relatives (the birds of paradise, Australian mud-nesters, and shrikes). They are also the core group of the Corvida, which includes the related groups, such as Old World orioles and vireos. [11]

Crested jayshrikes were thought to be in this family, but may be a type of helmetshrike instead. Haubenhaher1.jpg
Crested jayshrikes were thought to be in this family, but may be a type of helmetshrike instead.

Clarification of the interrelationships of the corvids has been achieved based on cladistic analysis of several DNA sequences. [11] [12] The jays and magpies do not constitute monophyletic lineages, but rather seem to split up into an American and Old World lineage, and an Holarctic and Oriental lineage, respectively. These are not closely related among each other. The position of the azure-winged magpie, which has always been of undistinguished lineage, is less clear than previously thought.

The crested jayshrike (Platylophus galericulatus) is traditionally included in the Corvidae, but is not a true member of this family, being closer to the helmetshrikes (Malaconotidae) or shrikes (Laniidae). [1] [13] Likewise, the Hume's ground "jay" (Pseudopodoces humilis) is, in fact, a member of the tit family, Paridae. [14] The following tree showing the phylogeny of the crow family is based on a molecular study by Jenna McCullough and collaborators published in 2023. [15]

Corvidae

Pyrrhocorax – choughs (2 species)

Crypsirininae

Crypsirina – treepies (2 species)

Dendrocitta – treepies (7 species)

Temnurus – ratchet-tailed treepie

Platysmurus – black magpies (2 species)

Cissinae

Cissa – green magpies (4 species)

Urocissa – blue magpies (5 species)

Perisoreinae

Cyanopica – magpies (2 species)

Perisoreus – grey jays (3 species)

Cyanocoracinae

Cyanolyca – jays (9 species)

Cyanocorax – New World jays (20 species)

Aphelocoma – jays and scrub jays (7 species)

Gymnorhinus – pinyon jay

Cyanocitta – jays (2 species)

Corvinae

Garrulus – Old World jays (3 species)

Ptilostomus – piapiac

Zavattariornis – Stresemann's bushcrow

Podoces – ground jays (4 species)

Pica – magpies (7 species)

Nucifraga – nutcrackers (4 species)

Coloeus – jackdaws (2 species) Eurasian jackdaw.png

Corvus – crows, ravens, rook (50 species) Carrion crow.png

Fossil record

The earliest corvid fossils date to mid-Miocene Europe, [16] about 17 million years ago; Miocorvus and Miopica may be ancestral to crows and some of the magpie lineage, respectively, or similar to the living forms, due to convergent evolution. The known prehistoric corvid genera appear to be mainly of the New World and Old World jay and Holarctic magpie lineages:

In addition, there are numerous fossil species of extant genera since the Mio–Pliocene, mainly European Corvus. [a]

Morphology

Skeleton of American crow (Corvus brachyrhynchos) on display at the Museum of Osteology. American Crow skeleton.jpg
Skeleton of American crow (Corvus brachyrhynchos) on display at the Museum of Osteology.

Corvids are large to very large passerines with a robust build and strong legs; all species, except the pinyon jay, have nostrils covered by bristle-like feathers. [18] Many corvids of temperate zones have mainly black or blue coloured plumage; however, some are pied black and white, some have a blue-purple iridescence, and many tropical species are brightly coloured. The sexes are very similar in color and size. Corvids have strong, stout bills and large wingspans. The family includes the largest members of the passerine order.

The smallest corvid is the dwarf jay (Cyanolyca nanus), at 41 g (1.4 oz) and 21.5 cm (8.5 in). The largest corvids are the common raven (Corvus corax) and the thick-billed raven (Corvus crassirostris), both of which regularly exceed 1,400 grams (3.1 pounds) and 65 cm (26 in).

Species can be identified based on size, shape, and geography; however, some, especially the Australian crows, are best identified by their raucous calls. [2]

Ecology

Corvids occur in most climatic zones. Most are sedentary, and do not migrate significantly. However, during a shortage of food, irruptive migration can occur. [2] When species are migratory, they will form large flocks in the fall (around August in the Northern Hemisphere) and travel south. [19]

One reason for the success of crows, compared to ravens, is their ability to overlap breeding territory. During breeding season, crows were shown to overlap breeding territory six times as much as ravens. This invasion of breeding ranges allowed a related increase in local population density. [20]

Since crows and magpies have benefited and even increased in numbers due to human development, it was suggested that this might cause increased rates of nest predation of smaller bird species, leading to declines. Several studies have shown this concern to be unfounded. One study examined American crows, which had increased in numbers, were a suspect in nest predation of threatened marbled murrelets. However, Steller's jays, which are successful independently of human development, are more efficient in plundering small birds' nests than American crows and common ravens. Therefore, the human relationship with crows and ravens did not significantly increase nest predation when compared to other factors, such as habitat destruction. [20] Similarly, a study examining the decline of British songbirds found no link between Eurasian magpie numbers and population changes of 23 songbird species. [21]

Behaviour

Some corvids have strong organization and community groups. Jackdaws, for example, have a strong social hierarchy, and are facultatively colonial during breeding. [22] Providing mutual aid has also been recorded within many of the corvid species.

Young corvids have been known to play and take part in elaborate social games. Documented group games follow "king of the mountain" or "follow the leader" patterns. Other play involves the manipulation, passing, and balancing of sticks. Corvids also take part in other activities, such as sliding down smooth surfaces. These games are understood to play a large role in the adaptive and survival ability of the birds. [23]

Mate selection is quite complex, and accompanied with much social play in the Corvidae. Youngsters of social corvid species undergo a series of tests, including aerobatic feats, before being accepted as a mate by the opposite sex. [19]

Some corvids can be aggressive. Blue jays, for example, are well known to attack anything that threatens their nest. Crows have been known to attack dogs, cats, ravens, and birds of prey. Most of the time, these assaults take place as a distraction long enough to allow an opportunity for stealing food. [19]

Food and feeding

Corvids are highly opportunistic foragers. Here, a jungle crow feeds on a shark carcass. Raven scavenging on a dead shark.jpg
Corvids are highly opportunistic foragers. Here, a jungle crow feeds on a shark carcass.

The natural diet of many corvid species is omnivorous, consisting of invertebrates, nestlings, small mammals, berries, fruits, seeds, and carrion. However, some corvids, especially the crows, have adapted well to human conditions, and have come to rely on human food sources. In a US study of American crows, common ravens, and Steller's jays around campgrounds and human settlements, the crows appeared to have the most diverse diet of all, taking anthropogenic foods, such as: bread, spaghetti, fried potatoes, dog food, sandwiches, and livestock feed. The increase in available human food sources is contributing to population rises in some corvid species. [20]

Some corvids are predators of other birds. During the wintering months, corvids typically form foraging flocks. [2] However, some crows also eat many agricultural pests, including cutworms, wireworms, grasshoppers, and harmful weeds. [19] Some corvids will eat carrion, and since they lack a specialized beak for tearing into flesh, they must wait until animals are opened, whether by other predators or as roadkill.

Reproduction

A Canada jay pair feeding their chicks. Perisoreus canadensis feeding at nest.jpg
A Canada jay pair feeding their chicks.

Many species of corvid are territorial, protecting territories throughout the year, or simply during the breeding season. In some cases, territories may only be guarded during the day, with the pair joining off-territory roosts at night. Some corvids are well-known communal roosters. Some groups of roosting corvids can be very large, with a roost of 65,000 rooks counted in Scotland. [24] Some, including the rook and the jackdaw, are also communal nesters.

The partner bond in corvids is extremely strong, and even lifelong in some species. This monogamous lifestyle, however, can still contain extra-pair copulations. [25] Males and females build large nests together in trees or on ledges; jackdaws are known to breed in buildings and in rabbit warrens. [22] The male will also feed the female during incubation. [26] The nests are constructed of a mass of bulky twigs lined with grass and bark. Corvids can lay between 3 and 10 eggs, typically ranging between 4 and 7. The eggs are usually greenish in colour with brown blotches. Once hatched, the young remain in the nests for up to 6–10 weeks depending on the species.

Corvids use several different forms of parental care, including bi-parental care and cooperative breeding. [27] Cooperative breeding takes place when parents are helped in raising their offspring, usually by relatives, but also sometimes by non-related adults. [28] [29] Such helpers at the nest in most cooperatively-breeding birds are males, while females join other groups. White-throated magpie-jays are cooperatively-breeding corvids where the helpers are mostly female.

Intelligence

Jerison (1973) has suggested that the degree of brain encephalization (the ratio of brain size to body size, EQ) may correlate with an animal's intelligence and cognitive skills. [30] Corvids and psittacids have higher EQ than other bird families, similar to that of the apes. Among the Corvidae, ravens possess the largest brain to body size ratio. [31] In addition to the high EQ, the Corvid's intelligence is boosted by their living environment. Firstly, Corvids are found in some of the harshest environments on Earth, where surviving requires higher intelligence and better adaptations. Secondly, most of the Corvids are omnivorous, suggesting that they are exposed to a wider variety of different stimuli and environments. Furthermore, many corvid species live in a large family group, and demonstrate high social complexities. [32]

Their intelligence is boosted by the long growing period of the young. [33] [34] [35] By remaining with the parents, the young have more opportunities to learn necessary skills.

When compared to dogs and cats in an experiment testing the ability to seek out food according to three-dimensional clues, corvids out-performed the mammals. [36] A meta-analysis testing how often birds invented new ways to acquire food in the wild found corvids to be the most innovative birds. [37] A 2004 review suggested that their cognitive abilities are on par with those of non-human great apes. [38] Despite structural differences, the brains of corvids and great apes both evolved the ability to make geometrical measurements.

Empathy-consolation

Ravens are found to show bystander affiliation, and solicited bystander affiliation after aggressive conflicts. [39] Most of the time, bystanders already sharing a valuable relationship with the victim are more likely to affiliate with the victim to alleviate the victim's distress ("consolation") as a representation of empathy. Ravens are believed to be able to be sensitive to other's emotions.

Empathy-emotional contagion

Emotion contagion refers to the emotional state matching between individuals. Adriaense et al. (2018) used a bias paradigm to quantify emotional valence, which along with emotional arousal, define emotions. [40] They manipulated the positive and negative affective states in the demonstrator ravens, which showed significantly different responses to the two states: behaving pessimism to the negative states, and optimism to the positive states. Then, the researchers trained another observer raven to first observe the demonstrator's responses. The observer raven was then presented with ambiguous stimuli. The experiment results confirmed the existence of negative emotional contagions in ravens, while the positive emotional contagion remained unclear. Therefore, ravens are capable of both discerning the negative emotions in their conspecifics and showing signs of empathy. [41]

Interspecific communications

Interspecific communications are evolutionarily beneficial for species living in the same environment. Facial expressions are the most widely used method to express emotions by humans. Tate et al. (2006) explored the issue of non-human mammals processing the visual cues from faces to achieve interspecific communication with humans. [42] Researchers also examined the avian species' capabilities to interpret this non-verbal communication, and their extent of sensitivity to human emotions. Based on the experimental subject of American Crows' behavioral changes to varying human gazes and facial expressions, Clucas et al. (2013) identified that crows are able to change their behaviors to the presence of direct human gaze, but did not respond differentially to human emotional facial expressions. They further suggested that the high intelligence of the crows enables them to adapt well to human-dominated environments. [43]

Personality conformity

It is considered difficult to study emotions in animals when humans could not communicate with them. One way to identify animal personality traits is to observe the consistency of the individual's behavior over time and circumstances. [44] [45] For group-living species, there are two opposing hypotheses regarding the assortment of personalities within a group: the social niche specialization hypothesis, and the conformity hypothesis. To test these two hypotheses, McCune et al. (2018) performed an experiment on the boldness of two species in Corvidae: the Mexican Jay and California Scrub-Jay. Their results confirmed the conformity hypothesis, supported by the significant differences in the group effects. [46]

Social construction

The individual personality is both determined by genetics and shaped by social contexts. [47] Miller et al. (2016) examined the role of the developmental and social environment in personality formation in common ravens and carrion crows, which are highly social corvids. The researchers highlighted the correlation between social contexts and an individual's consistent behavior over time (personality), by showing that conspecific presence promoted the behavioral similarities between individuals. Therefore, the researchers demonstrated that social contexts had a significant impact on the development of the raven's and crow's personalities. [48]

Social complexity

The social complexity hypothesis suggests that living in a social group enhances the cognitive abilities of animals. Corvid ingenuity is represented through their feeding skills, memorization abilities, use of tools, and group behaviour. Living in large social groups has long been connected with high cognitive ability. To live in a large group, a member must be able to recognize individuals, and track the social position and foraging of other members over time. Members must also be able to distinguish between sex, age, reproductive status, and dominance, and to update this information constantly. It might be that social complexity corresponds to their high cognition, as well as contributing to the spread of information between members of the group. [49]

Consciousness, culture-rudiments, and neurology

The Eurasian magpie is the only non-mammal species known to be able to recognize itself in a mirror test, [50] although later research could not replicate this finding. [51] Studies using very similar setups could not find such behaviour in other corvids (e.g., Carrion crows [52] [53] ). Magpies have been observed taking part in elaborate grieving rituals, which have been likened to human funerals, including laying grass wreaths. [54] Marc Bekoff, at the University of Colorado, argues that it shows that they are capable of feeling complex emotions, including grief. [54] Furthermore, carrion crows show a neuronal response that correlates with their perception of a stimulus, which some scientists have argued to be an empirical marker of (avian/corvid) sensory consciousness—the conscious perception of sensory input—in the crows which do not have a cerebral cortex. [55] [56] A related study shows that the birds' pallium's neuroarchitecture is reminiscent of the mammalian cortex. [57] [58]

Tool use, memory, and complex rational thought

A New Caledonian crow uses a tool to retrieve the correct tool to obtain food.

There are also specific examples of corvid cleverness. One carrion crow was documented cracking nuts by placing them on a crosswalk, letting the passing cars crack the shell, waiting for the light to turn red, and then safely retrieving the contents. [59] A group of crows in England took turns lifting garbage bin lids while their companions collected food.

Members of the corvid family have been known to watch other birds, remember where they hide their food, then return once the owner leaves. [60] [61] Corvids also move their food around between hiding places to avoid thievery—but only if they have previously been thieves themselves (that is, they remember previous relevant social contexts, use their own experience of having been a thief to predict the behavior of a pilferer, and can determine the safest course to protect their caches from being pilfered). Studies to assess similar cognitive abilities in apes have been inconclusive. [62]

The ability to hide food requires highly accurate spatial memories. Corvids have been recorded to recall their food's hiding places up to nine months later. It is suggested that vertical landmarks (like trees) are used to remember locations. There has also been evidence that California scrub jays, which store perishable foods, not only remember where they stored their food, but for how long. This has been compared to episodic memory, previously thought unique to humans. [3]

New Caledonian crows (Corvus moneduloides) are notable for their highly developed tool fabrication. They make angling tools of twigs and leaves trimmed into hooks, and then subsequently use the hooks to pull insect larvae from tree holes. Tools are engineered according to task, and apparently, also to learned preferences. Recent studies revealed abilities to solve complicated problems, which suggested high levels of innovation of a complex nature. [63] [64] Other corvids that have been observed using tools include: the American crow, blue jay, and green jay. Researchers have discovered that New Caledonian crows do not just use single objects as tools—they can also construct novel compound tools through assemblage of otherwise non-functional elements. [65] [66] Diversity in tool design among corvids suggests cultural variation. Again, great apes are the only other animals known to use tools in such a fashion. [3]

Clark's nutcrackers and jackdaws were compared in a 2002 study based on geometric rule learning. The corvids, along with a domestic pigeon, had to locate a target between two landmarks, while distances and landmarks were altered. The nutcrackers were more accurate in their searches than the jackdaws and pigeons. [67]

Implications and specific comparisons with other animals

Proportions of correct responses as a function of species (R: ravens; C: chimpanzees; O: orang-utans) Proportions of correct responses as a function of species.webp
Proportions of correct responses as a function of species (R: ravens; C: chimpanzees; O: orang-utans)

The scarecrow is an archetypal scare tactic in the agricultural business. However, due to corvids' quick wit, scarecrows are soon ignored, and used as perches. Despite farmers' efforts to rid themselves of corvid pests, their attempts have only expanded corvid territories, and strengthened their numbers. [19]

Contrary to earlier teleological classifications, in which they were seen as "highest" songbirds due to their intelligence, current systematics might place corvids—based on their total number of physical characteristics, instead of just their brains (which are the most developed of birds)—in the lower middle of the passerine evolutionary tree, dependent on which subgroup is chosen as the most derived. [11] As per one observer:

During the 19th century, there arose the belief that these were the "most advanced" birds, based upon the belief that Darwinian evolution brings "progress." In such a classification, the "most intelligent" of birds were listed last, reflecting their position "atop the pyramid." Modern biologists reject the concept of hierarchical "progress" in evolution [...]. [2]

The other major group of highly intelligent birds of the order Psittaciformes (which includes 'true' parrots, cockatoos, and New Zealand parrots) is not closely related to corvids.

A study found that four-month-old ravens can have physical and social cognitive skills similar to that of adult great apes, and concluded that the "dynamic of the different influences that, during ontogeny, contributes to adult cognition" is required for the study of cognition. [69] [68]

Disease

Corvids are reservoirs (carriers) for the West Nile virus in the United States. They are infected by mosquitoes (the vectors), primarily of the Culex species. Crows and ravens are quickly killed by this disease, so their deaths are an early-warning system when West Nile virus arrives in an area (as are horses and other bird-species deaths). One of the first signs that West Nile virus first arrived in the US in 1999 was the death of crows in New York. [70]

Relationship with humans

Several different corvids, particularly ravens, have occasionally served as pets, although they are not able to speak as readily as parrots, and are not suited to a caged environment.

It is illegal to own corvids, or any other migratory bird, without a permit in North America, due to the Migratory Bird Act.

Humans have been able to coexist with many members of the Corvidae family throughout history, most notably crows and ravens (see: "Role in myth and culture" section below). These positive interactions have extended into modern times.

Role in myth and culture

Folklore often represents corvids as clever, and even mystical, animals. Some Native Americans, such as the Haida, believed that a raven created the earth, and despite being a trickster spirit, ravens were popular on totems, credited with creating man, and considered responsible for placing the Sun in the sky. [71]

Due to their carrion diet, the Celtic peoples strongly associated corvids with war, death, and the battlefield; their great intelligence meant that they were often considered messengers, or manifestations of the gods, such as Bendigeidfran (Welsh for "Blessed Crow") or the Irish Morrigan (Middle Irish for "Great Queen"), [72] both who were underworld deities that may be related to the later Arthurian Fisher King. The Welsh Dream of Rhonabwy illustrates well the association of ravens with war. In many parts of Britain, gatherings of crows, or more often magpies, are counted using the divination rhyme: "one for sorrow, two for joy, three for a girl, four for a boy, five for silver, six for gold, seven for a secret never to be told." Another rhyme is: "one for sorrow, two for mirth, three for a funeral, four for a birth, five for heaven, six for hell, and seven for the Devil, his own sel." Cornish superstition holds that when a lone magpie is encountered, it must be loudly greeted with respect.

Various Germanic peoples highly revered the raven, and the raven was often depicted as a motif on shields or other war gear in Anglo-Saxon art, such as the Sutton Hoo burial, and Vendel period art. The major deity, Odin, was so commonly associated with ravens throughout history that he gained the kenning "Raven God," [b] and the raven banner was the flag of various Viking Age Scandinavian chieftains. Odin was also attended by Hugin and Munin, two ravens who flew all over the world, and whispered information they acquired into his ears. [73] The Valravn sometimes appeared in modern Scandinavian folklore. On a shield and purse lid excavated among the Sutton Hoo treasures, imagery of stylised corvids with scrolled beaks are meticulously detailed in the decorative enamel work. The corvid symbolism reflected their common totemic status to the Anglo-Saxons, whose pre-Christian indigenous beliefs were of the same origin as that of the aforementioned Vikings.

The sixth century BCE Greek scribe Aesop featured corvids as intelligent antagonists in many fables. Later, in western literature, popularized by American poet Edgar Allan Poe's work "The Raven", the common raven becomes a symbol of the main character's descent into madness.

The children's book Mrs. Frisby and the Rats of NIMH and its animated film adaptation features a protagonist crow named Jeremy.

The Hawaiian crow is extinct in the wild as a result of habitat loss and other factors. Corvus hawaiiensis FWS.jpg
The Hawaiian crow is extinct in the wild as a result of habitat loss and other factors.

Status and conservation

Unlike many other bird families, corvid fitness and reproduction, especially with many crows, has increased due to human development. The survival and reproductive success of certain crows and ravens is assisted by their close relationship with humans. [20]

Human development provides additional resources by clearing land, creating shrublands rich in berries and insects. When the cleared land naturally replenishes, jays and crows use the young dense trees for nesting sites. Ravens typically use larger trees in denser forest. [20]

Most corvids are not threatened, and many species are even increasing in population due to human activity. However, a few species are in danger. For example, the destruction of the Southeast Asian rainforest is endangering mixed-species feeding flocks with members from the family Corvidae. [74] Also, since its semiarid scrubland habitat is an endangered ecosystem, the Florida scrub jay has a small and declining population. [75] [76] A number of island species, which are more vulnerable to introduced species and habitat loss, have been driven to extinction, such as the New Zealand raven, or are threatened, like the Mariana crow.

The American crow population of the United States has grown over the years. It is possible that the American crow, due to humans increasing suitable habitat, will cause Northwestern crows and fish crows to decline. [77]

Species

FAMILY CORVIDAE

Rufous treepie, Dendrocitta vagabunda Rufous Treepie I IMG 9850.jpg
Rufous treepie, Dendrocitta vagabunda
Yellow-billed blue magpie, Urocissa flavirostris Yellow-billed Blue Magpie I IMG 7393.jpg
Yellow-billed blue magpie, Urocissa flavirostris
Eurasian jay (Garrulus glandarius) Orvani (1).JPG
Eurasian jay (Garrulus glandarius)
Eurasian magpie, Pica pica Elster wikipedia2.jpg
Eurasian magpie, Pica pica
Plush-crested jay, Cyanocorax chrysops Plush-crested Jay.jpg
Plush-crested jay, Cyanocorax chrysops
Common raven, Corvus corax Corvus corax ad berlin 090516.jpg
Common raven, Corvus corax
Hooded crow, Corvus cornix Wrona Siwa.jpg
Hooded crow, Corvus cornix
Thick-billed raven, Corvus crassirostris Flickr - don macauley - Corvus crassirostris.jpg
Thick-billed raven, Corvus crassirostris
Australian raven, Corvus coronoides Corvus coronoides.jpg
Australian raven, Corvus coronoides

Explanatory notes

  1. ↑ See the genus accounts for more.
  2. ↑ E.g. Icelandic: hrafnaguð, as per the Gylfaginning .

Related Research Articles

<span class="mw-page-title-main">Magpie</span> Large bird in the corvid family

Magpies are birds of various species of the family Corvidae. Like other members of their family, they are widely considered to be intelligent creatures. The Eurasian magpie, for instance, is thought to rank among the world's most intelligent creatures, and is one of the few nonmammalian species able to recognize itself in a mirror test. Magpies have shown the ability to make and use tools, imitate human speech, grieve, play games, and work in teams. They are particularly well known for their songs and were once popular as cagebirds. In addition to other members of the genus Pica, corvids considered magpies are in the genera Cissa, Urocissa, and Cyanopica.

<span class="mw-page-title-main">Eurasian magpie</span> Species of bird

The Eurasian magpie or common magpie is a resident breeding bird throughout the northern part of the Eurasian continent. It is one of several birds in the crow family (corvids) designated magpies, and belongs to the Holarctic radiation of "monochrome" magpies. In Europe, "magpie" is used by English speakers as a synonym for the Eurasian magpie: the only other magpie in Europe is the Iberian magpie, which is limited to the Iberian Peninsula. Despite having a shared name and similar colouration, it is not closely related to the Australian magpie.

<span class="mw-page-title-main">Carrion crow</span> Species of bird

The carrion crow is a passerine bird of the family Corvidae, native to western Europe and the eastern Palearctic.

<span class="mw-page-title-main">Australian raven</span> Passerine bird native to Australia

The Australian raven is a passerine corvid bird native to Australia. Measuring 46–53 centimetres (18–21 in) in length, it has an all-black plumage, beak and mouth, as well as strong, greyish-black legs and feet. The upperparts of its body are glossy, with a purple-blue, greenish sheen; its black feathers have grey bases. The Australian raven is distinguished from the Australian crow, and other related corvids, by its long chest feathers, or throat hackles, which are prominent in mature birds. Older individuals and subadults have white irises, while the younger birds' eyes display blue inner rims; hatchlings and young birds have brown, dark irises until about fifteen months of age, at which point their irises become hazel-coloured, with an inner blue rim around each pupil, this lasting until they are roughly 2.5 to 3 years of age. Nicholas Aylward Vigors and Thomas Horsfield described the Australian raven in 1827, its species name coronoides highlighting its similarity with the carrion crow. Two subspecies are recognised, which differ slightly in their vocalisations, and are quite divergent, genetically.

<span class="mw-page-title-main">Torresian crow</span> Species of bird

The Torresian crow, also called the Australian crow or Papuan crow, is a passerine bird in the crow family native to the north and west of Australia and nearby islands in Indonesia and Papua New Guinea. The species has a black plumage, beak and mouth with white irises. The base of the feathers on the head and neck are white. The Torresian crow is slightly larger with a more robust bill than the morphologically similar little crow.

<span class="mw-page-title-main">Little raven</span> Species of bird

The little raven is a species of the family Corvidae that is native to southeastern Australia. An adult individual is about 48–50 cm (19–19.5 in) in length, with completely black plumage, beak, and legs; as with all Australian species of Corvus, the black feathers have a grey base, and the iris of the adult bird is white. Although the little raven was first named by Gregory Mathews in 1912, it was only in 1967 that there was consensus to separate it from the Australian raven as a distinct species.

<span class="mw-page-title-main">Western jackdaw</span> Species of bird in the crow family Corvidae

The western jackdaw, also known as the Eurasian jackdaw, the European jackdaw, or simply the jackdaw, is a passerine bird in the crow family. Found across Europe, western Asia and North Africa; it is mostly resident, although northern and eastern populations migrate south in the winter. Four subspecies are recognised, which differ mainly in the colouration of the plumage on the head and nape. Linnaeus first described it formally, giving it the name Corvus monedula. The common name derives from the word jack, denoting "small", and daw, a less common synonym for "jackdaw", and the native English name for the bird.

<span class="mw-page-title-main">American crow</span> Species of bird

The American crow is a large passerine bird species of the family Corvidae. It is a common bird found throughout much of North America. American crows are the New World counterpart to the carrion crow and the hooded crow of Eurasia; they all occupy the same ecological niche. Although the American crow and the hooded crow are very similar in size, structure and behavior, their calls and visual appearance are different.

<i>Aphelocoma</i> Genus of birds

The passerine birds of the genus Aphelocoma include the scrub jays and their relatives. They are New World jays found in Mexico, western Central America and the western United States, with an outlying population in Florida. This genus belongs to the group of New World jays—possibly a distinct subfamily–which is not closely related to other jays, magpies or treepies. Within this group, according to a 2023 molecular analysis, Aphelocoma is the sister group to a clade consisting of Cyanocitta and Gymnorhinus. They live in open pine-oak forests, chaparral, and mixed evergreen forests.

<span class="mw-page-title-main">Rufous treepie</span> Species of bird

The rufous treepie is a treepie, native to the Indian Subcontinent and adjoining parts of Southeast Asia. It is a member of the crow family, Corvidae. It is long tailed and has loud musical calls making it very conspicuous. It is found commonly in open scrub, agricultural areas, forests as well as urban gardens. Like other corvids it is very adaptable, omnivorous and opportunistic in feeding.

<i>Crypsirina</i> Genus of birds

Crypsirina is a small genus of long-tailed passerine birds in the crow and jay family, Corvidae. The two species are highly arboreal and rarely come to the ground to feed. The generic name is derived from the Greek words kruptō, meaning "to conceal," and rhis or rhinos, meaning "nostrils".

<i>Cyanocorax</i> Genus of birds

Cyanocorax is a genus of New World jays, passerine birds in the family Corvidae. It contains several closely related species that primarily are found in wooded habitats, chiefly in lowland tropical rainforest but in some cases also in seasonally dry forest, grassland and montane forest. They occur from Mexico through Central into southern South America, with the green jay and brown jay just entering the United States in southernmost Texas, ad the Azure and Plush-crested jays occurring southwards to the lower Paraná River basin. This genus is considered especially close to Cyanolyca, an upland radiation occurring throughout the American Cordillera from Mexico to Peru and Bolivia, who look very similar to the blue-and-black species of Cyanocorax except for being a bit smaller. The North American blue jay genera Aphelocoma, Cyanocitta and Gymnorhinus seem to be slightly less closely related.

<i>Pica</i> (genus) Genus of birds

Pica is a genus of seven species of birds in the family Corvidae in both the New World and the Old. It is one of several corvid genera whose members are known as magpies.

<span class="mw-page-title-main">Azure jay</span> Species of bird

The azure jay is a passeriform bird of the crow family, Corvidae. It is found in the Atlantic Forest, especially with Araucaria angustifolia, in south-eastern Brazil, far eastern Paraguay and far north-eastern Argentina. It is the state bird of Paraná.

<span class="mw-page-title-main">Bird intelligence</span> Study of intelligence in birds

The difficulty of defining or measuring intelligence in non-human animals makes the subject difficult to study scientifically in birds. In general, birds have relatively large brains compared to their head size. Furthermore, bird brains have two-to-four times the neuron packing density of mammal brains, for higher overall efficiency. The visual and auditory senses are well developed in most species, though the tactile and olfactory senses are well realized only in a few groups. Birds communicate using visual signals as well as through the use of calls and song. The testing of intelligence in birds is therefore usually based on studying responses to sensory stimuli.

<span class="mw-page-title-main">White-bellied treepie</span> Species of bird

The white-bellied treepie is a bird of the crow family endemic to the forests of southern India. They overlap in distribution in some areas with the rufous treepie but are easy to tell apart both from appearance and call.

<i>Corvus</i> Genus of birds including crows, ravens and rooks

Corvus is a widely distributed genus of passerine birds ranging from medium-sized to large-sized in the family Corvidae. It includes species commonly known as crows, ravens, and rooks. The species commonly encountered in Europe are the carrion crow, hooded crow, common raven, and rook; those discovered later were named "crow" or "raven" chiefly on the basis of their size, crows generally being smaller. The genus name is Latin for "raven".

<span class="mw-page-title-main">Oriental magpie</span> Species of bird

The Oriental magpie is a species of magpie found from south-eastern Russia to eastern China, Korea, Taiwan, Japan and northern Indochina and Myanmar. Other names for the Oriental magpie include Korean magpie and Asian magpie.

<span class="mw-page-title-main">Rook (bird)</span> Species of bird in the crow family Corvidae

The rook is a member of the family Corvidae in the passerine order of birds. It is found in the Palearctic, its range extending from Scandinavia and western Europe to eastern Siberia. It is a large, gregarious, black-feathered bird, distinguished from similar species by the whitish featherless area on the face. Rooks nest collectively in the tops of tall trees, often close to farms or villages; the groups of nests are known as rookeries.

<span class="mw-page-title-main">Jackdaw</span> Two species of bird

Jackdaws are two species of bird in the genus Coloeus closely related to, but generally smaller than, crows and ravens (Corvus). They have a blackish crown, wings, and tail, with the rest of their plumage paler. The word Coloeus is Neo-Latin, from the Ancient Greek for jackdaws: koloiós (κολοιός). They come from Asia, Europe, Africa and Siberia.

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