The Devonian is a geologic period and system of the Paleozoic era during the Phanerozoic eon, spanning 60.3 million years from the end of the preceding Silurian period at 419.2 million years ago (Ma), to the beginning of the succeeding Carboniferous period at 358.9 Ma. It is named after Devon, South West England, where rocks from this period were first studied.
Paleontology, also spelled palaeontology or palæontology, is the scientific study of life that existed prior to the start of the Holocene epoch. It includes the study of fossils to classify organisms and study their interactions with each other and their environments. Paleontological observations have been documented as far back as the 5th century BC. The science became established in the 18th century as a result of Georges Cuvier's work on comparative anatomy, and developed rapidly in the 19th century. The term has been used since 1822 formed from Greek παλαιός, ὄν, and λόγος.
The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago (Mya), to the beginning of the Devonian Period, 419.2 Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out.
Paleobotany, also spelled as palaeobotany, is the branch of botany dealing with the recovery and identification of plant remains from geological contexts, and their use for the biological reconstruction of past environments (paleogeography), and the evolutionary history of plants, with a bearing upon the evolution of life in general. A synonym is paleophytology. It is a component of paleontology and paleobiology. The prefix palaeo- or paleo- means "ancient, old", and is derived from the Greek adjective παλαιός, palaios. Paleobotany includes the study of terrestrial plant fossils, as well as the study of prehistoric marine photoautotrophs, such as photosynthetic algae, seaweeds or kelp. A closely related field is palynology, which is the study of fossilized and extant spores and pollen.
Petrified wood, is the name given to a special type of fossilized wood, the fossilized remains of terrestrial vegetation. Petrifaction is the result of a tree or tree-like plants having been replaced by stone via a mineralization process that often includes permineralization and replacement. The organic materials making up cell walls have been replicated with minerals. In some instances, the original structure of the stem tissue may be partially retained. Unlike other plant fossils, which are typically impressions or compressions, petrified wood is a three-dimensional representation of the original organic material.
Lepidodendron is an extinct genus of primitive lycopodian vascular plants belonging the order Lepidodendrales. It is well preserved and common in the fossil record. Like other Lepidodendrales, species of Lepidodendron grew as large-tree-like plants in wetland coal forest environments. They sometimes reached heights of 50 metres, and the trunks were often over 1 m in diameter. They are often known as "scale trees", due to their bark having been covered in diamond shaped leaf-bases, from which leaves grew during earlier stages of growth. However, they are correctly defined as arborescent lycophytes. They thrived during the Carboniferous Period, and persisted until the end of the Permian around 252 million years ago. Sometimes erroneously called "giant club mosses", the genus was actually more closely related to modern quillworts than to modern club mosses. In the form classification system used in paleobotany, Lepidodendron is both used for the whole plant as well as specifically the stems and leaves.
Alfisols are a soil order in USDA soil taxonomy. Alfisols form in semi-arid to humid areas, typically under a hardwood forest cover. They have a clay-enriched subsoil and relatively high native fertility. "Alf" refers to aluminium (Al) and iron (Fe). Because of their productivity and abundance, Alfisols represent one of the more important soil orders for food and fiber production. They are widely used both in agriculture and forestry, and are generally easier to keep fertile than other humid-climate soils, though those in Australia and Africa are still very deficient in nitrogen and available phosphorus. Those in monsoonal tropical regions, however, have a tendency to acidify when heavily cultivated, especially when nitrogenous fertilizers are used.
Archaeopteris is an extinct genus of progymnosperm tree with fern-like leaves. A useful index fossil, this tree is found in strata dating from the Upper Devonian to Lower Carboniferous, the oldest fossils being 385 million years old, and had global distribution.
In the geological timescale, the Middle Devonian epoch occurred during the Devonian period, after the end of the Emsian age.
Wattieza was a genus of prehistoric trees that existed in the mid-Devonian that belong to the cladoxylopsids, close relatives of the modern ferns and horsetails. The 2005 discovery in Schoharie County, New York, of fossils from the Middle Devonian about 385 million years ago united the crown of Wattieza to a root and trunk known since 1870. The fossilized grove of "Gilboa stumps" discovered at Gilboa, New York, were described as Eospermatopteris, though the complete plant remained unknown. These fossils have been described as the earliest known trees, standing 8 m (26 ft) or more tall, resembling the unrelated modern tree fern.
Prototaxites is an extinct genus of terrestrial fungi dating from the Late Silurian until the Late Devonian periods. Prototaxites formed large trunk-like structures up to 1 metre (3 ft) wide, reaching 8 metres (26 ft) in length, made up of interwoven tubes around 50 micrometres (0.0020 in) in diameter, making it by far the largest land-dwelling organism of its time.
The evolution of plants has resulted in a wide range of complexity, from the earliest algal mats of unicellular archaeplastids evolved through endosymbiosis, through multicellular marine and freshwater green algae, to spore-bearing terrestrial bryophytes, lycopods and ferns, and eventually to the complex seed-bearing gymnosperms and angiosperms of today. While many of the earliest groups continue to thrive, as exemplified by red and green algae in marine environments, more recently derived groups have displaced previously ecologically dominant ones; for example, the ascendance of flowering plants over gymnosperms in terrestrial environments.
Gilboa Fossil Forest, New York, United States, is a petrified forest and one of the oldest known forests. Located near the Gilboa Dam in Schoharie County, New York, the region is home to tree trunks from the Devonian period. The fossils, some of the only survivors of their type in the world, are believed to have been from one of the first forests on Earth, and was part of the Earth's afforestation. Paleobotanists have been interested in the site since the 1920s when construction work for a water supply project found several large, vertical fossilized stumps. Some of these remain on display at the Gilboa Dam site and the New York Power Authority Blenheim-Gilboa Visitor's Center in Schoharie County and at the New York State Museum.
This article attempts to place key plant innovations in a geological context. It concerns itself only with novel adaptations and events that had a major ecological significance, not those that are of solely anthropological interest. The timeline displays a graphical representation of the adaptations; the text attempts to explain the nature and robustness of the evidence.
Fungi diverged from other life around 1.5 billion years ago, with the glomaleans branching from the "higher fungi" (dikaryans) at ~570 million years ago, according to DNA analysis. Fungi probably colonized the land during the Cambrian, over 500 million years ago,, and possibly 635 million years ago during the Ediacaran, but terrestrial fossils only become uncontroversial and common during the Devonian, 400 million years ago.
A seed plant or spermatophyte, also known as a phanerogam or a phaenogam, is any plant that produces seeds. It is a category of embryophyte that includes most of the familiar land plants, including the flowering plants and the gymnosperms, but not ferns, mosses, or algae.
Araucaria mirabilis is an extinct species of coniferous tree from Patagonia, Argentina. It belongs to the genus Araucaria.
The Silurian-Devonian Terrestrial Revolution, also known as the Devonian Plant Explosion (DePE) and the Devonian explosion, was a period of rapid colonization, diversification and radiation of land plants and fungi on dry lands that occurred 428 to 359 million years ago (Mya) during the Silurian and Devonian periods, with the most critical phase occurring during the Late Silurian and Early Devonian.
The Waterloo Farm lagerstätte is a Famennian lagerstätte in South Africa that constitutes the only known record of a near-polar Devonian coastal ecosystem.
Calamophyton is an extinct genus of tree, or "tree-sized plant", that was extant in the Middle Devonian period. As of 2024, a well-preserved fossilized forest of Calamophyton trees discovered in Somerset, England, represents the earliest-known forest.
