Pinus muricata | |
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Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Gymnospermae |
Division: | Pinophyta |
Class: | Pinopsida |
Order: | Pinales |
Family: | Pinaceae |
Genus: | Pinus |
Subgenus: | P. subg. Pinus |
Section: | P. sect. Trifoliae |
Subsection: | P. subsect. Australes |
Species: | P. muricata |
Binomial name | |
Pinus muricata | |
Natural range of Pinus muricata |
Pinus muricata, the bishop pine, is a pine with a very restricted range: mostly in California, including several offshore Channel Islands, and a few locations in Baja California, Mexico. It is always on or near the coast. [2]
In San Luis Obispo County it is found alone or in stands scattered on the coastal mountains and hills from Morro Bay to Shell Beach. A few stands of the tree are seen on the hills above the Sycamore Canyon Resort in Avila Beach. Within the City of San Luis Obispo, the Terrace Hill Open Space has several scattered specimens. Bishop pine seems to prefer already disturbed, unvegetated areas where it probably faces less competition from oaks and shrubs.
The common name "bishop pine" resulted from the tree having been first identified near the Mission of San Luis Obispo in San Luis Obispo, California. This tree has a large number of common names and other prior scientific names, due primarily to numerous variant forms. Other English names that have occasionally been used are prickle cone pine, Obispo pine, Santa Cruz pine and dwarf marine pine.
Pinus muricata is a coniferous evergreen tree growing to a height of 15 to 26 metres (49 to 85 ft), [3] rarely up to 34 metres (112 ft), with a trunk diameter of up to 1.2 metres (3.9 ft). The species is often smaller, stunted and twisted in coastal exposures. It is drought-tolerant and grows on dry, rocky soil.
The needles are in pairs, green to blue-green, and 8–16 cm (3–6.5 in) long. Cones occur in one to five clusters. [4] The cones are strongly reflexed down the branch, 5-to-10-centimetre-long (2.0 to 3.9 in) long; the scales are stiff, thin on the side of the cone facing the stem, but greatly thickened on the side facing away and with a stout 5 to 12 millimetres (0.20 to 0.47 in) spine; both features adaptive to minimise squirrel predation and fire damage to the cones. The cones remain unopened for many years until fire or strong heat causes them to open and release the seeds. [5]
There are two Pinus muricata forms:
The resin composition also differs. The dividing line between the two is very sharp, five miles (8 km) south of the boundary between Mendocino County and Sonoma County, California. Experimental attempts to hybridize the two forms have consistently failed, indicating that their taxonomic relationship may be more distant than the very small differences in appearance would suggest.
Pinus muricata is found with several oak and cypress associates within the California Coast Ranges. There are also a number of common understory flora associates including sword fern, salal and western poison oak. [6] Notable occurrences of P. muricata is in association with Mendocino cypress as a pygmy forest on coastal terraces in Mendocino County and Sonoma County, including one location within Salt Point State Park. It is classified an endangered species in Mexico.
Pinus muricata has been used in plantations with resultant growth rates higher than in the wild, but with adverse impacts to biodiversity.
This plant has ornamental value, and is cultivated in parks and gardens. It has gained the Royal Horticultural Society's Award of Garden Merit. [7]
Pseudotsuga macrocarpa, commonly called the bigcone spruce or bigcone Douglas-fir, is an evergreen conifer native to the mountains of southern California. It is notable for having the largest cones in the genus Pseudotsuga, hence the name.
The Douglas fir is an evergreen conifer species in the pine family, Pinaceae. It is native to western North America and is also known as Douglas-fir, Douglas spruce, Oregon pine, and Columbian pine. There are three varieties: coast Douglas-fir, Rocky Mountain Douglas-fir and Mexican Douglas-fir.
Pinus radiata, the Monterey pine, insignis pine or radiata pine, is a species of pine native to the Central Coast of California and Mexico. It is an evergreen conifer in the family Pinaceae.
Pinus balfouriana, the foxtail pine, is a rare high-elevation pine that is endemic to California, United States. It is closely related to the Great Basin and Rocky Mountain bristlecone pines, in the subsection Balfourianae.
Pinus sabiniana, with vernacular names including towani pine, foothill pine, gray pine, bull pine, and digger pine, is a pine endemic to California in the United States. Some sources discourage using the name "digger pine," considering it pejorative.
The Torrey pine is a rare pine species in California, United States. It is a critically endangered species growing only in coastal San Diego County, and on Santa Rosa Island, offshore from Santa Barbara in Santa Barbara County. The Torrey pine is endemic to the California coastal sage and chaparral ecoregion.
Pinus contorta, with the common names lodgepole pine and shore pine, and also known as twisted pine, and contorta pine, is a common tree in western North America. It is common near the ocean shore and in dry montane forests to the subalpine, but is rare in lowland rain forests. Like all pines, it is an evergreen conifer.
Pinus serotina, the pond pine, black bark pine, bay pine,marsh pine, or pocosin pine, is a pine tree found along the Southeastern portion of the Atlantic coastal plain of the United States, from southern New Jersey south to Florida and west to southern Alabama. Pond pine distribution may be starting to spread west towards Mississippi and Tennessee.
Pinus parviflora, also known as five-needle pine, or Japanese white pine, is a pine in the white pine group, Pinus subgenus Strobus, native to Japan.
Coulter pine, or big-cone pine, is a native of the coastal mountains of Southern California in the United States and northern Baja California in Mexico. Isolated groves are found as far north as Clearlake, California on the flanks of Mt. Konocti and Black Diamond Mines Regional Preserve. It is named after Thomas Coulter, an Irish botanist and physician. The Coulter pine produces the heaviest cone of any pine tree, up to 5 kg (11 lb) and among conifers is exceeded only by the cones of Araucaria bidwillii.
The knobcone pine, Pinus attenuata, is a tree that grows in mild climates on poor soils. It ranges from the mountains of southern Oregon to Baja California with the greatest concentration in northern California and the Oregon-California border.
Pinus ponderosa, commonly known as the ponderosa pine, bull pine, blackjack pine, western yellow-pine, or filipinus pine is a very large pine tree species of variable habitat native to mountainous regions of western North America. It is the most widely distributed pine species in North America.
Pinus aristata, the Rocky Mountain bristlecone pine, is a long-living species of bristlecone pine tree native to the United States. It appears in the Rocky Mountains in Colorado and northern New Mexico, with isolated populations in the San Francisco Peaks in Arizona and the Kaibab National Forest north of the Grand Canyon. It is usually found at very high altitudes, from 7,000–13,000 feet (2,100–4,000 m), in cold, dry subalpine climate conditions, often at the tree line, although it also forms extensive closed-canopy stands at somewhat lower elevations.
Cupressus arizonica, the Arizona cypress, is a North American species of tree in the cypress family Cupressaceae, native to the southwestern United States and Mexico. Populations may be scattered rather than in large, dense stands.
The Northern California coastal forests are a temperate coniferous forests ecoregion of coastal Northern California and southwestern Oregon.
Cupressus goveniana, now reclassified as Hesperocyparis goveniana, with the common names Californian cypress and Gowen cypress, is a species of cypress, that is endemic to California.
Cupressus pygmaea, the Mendocino cypress or pygmy cypress, is a taxon of disputed status in the genus Cupressus endemic to certain coastal terraces and coastal mountain ranges of Mendocino and Sonoma Counties in northwestern California. It is a variable tree, and closely related to Cupressus abramsiana and Cupressus goveniana, enough to sometimes be considered conspecific with them.
A Closed-cone conifer forest or woodland is a plant community occurring in coastal California and several offshore islands. The forests typically have a single-aged single-species conifer overstory with dense ladder fuels. Overstory species include coulter pine, monterey pine, bishop pine, shore pine, and several endemic cypresses, species which generally rely on fire to open their cones and release seeds. Closed-cone forests often grow in low nutrient and/or stressed soils, which can lead to slow growth.
Dwarf forest, elfin forest, or pygmy forest is an uncommon ecosystem featuring miniature trees, inhabited by small species of fauna such as rodents and lizards. They are usually located at high elevations, under conditions of sufficient air humidity but poor soil. There are two main dwarf forest ecosystem types, involving different species and environmental characteristics: coastal temperate and montane tropical regions. Temperate coastal dwarf forest is common for parts of Southern California. Montane tropical forests are found across tropical highlands of Central America, northern South America and Southeast Asia. There are also other isolated examples of dwarf forests scattered across the world, while the largest dwarf forest is found in the Philippines.
The California interior chaparral and woodlands ecoregion covers 24,900 square miles (64,000 km2) in an elliptical ring around the California Central Valley. It occurs on hills and mountains ranging from 300 feet (91 m) to 3,000 feet (910 m). It is part of the Mediterranean forests, woodlands, and scrub biome, with cool, wet winters and hot, dry summers. Temperatures within the coast can range from 53° to 65 °F and 32° to 60 °F within the mountains. Many plant and animal species in this ecoregion are adapted to periodic fire.