An ecosystem is a community of living organisms in conjunction with the nonliving components of their environment, interacting as a system.These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the system through photosynthesis and is incorporated into plant tissue. By feeding on plants and on one-another, animals play an important role in the movement of matter and energy through the system. They also influence the quantity of plant and microbial biomass present. By breaking down dead organic matter, decomposers release carbon back to the atmosphere and facilitate nutrient cycling by converting nutrients stored in dead biomass back to a form that can be readily used by plants and other microbes.
In ecology, a community is a group or association of populations of two or more different species occupying the same geographical area and in a particular time, also known as a biocoenosis. The term community has a variety of uses. In its simplest form it refers to groups of organisms in a specific place or time, for example, "the fish community of Lake Ontario before industrialization".
In biology and ecology, abiotic components or abiotic factors are non-living chemical and physical parts of the environment that affect living organisms and the functioning of ecosystems. Abiotic factors and the phenomena associated with them underpin all biology.
Biotic components or biotic factors, can be described as any living component that affects another organism, or shapes the ecosystem. This includes both animals that consume other organisms within their ecosystem, and the organism that is being consumed. Biotic factors also include human influence, pathogens, and disease outbreaks. Each biotic factor needs the proper amount of energy and nutrition to function day to day.
Ecosystems are controlled by external and internal factors. External factors such as climate, parent material which forms the soil and topography, control the overall structure of an ecosystem but are not themselves influenced by the ecosystem.Unlike external factors, internal factors are controlled, for example, decomposition, root competition, shading, disturbance, succession, and the types of species present.
Climate is defined as the average state of everyday's weather condition over a period of 30 years. It is measured by assessing the patterns of variation in temperature, humidity, atmospheric pressure, wind, precipitation, atmospheric particle count and other meteorological variables in a given region over long periods of time. Climate differs from weather, in that weather only describes the short-term conditions of these variables in a given region.
Parent material is the underlying geological material in which soil horizons form. Soils typically inherit a great deal of structure and minerals from their parent material, and, as such, are often classified based upon their contents of consolidated or unconsolidated mineral material that has undergone some degree of physical or chemical weathering and the mode by which the materials were most recently transported.
Topography is the study of the shape and features of land surfaces. The topography of an area could refer to the surface shapes and features themselves, or a description.
Ecosystems are dynamic entities—they are subject to periodic disturbances and are in the process of recovering from some past disturbance.Ecosystems in similar environments that are located in different parts of the world can end up doing things very differently simply because they have different pools of species present. Internal factors not only control ecosystem processes but are also controlled by them and are often subject to feedback loops.
Feedback occurs when outputs of a system are routed back as inputs as part of a chain of cause-and-effect that forms a circuit or loop. The system can then be said to feed back into itself. The notion of cause-and-effect has to be handled carefully when applied to feedback systems:
Simple causal reasoning about a feedback system is difficult because the first system influences the second and second system influences the first, leading to a circular argument. This makes reasoning based upon cause and effect tricky, and it is necessary to analyze the system as a whole.
Resource inputs are generally controlled by external processes like climate and parent material. Resource availability within the ecosystem is controlled by internal factors like decomposition, root competition or shading.Although humans operate within ecosystems, their cumulative effects are large enough to influence external factors like climate.
Biodiversity affects ecosystem functioning, as do the processes of disturbance and succession. Ecosystems provide a variety of goods and services upon which people depend.
Biodiversity refers to the variety and variability of life on Earth. Biodiversity is typically a measure of variation at the genetic, species, and ecosystem level. Terrestrial biodiversity is usually greater near the equator, which is the result of the warm climate and high primary productivity. Biodiversity is not distributed evenly on Earth, and is richest in the tropics. These tropical forest ecosystems cover less than 10 percent of earth's surface, and contain about 90 percent of the world's species. Marine biodiversity is usually highest along coasts in the Western Pacific, where sea surface temperature is highest, and in the mid-latitudinal band in all oceans. There are latitudinal gradients in species diversity. Biodiversity generally tends to cluster in hotspots, and has been increasing through time, but will be likely to slow in the future.
In ecology, a disturbance is a temporary change in environmental conditions that causes a pronounced change in an ecosystem. Disturbances often act quickly and with great effect, to alter the physical structure or arrangement of biotic and abiotic elements. Disturbance can also occur over a long period of time and can impact the biodiversity within an ecosystem. Major ecological disturbances may include fires, flooding, storms, insect outbreaks and trampling. Earthquakes, various types of volcanic eruptions, tsunami, firestorms, impact events, climate change, and the devastating effects of human impact on the environment such as clearcutting, forest clearing and the introduction of invasive species can be considered major disturbances. Not only invasive species can have a profound effect on an ecosystem, but also naturally occurring species can cause disturbance by their behavior. Disturbance forces can have profound immediate effects on ecosystems and can, accordingly, greatly alter the natural community. Because of these and the impacts on populations, disturbance determines the future shifts in dominance, various species successively becoming dominant as their life history characteristics, and associated life-forms, are exhibited over time.
Ecological succession is the process of change in the species structure of an ecological community over time. The time scale can be decades, or even millions of years after a mass extinction.
The term ecosystem was first used in 1935 in a publication by British ecologist Arthur Tansley.Tansley devised the concept to draw attention to the importance of transfers of materials between organisms and their environment. He later refined the term, describing it as "The whole system, ... including not only the organism-complex, but also the whole complex of physical factors forming what we call the environment". Tansley regarded ecosystems not simply as natural units, but as "mental isolates". Tansley later defined the spatial extent of ecosystems using the term ecotope.
Sir Arthur George Tansley FLS, FRS was an English botanist and a pioneer in the science of ecology.
Ecotopes are the smallest ecologically distinct landscape features in a landscape mapping and classification system. As such, they represent relatively homogeneous, spatially explicit landscape functional units that are useful for stratifying landscapes into ecologically distinct features for the measurement and mapping of landscape structure, function and change.
G. Evelyn Hutchinson, a limnologist who was a contemporary of Tansley's, combined Charles Elton's ideas about trophic ecology with those of Russian geochemist Vladimir Vernadsky. As a result, he suggested that mineral nutrient availability in a lake limited algal production. This would, in turn, limit the abundance of animals that feed on algae. Raymond Lindeman took these ideas further to suggest that the flow of energy through a lake was the primary driver of the ecosystem. Hutchinson's students, brothers Howard T. Odum and Eugene P. Odum, further developed a "systems approach" to the study of ecosystems. This allowed them to study the flow of energy and material through ecological systems.
Ecosystems are controlled both by external and internal factors. External factors, also called state factors, control the overall structure of an ecosystem and the way things work within it, but are not themselves influenced by the ecosystem. The most important of these is climate.Climate determines the biome in which the ecosystem is embedded. Rainfall patterns and seasonal temperatures influence photosynthesis and thereby determine the amount of water and energy available to the ecosystem.
Parent material determines the nature of the soil in an ecosystem, and influences the supply of mineral nutrients. Topography also controls ecosystem processes by affecting things like microclimate, soil development and the movement of water through a system. For example, ecosystems can be quite different if situated in a small depression on the landscape, versus one present on an adjacent steep hillside.
Other external factors that play an important role in ecosystem functioning include time and potential biota. Similarly, the set of organisms that can potentially be present in an area can also significantly affect ecosystems. Ecosystems in similar environments that are located in different parts of the world can end up doing things very differently simply because they have different pools of species present.The introduction of non-native species can cause substantial shifts in ecosystem function.
Unlike external factors, internal factors in ecosystems not only control ecosystem processes but are also controlled by them. Consequently, they are often subject to feedback loops. [ citation needed ]While the resource inputs are generally controlled by external processes like climate and parent material, the availability of these resources within the ecosystem is controlled by internal factors like decomposition, root competition or shading. Other factors like disturbance, succession or the types of species present are also internal factors.
Primary production is the production of organic matter from inorganic carbon sources. This mainly occurs through photosynthesis. The energy incorporated through this process supports life on earth, while the carbon makes up much of the organic matter in living and dead biomass, soil carbon and fossil fuels. It also drives the carbon cycle, which influences global climate via the greenhouse effect.
Through the process of photosynthesis, plants capture energy from light and use it to combine carbon dioxide and water to produce carbohydrates and oxygen. The photosynthesis carried out by all the plants in an ecosystem is called the gross primary production (GPP).About half of the GPP is consumed in plant respiration. The remainder, that portion of GPP that is not used up by respiration, is known as the net primary production (NPP). Total photosynthesis is limited by a range of environmental factors. These include the amount of light available, the amount of leaf area a plant has to capture light (shading by other plants is a major limitation of photosynthesis), rate at which carbon dioxide can be supplied to the chloroplasts to support photosynthesis, the availability of water, and the availability of suitable temperatures for carrying out photosynthesis.
Energy and carbon enter ecosystems through photosynthesis, are incorporated into living tissue, transferred to other organisms that feed on the living and dead plant matter, and eventually released through respiration.
The carbon and energy incorporated into plant tissues (net primary production) is either consumed by animals while the plant is alive, or it remains uneaten when the plant tissue dies and becomes detritus. In terrestrial ecosystems, roughly 90% of the net primary production ends up being broken down by decomposers. The remainder is either consumed by animals while still alive and enters the plant-based trophic system, or it is consumed after it has died, and enters the detritus-based trophic system.[ citation needed ]
In aquatic systems, the proportion of plant biomass that gets consumed by herbivores is much higher.In trophic systems photosynthetic organisms are the primary producers. The organisms that consume their tissues are called primary consumers or secondary producers—herbivores. Organisms which feed on microbes (bacteria and fungi) are termed microbivores. Animals that feed on primary consumers—carnivores—are secondary consumers. Each of these constitutes a trophic level.
The sequence of consumption—from plant to herbivore, to carnivore—forms a food chain. Real systems are much more complex than this—organisms will generally feed on more than one form of food, and may feed at more than one trophic level. Carnivores may capture some prey which are part of a plant-based trophic system and others that are part of a detritus-based trophic system (a bird that feeds both on herbivorous grasshoppers and earthworms, which consume detritus). Real systems, with all these complexities, form food webs rather than food chains.The food chain usually consist of four level of consumption which are producers, primary consumers, secondary consumers, and tertiary consumers.
The carbon and nutrients in dead organic matter are broken down by a group of processes known as decomposition. This releases nutrients that can then be re-used for plant and microbial production and returns carbon dioxide to the atmosphere (or water) where it can be used for photosynthesis. In the absence of decomposition, the dead organic matter would accumulate in an ecosystem, and nutrients and atmospheric carbon dioxide would be depleted.Approximately 90% of terrestrial net primary production goes directly from plant to decomposer.
Decomposition processes can be separated into three categories—leaching, fragmentation and chemical alteration of dead material. As water moves through dead organic matter, it dissolves and carries with it the water-soluble components. These are then taken up by organisms in the soil, react with mineral soil, or are transported beyond the confines of the ecosystem (and are considered lost to it).Newly shed leaves and newly dead animals have high concentrations of water-soluble components and include sugars, amino acids and mineral nutrients. Leaching is more important in wet environments and much less important in dry ones.
Fragmentation processes break organic material into smaller pieces, exposing new surfaces for colonization by microbes. Freshly shed leaf litter may be inaccessible due to an outer layer of cuticle or bark, and cell contents are protected by a cell wall. Newly dead animals may be covered by an exoskeleton. Fragmentation processes, which break through these protective layers, accelerate the rate of microbial decomposition.Animals fragment detritus as they hunt for food, as does passage through the gut. Freeze-thaw cycles and cycles of wetting and drying also fragment dead material.
The chemical alteration of the dead organic matter is primarily achieved through bacterial and fungal action. Fungal hyphae produce enzymes which can break through the tough outer structures surrounding dead plant material. They also produce enzymes which break down lignin, which allows them access to both cell contents and to the nitrogen in the lignin. Fungi can transfer carbon and nitrogen through their hyphal networks and thus, unlike bacteria, are not dependent solely on locally available resources.
Decomposition rates vary among ecosystems.The rate of decomposition is governed by three sets of factors—the physical environment (temperature, moisture, and soil properties), the quantity and quality of the dead material available to decomposers, and the nature of the microbial community itself. Temperature controls the rate of microbial respiration; the higher the temperature, the faster microbial decomposition occurs. It also affects soil moisture, which slows microbial growth and reduces leaching. Freeze-thaw cycles also affect decomposition—freezing temperatures kill soil microorganisms, which allows leaching to play a more important role in moving nutrients around. This can be especially important as the soil thaws in the spring, creating a pulse of nutrients which become available.
Decomposition rates are low under very wet or very dry conditions. Decomposition rates are highest in wet, moist conditions with adequate levels of oxygen. Wet soils tend to become deficient in oxygen (this is especially true in wetlands), which slows microbial growth. In dry soils, decomposition slows as well, but bacteria continue to grow (albeit at a slower rate) even after soils become too dry to support plant growth.
Ecosystems continually exchange energy and carbon with the wider environment. Mineral nutrients, on the other hand, are mostly cycled back and forth between plants, animals, microbes and the soil. Most nitrogen enters ecosystems through biological nitrogen fixation, is deposited through precipitation, dust, gases or is applied as fertilizer.
Since most terrestrial ecosystems are nitrogen-limited, nitrogen cycling is an important control on ecosystem production.
Until modern times, nitrogen fixation was the major source of nitrogen for ecosystems. Nitrogen-fixing bacteria either live symbiotically with plants or live freely in the soil. The energetic cost is high for plants which support nitrogen-fixing symbionts—as much as 25% of gross primary production when measured in controlled conditions. Many members of the legume plant family support nitrogen-fixing symbionts. Some cyanobacteria are also capable of nitrogen fixation. These are phototrophs, which carry out photosynthesis. Like other nitrogen-fixing bacteria, they can either be free-living or have symbiotic relationships with plants.Other sources of nitrogen include acid deposition produced through the combustion of fossil fuels, ammonia gas which evaporates from agricultural fields which have had fertilizers applied to them, and dust. Anthropogenic nitrogen inputs account for about 80% of all nitrogen fluxes in ecosystems.
When plant tissues are shed or are eaten, the nitrogen in those tissues becomes available to animals and microbes. Microbial decomposition releases nitrogen compounds from dead organic matter in the soil, where plants, fungi, and bacteria compete for it. Some soil bacteria use organic nitrogen-containing compounds as a source of carbon, and release ammonium ions into the soil. This process is known as nitrogen mineralization. Others convert ammonium to nitrite and nitrate ions, a process known as nitrification. Nitric oxide and nitrous oxide are also produced during nitrification.Under nitrogen-rich and oxygen-poor conditions, nitrates and nitrites are converted to nitrogen gas, a process known as denitrification.
Other important nutrients include phosphorus, sulfur, calcium, potassium, magnesium and manganese.Phosphorus enters ecosystems through weathering. As ecosystems age this supply diminishes, making phosphorus-limitation more common in older landscapes (especially in the tropics). Calcium and sulfur are also produced by weathering, but acid deposition is an important source of sulfur in many ecosystems. Although magnesium and manganese are produced by weathering, exchanges between soil organic matter and living cells account for a significant portion of ecosystem fluxes. Potassium is primarily cycled between living cells and soil organic matter.
Biodiversity plays an important role in ecosystem functioning. [ citation needed ]The reason for this is that ecosystem processes are driven by the number of species in an ecosystem, the exact nature of each individual species, and the relative abundance organisms within these species. Ecosystem processes are broad generalizations that actually take place through the actions of individual organisms. The nature of the organisms—the species, functional groups and trophic levels to which they belong—dictates the sorts of actions these individuals are capable of carrying out and the relative efficiency with which they do so.
Ecological theory suggests that in order to coexist, species must have some level of limiting similarity—they must be different from one another in some fundamental way, otherwise one species would competitively exclude the other.Despite this, the cumulative effect of additional species in an ecosystem is not linear—additional species may enhance nitrogen retention, for example, but beyond some level of species richness, additional species may have little additive effect.
The addition (or loss) of species which are ecologically similar to those already present in an ecosystem tends to only have a small effect on ecosystem function. Ecologically distinct species, on the other hand, have a much larger effect. Similarly, dominant species have a large effect on ecosystem function, while rare species tend to have a small effect. Keystone species tend to have an effect on ecosystem function that is disproportionate to their abundance in an ecosystem. [ citation needed ]Similarly, an ecosystem engineer is any organism that creates, significantly modifies, maintains or destroys a habitat.
Ecosystems are dynamic entities. They are subject to periodic disturbances and are in the process of recovering from some past disturbance.When a perturbation occurs, an ecosystem responds by moving away from its initial state. The tendency of an ecosystem to remain close to its equilibrium state, despite that disturbance, is termed its resistance. On the other hand, the speed with which it returns to its initial state after disturbance is called its resilience. Time plays a role in the development of soil from bare rock and the recovery of a community from disturbance.
From one year to another, ecosystems experience variation in their biotic and abiotic environments. A drought, a colder than usual winter, and a pest outbreak all are short-term variability in environmental conditions. Animal populations vary from year to year, building up during resource-rich periods and crashing as they overshoot their food supply. These changes play out in changes in net primary production decomposition rates, and other ecosystem processes.Longer-term changes also shape ecosystem processes—the forests of eastern North America still show legacies of cultivation which ceased 200 years ago, while methane production in eastern Siberian lakes is controlled by organic matter which accumulated during the Pleistocene.
Disturbance also plays an important role in ecological processes. F. Stuart Chapin and coauthors define disturbance as "a relatively discrete event in time and space that alters the structure of populations, communities, and ecosystems and causes changes in resources availability or the physical environment".This can range from tree falls and insect outbreaks to hurricanes and wildfires to volcanic eruptions. Such disturbances can cause large changes in plant, animal and microbe populations, as well soil organic matter content. Disturbance is followed by succession, a "directional change in ecosystem structure and functioning resulting from biotically driven changes in resources supply."
The frequency and severity of disturbance determine the way it affects ecosystem function. A major disturbance like a volcanic eruption or glacial advance and retreat leave behind soils that lack plants, animals or organic matter. Ecosystems that experience such disturbances undergo primary succession. A less severe disturbance like forest fires, hurricanes or cultivation result in secondary succession and a faster recovery.More severe disturbance and more frequent disturbance result in longer recovery times.
Ecosystem ecology studies the processes and dynamics of ecosystems, and the way the flow of matter and energy through them structures natural systems. The study of ecosystems can cover 10 orders of magnitude, from the surface layers of rocks to the surface of the planet.
There is no single definition of what constitutes an ecosystem.German ecologist Ernst-Detlef Schulze and coauthors defined an ecosystem as an area which is "uniform regarding the biological turnover, and contains all the fluxes above and below the ground area under consideration." They explicitly reject Gene Likens' use of entire river catchments as "too wide a demarcation" to be a single ecosystem, given the level of heterogeneity within such an area. Other authors have suggested that an ecosystem can encompass a much larger area, even the whole planet. Schulze and coauthors also rejected the idea that a single rotting log could be studied as an ecosystem because the size of the flows between the log and its surroundings are too large, relative to the proportion cycles within the log. Philosopher of science Mark Sagoff considers the failure to define "the kind of object it studies" to be an obstacle to the development of theory in ecosystem ecology.
Ecosystems can be studied through a variety of approaches—theoretical studies, studies monitoring specific ecosystems over long periods of time, those that look at differences between ecosystems to elucidate how they work and direct manipulative experimentation.Studies can be carried out at a variety of scales, ranging from whole-ecosystem studies to studying microcosms or mesocosms (simplified representations of ecosystems). American ecologist Stephen R. Carpenter has argued that microcosm experiments can be "irrelevant and diversionary" if they are not carried out in conjunction with field studies done at the ecosystem scale. Microcosm experiments often fail to accurately predict ecosystem-level dynamics.
The Hubbard Brook Ecosystem Study started in 1963 to study the White Mountains in New Hampshire. It was the first successful attempt to study an entire watershed as an ecosystem. The study used stream chemistry as a means of monitoring ecosystem properties, and developed a detailed biogeochemical model of the ecosystem.Long-term research at the site led to the discovery of acid rain in North America in 1972. Researchers documented the depletion of soil cations (especially calcium) over the next several decades.
Human activities are important in almost all ecosystems. Although humans exist and operate within ecosystems, their cumulative effects are large enough to influence external factors like climate.
Ecosystems provide a variety of goods and services upon which people depend.Ecosystem goods include the "tangible, material products" of ecosystem processes such as food, construction material, medicinal plants. They also include less tangible items like tourism and recreation, and genes from wild plants and animals that can be used to improve domestic species.
Ecosystem services, on the other hand, are generally "improvements in the condition or location of things of value".These include things like the maintenance of hydrological cycles, cleaning air and water, the maintenance of oxygen in the atmosphere, crop pollination and even things like beauty, inspiration and opportunities for research. While material from the ecosystem had traditionally been recognized as being the basis for things of economic value, ecosystem services tend to be taken for granted.
When natural resource management is applied to whole ecosystems, rather than single species, it is termed ecosystem management.Although definitions of ecosystem management abound, there is a common set of principles which underlie these definitions. A fundamental principle is the long-term sustainability of the production of goods and services by the ecosystem; "intergenerational sustainability [is] a precondition for management, not an afterthought".
While ecosystem management can be used as part of a plan for wilderness conservation, it can also be used in intensively managed ecosystems(see, for example, agroecosystem and close to nature forestry).
As human population and per capita consumption grow, so do the resource demands imposed on ecosystems and the effects of the human ecological footprint. Natural resources are vulnerable and limited. The environmental impacts of anthropogenic actions are becoming more apparent. Problems for all ecosystems include: environmental pollution, climate change and biodiversity loss. For terrestrial ecosystems further threats include air pollution, soil degradation, and deforestation. For aquatic ecosystems threats include also unsustainable exploitation of marine resources (for example overfishing of certain species), marine pollution, microplastics pollution, water pollution, the warming of oceans, and building on coastal areas.
Society is increasingly becoming aware that ecosystem services are not only limited but also that they are threatened by human activities. The need to better consider long-term ecosystem health and its role in enabling human habitation and economic activity is urgent. To help inform decision-makers, many ecosystem services are being assigned economic values, often based on the cost of replacement with anthropogenic alternatives. The ongoing challenge of prescribing economic value to nature, for example through biodiversity banking, is prompting transdisciplinary shifts in how we recognize and manage the environment, social responsibility, business opportunities, and our future as a species.[ citation needed ]
Decomposition is the process by which organic substances are broken down into a more simple organic matter. The process is a part of the nutrient cycle and is essential for recycling the finite matter that occupies physical space in the biosphere. Bodies of living organisms begin to decompose shortly after death. Animals, such as worms, also help decompose the organic materials. Organisms that do this are known as decomposers. Although no two organisms decompose in the same way, they all undergo the same sequential stages of decomposition. The science which studies decomposition is generally referred to as taphonomy from the Greek word taphos, meaning tomb.
Decomposers are organisms that break down dead or decaying organisms, and in doing so, they carry out the natural process of decomposition. Like herbivores and predators, decomposers are heterotrophic, meaning that they use organic substrates to get their energy, carbon and nutrients for growth and development. While the terms decomposer and detritivore are often interchangeably used, detritivores must ingest and digest dead matter via internal processes while decomposers can directly absorb nutrients through chemical and biological processes hence breaking down matter without ingesting it. Thus, invertebrates such as earthworms, woodlice, and sea cucumbers are technically detritivores, not decomposers, since they must ingest nutrients and are unable to absorb them externally.
Organic matter, organic material, or natural organic matter (NOM) refers to the large pool of carbon-based compounds found within natural and engineered, terrestrial and aquatic environments. It is matter composed of organic compounds that have come from the remains of organisms such as plants and animals and their waste products in the environment. Organic molecules can also be made by chemical reactions that don't involve life. Basic structures are created from cellulose, tannin, cutin, and lignin, along with other various proteins, lipids, and carbohydrates. Organic matter is very important in the movement of nutrients in the environment and plays a role in water retention on the surface of the planet.
Soil fertility refers to the ability of soil to sustain agricultural plant growth, i.e. to provide plant habitat and result in sustained and consistent yields of high quality. A fertile soil has the following properties:
Primary succession is one of two types of biological and ecological succession of plant life, occurring in an environment in which new substrate devoid of vegetation and other organisms usually lacking soil, such as a lava flow or area left from retreated glacier, is deposited. In other words, it is the gradual growth of an ecosystem over a longer period of time.
Ecosystem ecology is the integrated study of living (biotic) and non-living (abiotic) components of ecosystems and their interactions within an ecosystem framework. This science examines how ecosystems work and relates this to their components such as chemicals, bedrock, soil, plants, and animals.
Fire ecology is a scientific discipline concerned with natural processes involving fire in an ecosystem and the ecological effects, the interactions between fire and the abiotic and biotic components of an ecosystem, and the role as an ecosystem process. Many ecosystems, particularly prairie, savanna, chaparral and coniferous forests, have evolved with fire as an essential contributor to habitat vitality and renewal. Many plant species in fire-affected environments require fire to germinate, establish, or to reproduce. Wildfire suppression not only eliminates these species, but also the animals that depend upon them.
In biology, detritus is dead particulate organic material. It typically includes the bodies or fragments of dead organisms as well as fecal material. Detritus is typically colonized by communities of microorganisms which act to decompose the material. In terrestrial ecosystems, it is encountered as leaf litter and other organic matter intermixed with soil, which is denominated "soil organic matter". Detritus of aquatic ecosystems is organic material suspended in water and piling up on seabed floors, which is referred to as marine snow.
Primary producers convert an abiotic source of energy into energy stored in organic compounds, which can be used by other organisms. The primary producers can convert the energy in the light or the energy in inorganic chemical compounds (chemolithotrophs) to build organic molecules, which is usually accumulated in the form of biomass and will be used as carbon and energy source by other organisms. The photoautotrophs are the main primary producers, converting the energy of the light into chemical energy through photosynthesis, ultimately building organic molecules from carbon dioxide, an inorganic carbon source. Examples of chemolithotrophs are some archaea and bacteria that produce biomass from the oxidation of inorganic chemical compounds, these organisms are called chemoautotrophs, and are frequently found in hydrothermal vents in the deep ocean. Primary producers ares at the lowest trophic level, and are the reasons why Earth is sustainable for life to this day.
Soil biology is the study of microbial and faunal activity and ecology in soil. Soil life, soil biota, soil fauna, or edaphon is a collective term that encompasses all organisms that spend a significant portion of their life cycle within a soil profile, or at the soil-litter interface. These organisms include earthworms, nematodes, protozoa, fungi, bacteria, different arthropods, as well as some reptiles, and species of burrowing mammals like gophers, moles and prairie dogs. Soil biology plays a vital role in determining many soil characteristics. The decomposition of organic matter by soil organisms has an immense influence on soil fertility, plant growth, soil structure, and carbon storage. As a relatively new science, much remains unknown about soil biology and its effect on soil ecosystems.
Soil ecology is the study of the interactions among soil biology, and between biotic and abiotic aspects of the soil environment. It is particularly concerned with the cycling of nutrients, formation and stabilization of the pore structure, the spread and vitality of pathogens, and the biodiversity of this rich biological community.
The following outline is provided as an overview of and topical guide to ecology:
The microbial loop describes a trophic pathway in the marine microbial food web where dissolved organic carbon (DOC) is returned to higher trophic levels via its incorporation into bacterial biomass, and then coupled with the classic food chain formed by phytoplankton-zooplankton-nekton. The term microbial loop was coined by Farooq Azam and Tom Fenchel et al. to include the role played by bacteria in the carbon and nutrient cycles of the marine environment.
Soil respiration refers to the production of carbon dioxide when soil organisms respire. This includes respiration of plant roots, the rhizosphere, microbes and fauna.
Soil organic matter (SOM) is the organic matter component of soil, consisting of plant and animal residues at various stages of decomposition, cells and tissues of soil organisms, and substances synthesized by soil organisms. SOM exerts numerous positive effects on soil physical and chemical properties, as well as the soil’s capacity to provide regulatory ecosystem services. Particularly, the presence of SOM is regarded as being critical for soil functions and soil quality.
Litterfall, plant litter, leaf litter, tree litter, soil litter, or duff, is dead plant material that have fallen to the ground. This detritus or dead organic material and its constituent nutrients are added to the top layer of soil, commonly known as the litter layer or O horizon. Litter has occupied the attention of ecologists at length for the reasons that it is an instrumental factor in ecosystem dynamics, it is indicative of ecological productivity, and may be useful in predicting regional nutrient cycling and soil fertility.
Invasive species of earthworms from the suborder Lumbricina have been expanding their range in North America. Their introduction can have marked effects on the nutrient cycles in temperate forests. These earthworms increase the cycling and leaching of nutrients by breaking up decaying organic matter and spreading it into the soil. Since plants native to these northern forests are evolutionarily adapted to the presence of thick layers of decaying organic matter, the introduction of worms can lead to loss of biodiversity as young plants face less nutrient-rich conditions. Some species of trees and other plants may be incapable of surviving such changes in available nutrients. This change in the plant diversity in turn affects other organisms and often leads to increased invasions of other exotic species as well as overall forest decline.
Soil microbiology is the study of microorganisms in soil, their functions, and how they affect soil properties. It is believed that between two and four billion years ago, the first ancient bacteria and microorganisms came about on Earth's oceans. These bacteria could fix nitrogen, in time multiplied, and as a result released oxygen into the atmosphere. This led to more advanced microorganisms, which are important because they affect soil structure and fertility. Soil microorganisms can be classified as bacteria, actinomycetes, fungi, algae and protozoa. Each of these groups has characteristics that define them and their functions in soil.
Soil carbon storage is an important function of terrestrial ecosystems. Soil contains more carbon than plants and the atmosphere combined. Understanding what maintains the soil carbon pool is important to understand the current distribution of carbon on Earth, and how it will respond to environmental change. While much research has been done on how plants, free-living microbial decomposers, and soil minerals affect this pool of carbon, it is recently coming to light that mycorrhizal fungi—symbiotic fungi that associate with roots of almost all living plants—may play an important role in maintaining this pool as well. Measurements of plant carbon allocation to mycorrhizal fungi have been estimated to be 5-20% of total plant carbon uptake, and in some ecosystems the biomass of mycorrhizal fungi can be comparable to the biomass of fine roots. Recent research has shown that mycorrhizal fungi hold 50 to 70 percent of the total carbon stored in leaf litter and soil on forested islands in Sweden. Turnover of mycorrhizal biomass into the soil carbon pool is thought to be rapid and has been shown in some ecosystems to be the dominant pathway by which living carbon enters the soil carbon pool.
The fungal loop hypothesis suggests that soil fungi in arid ecosystems connect the metabolic activity of plants and biological soil crusts which respond to different soil moisture levels. Compiling diverse evidence such as limited accumulation of soil organic matter, high phenol oxidative and proteolytic enzyme potentials due to microbial activity, and symbioses between plants and fungi, the fungal loop hypothesis suggests that carbon and nutrients are cycled in biotic pools rather than leached or effluxed to the atmosphere during and between pulses of precipitation.
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