Pied raven | |
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Pied raven type specimen, shot 24 September 1869. Zoologisk Museum, Copenhagen | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Order: | Passeriformes |
Family: | Corvidae |
Genus: | Corvus |
Species: | C. corax |
Subspecies: | C. c. varius |
Population: | †Corvus corax varius morpha leucophaeus |
The pied raven (Corvus corax varius morpha leucophaeus) is an extinct colour morph [1] of the North Atlantic subspecies of the common raven that was only found on the Faroe Islands; the last confirmed record was in 1902. [2] The pied raven had large areas of white feathering, most frequently on the head, the wings and the belly, and its beak was light brown. Apart from that, it looked like the all-black North Atlantic ravens (C. c. varius morpha typicus), [3] which remain widespread in the Faroe Islands and are also found in Iceland. [4]
The pied raven received binomial names such as Corvus leucophaeus (by Vieillot, 1817) and Corvus leucomelas (by Wagler, 1827). It is currently referred to as Corvus corax varius morpha leucophaeus.
In modern Faroese, the bird is called hvítravnur ("white raven"), older name gorpur bringu hvíti ("white-chested corbie"). Normal individuals of the subspecies varius, which is found on Iceland and the Faroe Islands, already show a tendency towards more extensive white feather bases compared with the nominate subspecies. But, only on the Faroes, a mutation in the melanin metabolism would become fixed in the population, causing some birds to have about half of their feathers entirely white. While albinotic specimens sometimes occur in bird populations, the pied raven seems not to have been based on such occasional "sports", but on a constantly or at least regularly present part of the local raven population. [5]
The first record of the pied raven seems to be in the pre-1500 kvæði Fuglakvæði eldra ("The elder ballad of birds") which mentions 40 local species, including the great auk. Later, the pied raven is mentioned in the reports of Lucas Debes (1673 [6] ) and Jens Christian Svabo (1781/82 [7] ). Carl Julian von Graba in 1828 [8] speaks of ten individuals he saw himself and states that these birds, while less numerous than the black morph, were quite common.
Díðrikur á Skarvanesi, the Faroe painter, painted the Fuglar series, a number of portrayal of birds. On his 18 fuglar ("18 birds"), the animal in the lower right corner can be identified as a pied raven. The painting is currently on display in the Listaskálin museum of Faroe art in Tórshavn. [9]
As exemplified by Skarvanesi's painting, which obviously was done from stuffed birds, the pied raven was an object of interest to collectors. During the nineteenth century, the pied birds were selectively shot because they could fetch high prices; the sýslumaður (sheriff) of Streymoy, Hans Christopher Müller once paid two Danish rigsdaler for a stuffed specimen from Nólsoy. Such sums, a healthy amount of money for the impoverished Faroe farmers, made shooting a pied raven a profitable enterprise. Additionally, ravens in general were hunted as pests. In the mid-eighteenth century, every Faroe male of hunting age was ordered by royal decree (see Naebbetold) to shoot at least one raven or two other predatory birds per year or be fined four skillings. The last confirmed pied raven was shot on November 2, 1902, on Mykines. Subsequently, there were a few reported sightings of white ravens: in the autumn of 1916 on Velbastaður and Koltur, in the winter of 1947 on Nólsoy and again sighted the next year, and in the spring of 1965 on Sandvík. [3] Because none of these sight records mentioned the unique black-and-white pattern and a small number of diluted, all-whitish ravens have been observed in recent decades in the Faroe Islands, none of these are officially recognised as pied ravens. Consequently, 1902 is generally considered as the year of extinction for the pied raven. [3]
The pied raven, being a colour variation, likely only differed in one or very few alleles (as opposed to numerous genes in a true subspecies) from the black birds. The "piebald" allele(s) was or were presumably recessive or (if more than one) only caused the novel coloration if they were all present. This is supported by the last sightings which occurred in the absence of a regular breeding population of piebald birds, and the observations of H. C. Müller. [10] Thus, it is not certain that the form is indeed extinct, if one can speak of "extinction" in any but a population genetical sense anyway. Theoretically, the allele(s) could still be present but hidden in black individuals of the subspecies and thus, a pied raven could once again be born one day. Although the raven remains fairly common in the Faroes, its population in the small archipelago is, however, only 200–300 breeding pairs. [4] [11] Because ravens with a diluted, all-whitish plumage have been sighted in the Faroes, also in recent decades, the alleles for that aberrancy still exists in the archipelago, but this is unlikely to be the same genes involved in the pied pattern. Corvids with a diluted plumage occur with some regularity in many countries. [3]
Today, 16 museum specimens of the pied raven are known: six in the Zoologisk Museum (Copenhagen, Denmark); four in the American Museum of Natural History (New York, United States); two in the Museum of Evolution (Uppsala, Sweden); one in the Naturalis Biodiversity Center (Leiden, the Netherlands); one in State Natural History Museum (Braunschweig, Germany); one in the State Museum of Zoology (Dresden, Germany); and one in the Manchester Museum (Manchester, United Kingdom). On June 12, 1995, the Postverk Føroya issued the postal stamp FR 276, which featured a pied raven. It was designed by the famous Faroese artist and scientific illustrator Astrid Andreasen.
The peppered moth is a temperate species of night-flying moth. It is mostly found in the northern hemisphere in places like Asia, Europe and North America. Peppered moth evolution is an example of population genetics and natural selection.
A raven is any of several larger-bodied passerine bird species in the genus Corvus. These species do not form a single taxonomic group within the genus. There is no consistent distinction between crows and ravens. Names are assigned to different species chiefly based on their size.
Corvidae is a cosmopolitan family of oscine passerine birds that contains the crows, ravens, rooks, magpies, jackdaws, jays, treepies, choughs, and nutcrackers. In colloquial English, they are known as the crow family or corvids. Currently, 135 species are included in this family. The genus Corvus containing 47 species makes up over a third of the entire family. Corvids (ravens) are the largest passerines.
The pied crow is a widely distributed African bird species in the crow genus of the family Corvidae.
The brown-necked raven is a larger bird than the carrion crow though not as large as the common raven. It has similar proportions to the common raven but the bill is not so large or deep and the wings tend to be a little more pointed in profile. The head and throat are a distinct brownish-black giving the bird its English name, while the rest of the plumage is black glossed with purple, blue or purplish-blue. Like the common raven, thick-billed raven and white-necked raven, it is one of the larger raven species. The feathers of this species often fade quite quickly to a brownish black and the bird can look distinctly brown by the time it moults. The feet, legs and bill are black. The dwarf raven was formerly considered a subspecies but this bird now appears to be closer to the pied crow than this species.
The grey currawong is a large passerine bird native to southern Australia, including Tasmania. One of three currawong species in the genus Strepera, it is closely related to the butcherbirds and Australian magpie of the family Artamidae. It is a large crow-like bird, around 48 cm (19 in) long on average; with yellow irises, a heavy bill, dark plumage with white undertail and wing patches. The male and female are similar in appearance. Six subspecies are recognised and are distinguished by overall plumage colour, which ranges from slate-grey for the nominate from New South Wales and eastern Victoria and subspecies plumbea from Western Australia, to sooty black for the clinking currawong of Tasmania and subspecies halmaturina from Kangaroo Island. All grey currawongs have a loud distinctive ringing or clinking call.
The black currawong, also known locally as the black jay, is a large passerine bird endemic to Tasmania and the nearby islands within the Bass Strait. One of three currawong species in the genus Strepera, it is closely related to the butcherbirds and Australian magpie within the family Artamidae. It is a large crow-like bird, around 50 cm (20 in) long on average, with yellow irises, a heavy bill, and black plumage with white wing patches. The male and female are similar in appearance. Three subspecies are recognised, one of which, Strepera fuliginosa colei of King Island, is vulnerable to extinction.
The pied currawong is a black passerine bird native to eastern Australia and Lord Howe Island. One of three currawong species in the genus Strepera, it is closely related to the butcherbirds and Australian magpie of the family Artamidae. Six subspecies are recognised. It is a robust crowlike bird averaging around 48 cm (19 in) in length, black or sooty grey-black in plumage with white undertail and wing patches, yellow irises, and a heavy bill. The male and female are similar in appearance. Known for its melodious calls, the species' name currawong is believed to be of indigenous origin.
The little pied cormorant, little shag or kawaupaka is a common species of Australasian waterbird, found around the coasts, islands, estuaries, and inland waters of Australia, New Guinea, New Zealand, Timor-Leste and Indonesia, and around the islands of the south-western Pacific and the subantarctic. It is a small short-billed cormorant usually black above and white below with a yellow bill and small crest, although a mostly black white-throated form predominates in New Zealand. Three subspecies are recognised. Until recently most authorities referred to this species as Phalacrocorax melanoleucos.
The Australian pied cormorant, also known as the pied cormorant, pied shag, or great pied cormorant, is a medium-sized member of the cormorant family. It is found around the coasts of Australasia. In New Zealand, it is usually known either as the pied shag or by its Māori name of kāruhiruhi. Older sources may refer to it as the "yellow-faced cormorant".
The common raven is a large all-black passerine bird. It is the most widely distributed of all corvids, found across the Northern Hemisphere. It is a raven known by many names at the subspecies level; there are at least eight subspecies with little variation in appearance, although recent research has demonstrated significant genetic differences among populations from various regions. It is one of the two largest corvids, alongside the thick-billed raven, and is possibly the heaviest passerine bird; at maturity, the common raven averages 63 centimetres in length and 1.47 kilograms in mass. Although their typical lifespan is considerably shorter, common ravens can live more than 23 years in the wild. Young birds may travel in flocks but later mate for life, with each mated pair defending a territory.
The white-necked crow is the largest of the four Caribbean corvids. It is endemic to the island of Hispaniola ; it was formerly also extant on Puerto Rico and Saint Croix in the United States Virgin Islands, but has been extirpated from both islands due to considerable forest clearance and hunting for meat.
The New Zealand raven was native to New Zealand, but has been extinct since the 16th century. There were three subspecies: the North Island raven, South Island raven, and Chatham raven from the Chatham Islands.
Astrid Jóhanna Andreasen is a Faroese artist, illustrator and postage stamp designer. As a scientific illustrator, she specialises in marine animals.
The fauna of the Faroe Islands is characterized by the remote location of the islands in the North Atlantic Ocean. There are few terrestrial species, but an abundance of breeding seabirds and marine animals. Some subspecies and breeds are endemic. All land mammals were introduced by humans.
The New Zealand fantail is a small insectivorous bird, the only species of fantail in New Zealand. It has four subspecies: R. f. fuliginosa in the South Island, R. f. placabilis in the North Island, R. f. penita in the Chatham Islands, and the now-extinct R. f. cervina formerly on Lord Howe Island. It is also known by its Māori names, pīwakawaka, tīwakawaka or piwaiwaka, and the Chatham Island subspecies by the Moriori name tchitake; the common pied morph is also known as pied fantail, and the uncommon dark morph is also known as black fantail. The species has been considered by many to be conspecific as the grey fantail of Australia and New Caledonia; however, due to significant differences in its calls, many authorities now treat it as a separate species.
Carl Julian (von) Graba was a German lawyer and Royal Danish judicial councillor, and was also a keen ornithologist and one of the first modern researchers to visit and study the Faroe Islands, where he described the local puffin which was subsequently named Fratercula arctica grabae after him. Graba's findings were mentioned in 1872 by Charles Darwin in his book On the Origin of Species.
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