Calocedrus

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Incense-cedar
Calocedrus decurrens 7947.jpg
Calocedrus decurrens
California incense cedar
Scientific classification OOjs UI icon edit-ltr.svg
Kingdom: Plantae
Clade: Tracheophytes
Clade: Gymnospermae
Division: Pinophyta
Class: Pinopsida
Order: Cupressales
Family: Cupressaceae
Subfamily: Cupressoideae
Genus: Calocedrus
Kurz
Type species
Calocedrus macrolepis
Synonyms [1]

HeyderiaK.Koch 1873 non Link 1833

California incense cedar, in Lassen Volcanic National Park Incense Cedar in Lassen VNP.jpg
California incense cedar, in Lassen Volcanic National Park

Calocedrus, the incense cedar (alternatively spelled incense-cedar), is a genus of coniferous trees in the cypress family Cupressaceae first described as a genus in 1873. [2] [3] Three species are native to eastern Asia and one to western North America. [1]

Contents

Description

The genus is related to Thuja , and has similar overlapping scale-leaves. Calocedrus differs from Thuja in the scale leaves being in apparent whorls of four (actually opposite decussate pairs like Thuja, but not evenly spaced apart as in Thuja, instead with the successive pairs closely then distantly spaced), and in the cones having just 2–3 pairs of moderately thin, erect scales, rather than 4–6 pairs of very thin scales in Thuja.

Taxonomy

The generic name Calocedrus means "beautiful cedar".

Cladogram showing the evolutionary relationships:

Stull et al. 2021 [4] [5]
Calocedrus

C. decurrens (Torrey) Florin

C. macrolepis Kurz

C. formosana (Florin) Florin

C. rupestris Aver., Nguyên & Lôc

C. decurrens foliage and male cones Calocedrus decurrens 7976.jpg
C. decurrens foliage and male cones

Species

Extant species

ImageNameDescriptionDistribution
Calocedrus decurrens (young female cones).jpg Calocedrus decurrens , California incense cedar (syn. Libocedrus decurrens)It is a large tree, typically reaching heights of 40–60 m and a trunk diameter of up to 3 m (maxima, 69 m tall and 3.9 m diameter), and with a broad conic crown of spreading branches. The leaves are bright green on both sides of the shoots, and the cones 2–2.5 cm long. It is by far the most widely known species in the genus, and is often simply called "incense cedar" without the regional qualifier.western North America
Calocedrus macrolepis var formosana4.jpg Calocedrus formosana , Taiwan incense cedarIt is very similar to C. macrolepis, and some botanists treat it as a variety of that, C. macrolepis var. formosana. It is a medium-sized tree, growing to 25–30 m tall, and is rare in the wild, occurring only as scattered trees in mixed forests. The leaves are glaucous green on the upper side of the shoots, and conspicuously marked with bright white stomatal patches on the underside. The cones are 1.5–2 cm long, carried on a 1–1.5 cm stem. [6] Taiwan
Calocedrus macrolepis kz1.jpg Calocedrus macrolepis , Chinese incense cedarIt is also a medium-size tree to 25–30 m tall, and like C. formosana, is rare in the wild. The leaves and cones are similar to C. formosana, differing most obviously in the shorter cone stem, only 0.5 cm long. [7] southwest China (from Guangdong west to Yunnan), and also in northern Vietnam, northern Laos, extreme northern Thailand and northeastern Myanmar
Calocedrus rupestris The most recently discovered living species of Calocedrus, first described in 2004. It occurs exclusively on rocky limestone (karst) terrain, a habitat that has a very high level of endemism. The close proximity of these populations to the Chinese and Laotian borders indicates that the species may occur in those countries as well. It is an evergreen, monoecious tree up to 25 m tall with a broadly rounded crown. The epithet "rupestris" means "rock-dwelling". [8] [9] Vietnam

Extinct species

NameDescriptionDistribution
Calocedrus huashanensis Described in 2012. It is known from compression fossils found in the Oligocene age Ningming Formation of southern China. Calocedrus huashanensis is known from branches and leaves.southern China
Calocedrus suleticensis known from fossils found in the Early Oligocene of Probostov (Holy Kluk Hill) in the volcanic complex of the Ceske stredohori Mts., Bohemia. Calocedrus suleticensis is known from a cone. Czech Republic

Uses

Archery

Incense cedar was one of the favored varieties of wood used to make bows by Native Americans in California. Like juniper, and Pacific yew, the other two coveted bow woods among Pacific Natives, this wood has excellent flexibility and compression strength-weight ratio. When backed with sinew, it produces extremely flexible, fast, hard-hitting bows, which are rivaled only by horn-sinew composite bows for their ability to store and release elastic energy. The archer Saxton Pope observed that Ishi used this wood to produce short bows. [10]

Lumber

The wood of Calocedrus is soft, moderately decay-resistant, and with a strong spicy-resinous fragrance. That of C. decurrens is the primary material for wooden pencils, because it is soft and tends to sharpen easily without forming splinters. The two Asian species were (at least in the past) in very high demand for coffin manufacture in China, due to the scent of the wood and its decay resistance. It is likely that past over-exploitation is responsible for their current rarity.

Incense cedar was the preferred hearth board of the Native Peoples of Northern California for lighting fires by friction.

Cultivation

Calocedrus decurrens, the California incense cedar, is a popular ornamental tree, grown particularly in locations with cool summer climates like Britain, Washington and British Columbia. Its very narrow columnar crown in landscape settings, an unexplained consequence of the climatic conditions in these areas, is not shown by trees in their native 'wild' habitat. The California incense cedar is also valued for its drought tolerance. The Asian species are rarely cultivated. [11]

Related Research Articles

<i>Cedrus</i> Genus of plants (coniferous trees)

Cedrus, with the common English name cedar, is a genus of coniferous trees in the plant family Pinaceae. They are native to the mountains of the western Himalayas and the Mediterranean region, occurring at altitudes of 1,500–3,200 m in the Himalayas and 1,000–2,200 m in the Mediterranean.

<i>Thuja</i> Genus of conifers

Thuja is a genus of coniferous tree or shrub in the Cupressaceae. There are five species in the genus, two native to North America and three native to eastern Asia. The genus is monophyletic and sister to Thujopsis. Members are commonly known as arborvitaes, thujas or cedars.

<span class="mw-page-title-main">Cupressaceae</span> Cypress family of conifers

Cupressaceae is a conifer family, the cypress, with worldwide distribution. The family includes 27–30 genera, which include the junipers and redwoods, with about 130–140 species in total. They are monoecious, subdioecious or (rarely) dioecious trees and shrubs up to 116 m (381 ft) tall. The bark of mature trees is commonly orange- to red-brown and of stringy texture, often flaking or peeling in vertical strips, but smooth, scaly or hard and square-cracked in some species.

<i>Cupressus</i> Several genera of evergreen conifers

Cupressus is one of several genera of evergreen conifers within the family Cupressaceae that have the common name cypress; for the others, see cypress. It is considered a polyphyletic group. Based on genetic and morphological analysis, the genus Cupressus is found in the subfamily Cupressoideae. The common name "cypress" comes via the Old French cipres from the Latin cyparissus, which is the latinisation of the Greek κυπάρισσος (kypárissos).

<i>Chamaecyparis</i> Genus of conifers

Chamaecyparis, common names cypress or false cypress, is a genus of conifers in the cypress family Cupressaceae, native to eastern Asia and to the western and eastern margins of the United States. The name is derived from the Greek khamai (χαμαί), meaning "on the earth", and kuparissos (κυπάρισσος) for "cypress".

<i>Tsuga</i> Genus of conifers

Tsuga is a genus of conifers in the subfamily Abietoideae of Pinaceae, the pine family. The common name hemlock is derived from a perceived similarity in the smell of its crushed foliage to that of the unrelated plant poison hemlock. Unlike the latter, Tsuga species are not poisonous.

<i>Callitropsis nootkatensis</i> Species of conifer

Callitropsis nootkatensis, formerly known as Cupressus nootkatensis, is a species of tree in the cypress family native to the coastal regions of northwestern North America. This species goes by many common names including: Nootka cypress, yellow cypress, Alaska cypress, Nootka cedar, yellow cedar, Alaska cedar, and Alaska yellow cedar. The specific epithet nootkatensis is derived from its discovery by Europeans on the lands of a First Nation of Canada, the Nuu-chah-nulth people of Vancouver Island, British Columbia, who were formerly referred to as the Nootka.

<i>Platycladus</i> Genus of conifers

Platycladus is a monotypic genus of evergreen coniferous trees in the cypress family Cupressaceae, containing only one species, Platycladus orientalis, also known as Chinese thuja, Oriental arborvitae, Chinese arborvitae, biota or Oriental thuja. It is native to northeastern parts of East Asia and North Asia, but is also now naturalised as an introduced species in other regions of the Asian continent.

<i>Afrocarpus</i> Genus of conifers

Afrocarpus is a genus of conifers of the family Podocarpaceae. Two to six species are recognized. They are evergreen trees native to Africa. Afrocarpus was designated a genus in 1989, when several species formerly classified in Podocarpus and Nageia were reclassified.

<i>Taiwania</i> Genus of conifers

Taiwania, with the single living species Taiwania cryptomerioides, is a large coniferous tree in the cypress family Cupressaceae.

<i>Actinostrobus</i> Genus of conifers

Actinostrobus is a genus of coniferous trees in the Cupressaceae. Common names include cypress, sandplain-cypress and cypress-pine, the last of these shared by the closely related genus Callitris.

<i>Libocedrus</i> Genus of conifers

Libocedrus is a genus of five species of coniferous trees in the cypress family Cupressaceae, native to New Zealand and New Caledonia. The genus is closely related to the South American genera Pilgerodendron and Austrocedrus, and the New Guinean genus Papuacedrus, both of which are included within Libocedrus by some botanists. These genera are rather similar to the Northern Hemisphere genera Calocedrus and Thuja: in earlier days, what is now Calocedrus was sometimes included in Libocedrus. They are much less closely related, as recently confirmed. The generic name means "teardrop cedar", apparently referring to drops of resin.

<i>Microbiota decussata</i> Species of plant

Microbiota is a monotypic genus of evergreen coniferous shrubs in the cypress family Cupressaceae, containing only one species, Microbiota decussata. The plant is native and endemic to a limited area of the Sikhote-Alin mountains in Primorskiy Krai in the Russian Far East. Microbiota is not to be confused with the range of microorganisms of the same name. The genus name was derived from micro-, meaning "small", + Biota, the genus name for a closely related conifer, a species formerly called Biota orientalis, now renamed Platycladus orientalis.

<i>Glyptostrobus pensilis</i> Species of conifer

Glyptostrobus pensilis, known in Chinese as 水松, and also Chinese swamp cypress, is an endangered conifer, and the sole living species in the genus Glyptostrobus.

<i>Retrophyllum</i> Genus of conifers

Retrophyllum is a genus of conifers in the family Podocarpaceae. It contains five generally recognized extant species with a disjunct distribution in the Southern Hemisphere, found in Papuasia and also in South America. Retrophyllum are evergreen trees typically occurring in tropical rainforests and cloud forests.

Thuja sutchuenensis, the Sichuan thuja, is a species of Thuja, an evergreen coniferous tree in the cypress family Cupressaceae. It is native to China, where it is an endangered local endemic in Chengkou County, on the southern slope of the Daba Mountains.

<i>Calocedrus decurrens</i> Species of conifer

Calocedrus decurrens, with the common names incense cedar and California incense cedar, is a species of coniferous tree native to western North America. It is the most widely known species in the genus, and is often simply called incense cedar without the regional qualifier.

<i>Cupressus torulosa</i> Species of conifer

Cupressus torulosa, commonly known as the Himalayan cypress or Bhutan cypress, is a species of cypress tree native to the mountainous northern regions of the Indian subcontinent, in the western Himalayas.

<i>Calocedrus formosana</i> Species of conifer

Calocedrus formosana is a conifer endemic to Taiwan.

<i>Calocedrus macrolepis</i> Species of conifer

Calocedrus macrolepis, commonly known as Chinese incense-cedar; is a species of conifer native to southwest China, northern Vietnam, northern Laos, extreme northern Thailand and northeastern Myanmar.

References

  1. 1 2 "Calocedrus". World Checklist of Selected Plant Families (WCSP). Royal Botanic Gardens, Kew.
  2. Kurz, Wilhelm Sulpiz (1873). "On a few new plants from Yunan". Journal of Botany, British and Foreign. 11: 196. OCLC   1642195.
  3. "Calocedrus Kurz". Tropicos . Missouri Botanical Garden.
  4. Stull, Gregory W.; Qu, Xiao-Jian; Parins-Fukuchi, Caroline; Yang, Ying-Ying; Yang, Jun-Bo; Yang, Zhi-Yun; Hu, Yi; Ma, Hong; Soltis, Pamela S.; Soltis, Douglas E.; Li, De-Zhu; Smith, Stephen A.; Yi, Ting-Shuang; et al. (2021). "Gene duplications and phylogenomic conflict underlie major pulses of phenotypic evolution in gymnosperms". Nature Plants. 7 (8): 1015–1025. bioRxiv   10.1101/2021.03.13.435279 . doi:10.1038/s41477-021-00964-4. PMID   34282286. S2CID   232282918.
  5. Stull, Gregory W.; et al. (2021). "main.dated.supermatrix.tree.T9.tre". Figshare. doi:10.6084/m9.figshare.14547354.v1.{{cite journal}}: Cite journal requires |journal= (help)
  6. Fu, Liguo; Yu, Yong-fu; Adams, Robert P.; Farjon, Aljos. "Calocedrus macrolepis var. formosana". Flora of China. Vol. 4 via eFloras.org, Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
  7. Fu, Liguo; Yu, Yong-fu; Adams, Robert P.; Farjon, Aljos. "Calocedrus macrolepis". Flora of China. Vol. 4 via eFloras.org, Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
  8. Averyanov, H.T. Nguyen & L.K. Phan. Issues of Basic Research in Life Sciences with direction in upland agriculture and forestry. Proceedings, the 2004th [sic] National Conference on Life Sciences Thai Nguyen University, September 23, 2004 41–43, 1.
  9. "Calocedrus rupestris". World Checklist of Selected Plant Families (WCSP). Royal Botanic Gardens, Kew.
  10. "How Ishi made his bow and his method of shooting, from: Hunting with the Bow and Arrow by Saxton Pope, 1923". 11 February 2019.
  11. Munz, P. A. 1974. Flora of Southern California 1–1086. University of California Press, Berkeley.