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In forestry and ecology, understory (American English), or understorey (Commonwealth English), also known as underbrush or undergrowth, includes plant life growing beneath the forest canopy without penetrating it to any great extent, but above the forest floor. Only a small percentage of light penetrates the canopy so understory vegetation is generally shade-tolerant. The understory typically consists of trees stunted through lack of light, other small trees with low light requirements, saplings, shrubs, vines and undergrowth. Small trees such as holly and dogwood are understory specialists.
In temperate deciduous forests, many understory plants start into growth earlier in the year than the canopy trees, to make use of the greater availability of light at that particular time of year. A gap in the canopy caused by the death of a tree stimulates the potential emergent trees into competitive growth as they grow upwards to fill the gap. These trees tend to have straight trunks and few lower branches. At the same time, the bushes, undergrowth, and plant life on the forest floor become denser. The understory experiences greater humidity than the canopy, and the shaded ground does not vary in temperature as much as open ground. This causes a proliferation of ferns, mosses, and fungi and encourages nutrient recycling, which provides favorable habitats for many animals and plants.
The understory is the underlying layer of vegetation in a forest or wooded area, especially the trees and shrubs growing between the forest canopy and the forest floor. Plants in the understory comprise an assortment of seedlings and saplings of canopy trees together with specialist understory shrubs and herbs. Young canopy trees often persist in the understory for decades as suppressed juveniles until an opening in the forest overstory permits their growth into the canopy. In contrast understory shrubs complete their life cycles in the shade of the forest canopy. Some smaller tree species, such as dogwood and holly, rarely grow tall and generally are understory trees.
The canopy of a tropical forest is typically about 10 m (33 ft) thick, and intercepts around 95% of the sunlight. [1] The understory therefore receives less intense light than plants in the canopy and such light as does penetrate is impoverished in wavelengths of light that are most effective for photosynthesis. Understory plants therefore must be shade tolerant—they must be able to photosynthesize adequately using such light as does reach their leaves. They often are able to use wavelengths that canopy plants cannot. In temperate deciduous forests towards the end of the leafless season, understory plants take advantage of the shelter of the still leafless canopy plants to "leaf out" before the canopy trees do. This is important because it provides the understory plants with a window in which to photosynthesize without the canopy shading them. This brief period (usually 1–2 weeks) is often a crucial period in which the plant can maintain a net positive carbon balance over the course of the year.
As a rule forest understories also experience higher humidity than exposed areas. The forest canopy reduces solar radiation, so the ground does not heat up or cool down as rapidly as open ground. Consequently, the understory dries out more slowly than more exposed areas do. The greater humidity encourages epiphytes such as ferns and mosses, and allows fungi and other decomposers to flourish. This drives nutrient cycling, and provides favorable microclimates for many animals and plants, such as the pygmy marmoset. [2]
In the fields of horticulture and botany, the term deciduous means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, after flowering; and to the shedding of ripe fruit. The antonym of deciduous in the botanical sense is evergreen.
Shade is the blocking of sunlight by any object, and also the shadow created by that object. It may refer to blocking of sunlight by a roof, a tree, an umbrella, a window shade or blind, wall, curtains, or other objects.
Temperate broadleaf and mixed forest is a temperate climate terrestrial habitat type defined by the World Wide Fund for Nature, with broadleaf tree ecoregions, and with conifer and broadleaf tree mixed coniferous forest ecoregions.
An old-growth forest or primary forest is a forest that has developed over a long period of time without disturbance. Due to this, old-growth forests exhibit unique ecological features. The Food and Agriculture Organization of the United Nations defines primary forests as naturally regenerated forests of native tree species where there are no clearly visible indications of human activity and the ecological processes are not significantly disturbed. One-third of the world's forests are primary forests. Old-growth features include diverse tree-related structures that provide diverse wildlife habitats that increases the biodiversity of the forested ecosystem. Virgin or first-growth forests are old-growth forests that have never been logged. The concept of diverse tree structure includes multi-layered canopies and canopy gaps, greatly varying tree heights and diameters, and diverse tree species and classes and sizes of woody debris.
Forest dynamics are the underlying physical and biological forces that shape and change a forest ecosystem. The continuous state of change in forests can be summarized with two basic elements: disturbance and succession.
Aspen parkland refers to a very large area of transitional biome between prairie and boreal forest in two sections, namely the Peace River Country of northwestern Alberta crossing the border into British Columbia, and a much larger area stretching from central Alberta, all across central Saskatchewan to south central Manitoba and continuing into small parts of the US states of Minnesota and North Dakota. Aspen parkland consists of groves of aspen, poplar and spruce, interspersed with areas of prairie grasslands, also intersected by large stream and river valleys lined with aspen-spruce forests and dense shrubbery. This is the largest boreal-grassland transition zone in the world and is a zone of constant competition and tension as prairie and woodlands struggle to overtake each other within the parkland.
Forest ecology is the scientific study of the interrelated patterns, processes, flora, fauna, funga, and ecosystems in forests. The management of forests is known as forestry, silviculture, and forest management. A forest ecosystem is a natural woodland unit consisting of all plants, animals, and micro-organisms in that area functioning together with all of the non-living physical (abiotic) factors of the environment.
In biology, the canopy is the aboveground portion of a plant cropping or crop, formed by the collection of individual plant crowns. In forest ecology, the canopy is the upper layer or habitat zone, formed by mature tree crowns and including other biological organisms. The communities that inhabit the canopy layer are thought to be involved in maintaining forest diversity, resilience, and functioning. Shade trees normally have a dense canopy that blocks light from lower growing plants.
Cornus alternifolia is a species of flowering plant in the dogwood family Cornaceae, native to eastern North America, from Newfoundland west to southern Manitoba and Minnesota, and south to northern Florida and Mississippi. It is rare in the southern United States. It is commonly known as green osier, alternate-leaved dogwood, and pagoda dogwood.
A treefall gap is a distinguishable hole in the canopy of a forest with vertical sides extending through all levels down to an average height of 2 m (6.6 ft) above ground. These holes occur as result of a fallen tree or large limb. The ecologist who developed this definition used two meters because he believed that "a regrowth height of 2 m was sufficient" for a gap to be considered closed, but not all scientists agree. For example, Runkle believed that regrowth should be 10–20 m (33–66 ft) above the ground. Alternatively, a treefall gap is "the smallest gap [that must] be readily distinguishable amid the complexity of forest structure."
In ecology, shade tolerance is a plant's ability to tolerate low light levels. The term is also used in horticulture and landscaping, although in this context its use is sometimes imprecise, especially in labeling of plants for sale in commercial nurseries.
Temperate deciduous or temperate broad-leaf forests are a variety of temperate forest 'dominated' by deciduous trees that lose their leaves each winter. They represent one of Earth's major biomes, making up 9.69% of global land area. These forests are found in areas with distinct seasonal variation that cycle through warm, moist summers, cold winters, and moderate fall and spring seasons. They are most commonly found in the Northern Hemisphere, with particularly large regions in eastern North America, East Asia, and a large portion of Europe, though smaller regions of temperate deciduous forests are also located in South America. Examples of trees typically growing in the Northern Hemisphere's deciduous forests include oak, maple, basswood, beech and elm, while in the Southern Hemisphere, trees of the genus Nothofagus dominate this type of forest. Temperate deciduous forests provide several unique ecosystem services, including habitats for diverse wildlife, and they face a set of natural and human-induced disturbances that regularly alter their structure.
The crown of a plant is the total of an individual plant's aboveground parts, including stems, leaves, and reproductive structures. A plant community canopy consists of one or more plant crowns growing in a given area.
Frangula alnus, commonly known as alder buckthorn, glossy buckthorn, or breaking buckthorn, is a tall deciduous shrub in the family Rhamnaceae. Unlike other "buckthorns", alder buckthorn does not have thorns. It is native to Europe, northernmost Africa, and western Asia, from Ireland and Great Britain north to the 68th parallel in Scandinavia, east to central Siberia and Xinjiang in western China, and south to northern Morocco, Turkey, and the Alborz in Iran and the Caucasus Mountains; in the northwest of its range, it is rare and scattered. It is also introduced and naturalised in eastern North America.
A woody plant is a plant that produces wood as its structural tissue and thus has a hard stem. In cold climates, woody plants further survive winter or dry season above ground, as opposed to herbaceous plants that die back to the ground until spring.
Monodominance is an ecological condition in which more than 60% of the tree canopy comprises a single species of tree. Monodominant forests are quite common under conditions of extra-tropical climate types. Although monodominance is studied across different regions, most research focuses on the many prominent species in tropical forests. Connel and Lowman, originally called it single-dominance. Conventional explanations of biodiversity in tropical forests in the decades prior to Connel and Lowman's work either ignored monodominance entirely or predicted that it would not exist.
Gap dynamics refers to the pattern of plant growth that occurs following the creation of a forest gap, a local area of natural disturbance that results in an opening in the canopy of a forest. Gap dynamics are a typical characteristic of both temperate and tropical forests and have a wide variety of causes and effects on forest life.
Dry sclerophyll forests occur throughout northern and eastern Tasmania. Characterised by the population of hard-leafed (sclerophyll) and often spiky, drought-adapted plants, dry sclerophyll forests are found in regions of where annual rainfall is below 1000mm.
In ecology, stratification refers to the vertical layering of a habitat; the arrangement of vegetation in layers. It classifies the layers of vegetation largely according to the different heights to which their plants grow. The individual layers are inhabited by different animal and plant communities (stratozones).
In ecology, a light gap is a break in forest canopy or similar barrier that allows young plants to grow where they would be otherwise inhibited by the lack of light reaching the seedbed. Light gaps form predominantly when a tree falls, and thus produces an opening in the forest canopy. Light gaps are important for maintaining diversity in species-rich ecosystems.