Xerosere

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Xerosere is a plant succession that is limited by water availability. It includes the different stages in a xerarch succession. Xerarch succession of ecological communities originated in extremely dry situation such as sand deserts, sand dunes, salt deserts, rock deserts etc. A xerosere may include lithoseres (on rock) and psammoseres (on sand). [1]

Contents

Stages

Bare rocks

Lava enters the Pacific at the Big Island of Hawaii Three Waikupanaha and one Ki lava ocean entries w-edit2.jpg
Lava enters the Pacific at the Big Island of Hawaii

Bare rocks are produced when glaciers recede or volcanoes erupt. Erosion of these rocks is brought by rain water and wind loaded with soil particles. The rain water combines with atmospheric carbon dioxide that corrodes the surface of the rocks and produce crevices. Water enters these crevices, freezes and expands to separate boulders. These boulders move down under the influence of gravity and wear particles from the rocks. Also when the wind loaded with soil particles strikes against the rocks, it removes soil particles. All these processes lead to formation of a little soil at the surface of these bare rocks. Animals such as spiders which can hide between boulders or stones invade these rocks. These animals live by feeding on insects which have been blown in or flown in. Algal and fungal spores reach these rocks by air from the surrounding areas. These spores grow and form symbiotic association, the lichen, which act as pioneer species of bare rocks. The process of succession starts when autotrophic organisms start living in the rocks.

Foliose and fruticose lichen stage

Foliose lichens have leaf-like thalli, while the fruticose lichens are like small bushes. They are attached to the substratum at one point only, therefore, do not cover the soil completely. They can absorb and retain more water and are able to accumulate more dust particles. Their dead remains are decomposed to humus which mixes with soil particles and help building substratum and improving soil moisture contents further. The shallow depressions in the rocks and crevices become filled with soil and topsoil layer increases further. These autogenic changes favor growth and establishment of mosses.

Moss stage

The spores of xerophytic mosses, such as Polytrichum , Tortula , and Grimmia , are brought to the rock where they succeed lichens. Their rhizoids penetrate soil among the crevices, secrete acids and corrode the rocks. The bodies of mosses are rich in organic and inorganic compounds. When these die they add these compounds to the soil, increasing the fertility of the soil. As mosses develop in patches they catch soil particles from the air and help increase the amount of substratum. The changing environment leads to migration of lichens and helps invasion of herbaceous vegetation that can out-compete mosses.

Herb stage

Encroaching herbs and shrubs Rangitotolavapath.jpg
Encroaching herbs and shrubs

Herbaceous weeds, mostly annuals such as asters, evening primroses, and milk weeds, invade the rock. Their roots penetrate deep down, secrete acids and enhance the process of weathering. Leaf litter and death of herbs add humus to the soil. Shading of soil results in decrease in evaporation and there is a slight increase in temperature. As a result, the xeric conditions begin to change and biennial and perennial herbs and xeric grasses such as Aristida , Festuca , and Poa , begin to inhabit. These climatic conditions favor growth of bacterial and fungal populations, resulting in increase in decomposition activities.

Shrub stage

The herb and grass mixture is invaded by shrub species, such as Rhus and Physocarpus . Early invasion of shrub is slow, but once a few bushes have become established, birds invade the area and help disperse scrub seeds. This results in dense scrub growth shading the soil and making conditions unfavorable for the growth of herbs, which then begin to migrate. The soil formation continues and its moisture content increases.

Tree stage

Change in environment favors colonization of tree species. The tree saplings begin to grow among the scrubs and establish themselves. The kind of tree species inhabiting the area depends upon the nature of the soil. In poorly drained soils oaks establish themselves. The trees form canopy and shade the area. Shade-loving scrubs continue to grow as secondary vegetation. Leaf litter and decaying roots weather the soil further and add humus to it making the habitat more favorable for growth to trees. Mosses and ferns make their appearance and fungi population grows abundantly.

Climax stage

The succession culminates in a climax community, the forest. Many intermediate tree stages develop prior to establishment of a climax community. The forest type depends upon climatic conditions. The climax forest may be:

Oak-hickory climax forest
In dry habitat oaks and hickories are climax vegetation. There is only one tree stage and forests are characterized by presence of scrubs, herbs, ferns, and mosses.
Beech-hemlock climax forest
These climax forests develop in mesic climates. The dominant vegetation is beech and hemlock. There are many intermediate tree stages. The other vegetation types include herbs, ferns, and mosses.
Beech-maple climax forest
These climax forests develop in mesic climates in the Northeastern United States. The dominant vegetation is American beech and sugar maple.
Spruce-alpine fir climax forest
At high altitudes in Rocky Mountains the climax forest is dominated by spruces and alpine firs.

See also

Related Research Articles

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<span class="mw-page-title-main">Pioneer species</span> First species to colonize or inhabit damaged ecosystems

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<span class="mw-page-title-main">Ecological succession</span> Process of change in the species structure of an ecological community over time

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<span class="mw-page-title-main">Primary succession</span> Gradual growth and change of an ecosystem on new substrate

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<span class="mw-page-title-main">Shrubland</span> Vegetation dominated by shrubs

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<span class="mw-page-title-main">Hydrosere</span>

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<span class="mw-page-title-main">Wistman's Wood</span> English nature reserve on Dartmoor in Devon, England

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<i>Ceratodon purpureus</i> Species of moss

Ceratodon purpureus is a dioicous moss with a color ranging from yellow-green to red. The height amounts to 3 centimeters. It is found worldwide, mainly in urban areas and next to roads on dry sand soils. It can grow in a very wide variety of habitats, from polluted highway shoulders and mine tailings to areas recently denuded by wildfire to the bright slopes of Antarctica. Its common names include redshank, purple forkmoss, ceratodon moss, fire moss, and purple horn toothed moss.

<span class="mw-page-title-main">Beech–maple forest</span> Closed canopy hardwood forest

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<span class="mw-page-title-main">Eastern forest–boreal transition</span> Temperate broadleaf and mixed forest ecoregion in Canada and the United States

The eastern forest–boreal transition is a temperate broadleaf and mixed forests ecoregion of North America, mostly in eastern Canada. It is a transitional zone or region between the predominantly coniferous Boreal Forest and the mostly deciduous broadleaf forest region further south.

<span class="mw-page-title-main">Crustose lichen</span> Growth form of lichen as a continuously adherent crust

Crustose lichens are lichens that form a crust which strongly adheres to the substrate, making separation from the substrate impossible without destruction. The basic structure of crustose lichens consists of a cortex layer, an algal layer, and a medulla. The upper cortex layer is differentiated and is usually pigmented. The algal layer lies beneath the cortex. The medulla fastens the lichen to the substrate and is made up of fungal hyphae. The surface of crustose lichens is characterized by branching cracks that periodically close in response to climatic variations such as alternate wetting and drying regimes.

<i>Rhytidiadelphus loreus</i> Species of moss in the family Hylocomiaceae

Rhytidiadelphus loreus, also commonly known as lanky moss and Little Shaggy Moss, is a nonvascular "feather moss" species that is a key component of a healthy, thriving forest ecosystem. Lanky moss grows in North America, Canada and Europe. It is primarily a coastal species that grows in moist, coniferous and deciduous forests, acidic grasslands and in the UK, heathy slopes on mountains. It grows on decaying logs, the forest floor and as an epiphyte on living trees. Its key functions in the ecosystem are water retention and temperature insulation.

Located in the Scandinavian Peninsula, Sweden is a mountainous country dominated by lakes and forests. Its habitats include mountain heath, montane forests, tundra, taiga, beech forests, rivers, lakes, bogs, brackish, marine coasts, and cultivated land. The climate of Sweden is mild for a country at this latitude, largely owing to the significant maritime influence.

<i>Timmia megapolitana</i> Genus of mosses

Timmia megapolitana, also known as Indian feather moss and warrior moss is a genus of moss in the family Timmiaceae and order Timmiales.

References

  1. Cooper, W.S. (January 1, 1913). The climax forest of Isle Royale, Lake Superior, and its development. Bot. gazette.