Pinus heldreichii | |
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Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Gymnospermae |
Division: | Pinophyta |
Class: | Pinopsida |
Order: | Pinales |
Family: | Pinaceae |
Genus: | Pinus |
Subgenus: | P. subg. Pinus |
Section: | P. sect. Pinus |
Subsection: | Pinus subsect. Pinaster |
Species: | P. heldreichii |
Binomial name | |
Pinus heldreichii | |
Natural range of Pinus heldreichii |
Pinus heldreichii (synonym P. leucodermis; family Pinaceae), the Bosnian pine or Heldreich's pine, [2] is a species of pine native to mountainous areas of the Balkans and southern Italy. [3]
It is an evergreen tree up to 25–35 metres (82–115 feet) in height, and 2 m (6+1⁄2 ft) in trunk diameter.
It is a member of the hard pine group, Pinus subgenus Pinus, with leaves ('needles') in fascicles (bundles) of two, with a persistent sheath. They are 4.5–10 centimetres (1+3⁄4–4 inches) long and 1.5–2 millimetres (1⁄16–3⁄32 in) thick. Cones are 5–9 cm (2–3+1⁄2 in) long, with thin, fragile scales; they are dark blue-purple before maturation, turning brown when ripe about 16–18 months after pollination. The 6–7 mm (1⁄4–9⁄32 in) long seeds have a 2–2.5 cm (3⁄4–1 in) wing and are wind-dispersed.
A tree in Northern Greece was dated as 1,075 years old in 2016. [4]
What is believed to be the oldest known living tree in Europe has been discovered in a remote mountainous area of the Pollino National Park in southern Italy. It is a Heldreich's pine estimated at 1,230 years. Much of its core has turned to dust, but there is enough new growth to confirm it is still alive. [2]
A notable specimen in the Pirin Mountains of Bulgaria, known as Baikushev's pine, is 24 m (79 ft) tall, 2.2 m (7 ft 3 in) in diameter, and is estimated to be over 1300 years old. [5]
The species was first described as Pinus heldreichii by the Swiss botanist K. Hermann Christ in honour of Theodor von Heldreich in 1863 from specimens collected on Mount Olympus, and then described a second time as P. leucodermis in 1864; the author of the second description (the Austrian botanist F. Antoine who found it on Orjen above the Bay of Kotor in Montenegro) being unaware of the slightly earlier publication by Christ. Some minor morphological differences have been claimed between the two descriptions (leading to the maintenance of both as separate taxa by a few botanists), but this is not supported by modern studies of the species, which show that both names refer to the same taxon. The discrepancies in the descriptions are largely due to Christ's cone specimens being immature and shrunken after drying, having been collected in July, four months before maturity.
It can be found in the mountains of Bosnia and Herzegovina, Montenegro, Croatia, southwestern Bulgaria, Albania, North Macedonia, Kosovo, northern Greece (Valia Kalda, Smolikas and Vasilitsa, Mount Olympus and in other high mountains), and locally in southern Italy (it is the symbol of the Pollino National Park), growing at 1,500–2,500 m (4,900–8,200 ft) altitude. It reaches the alpine tree line in these areas.
Bosnian pine is a popular ornamental tree in parks and large gardens, giving reliable, steady, though not fast, growth on a wide range of sites, and with a very neat, conical crown. It is also noted for its very decorative purple cones. The cultivars ‘Smidtii’ [6] and ‘Compact Gem’ [7] have been given the Royal Horticultural Society's Award of Garden Merit. [8] It is hardy down to at least −45 °C (−49 °F), and tolerant of severe wind exposure. Many in cultivation are still grown under the name Pinus leucodermis or Pinus heldreichii var. leucodermis.
P. heldreichii is able to adapt to extreme environmental conditions and is also a great colonizer. It is resistant to sulphur dioxide, hydrogen fluoride, nitrogen dioxide and ozone pollution and is further able to withstand wind, ice and heavy snow. These abilities makes it suitable for reforestation of extensive dry and high-altitude areas. In the south of Italy it is planted because it is less susceptible to pests than other pine species. [9]
Pinus sylvestris, the Scots pine (UK), Scotch pine (US), Baltic pine, or European red pine is a species of tree in the pine family Pinaceae that is native to Eurasia. It can readily be identified by its combination of fairly short, blue-green leaves and orange-red bark.
Western white pine, also called silver pine and California mountain pine, is a species of pine in the family Pinaceae. It occurs in mountain ranges of northwestern North America and is the state tree of Idaho.
Pinus taeda, commonly known as loblolly pine, is one of several pines native to the Southeastern United States, from East Texas to Florida, and north to southern New Jersey. The wood industry classifies the species as a southern yellow pine. U.S. Forest Service surveys found that loblolly pine is the second-most common species of tree in the United States, after red maple. For its timber, the pine species is regarded as the most commercially important tree in the Southeastern U.S. The common name loblolly is given because the pine species is found mostly in lowlands and swampy areas.
Pinus mugo, known as dwarf mountain pine, mountain pine, scrub mountain pine, Swiss mountain pine, bog pine, creeping pine, or mugo pine, is a species of conifer, native to high elevation habitats from southwestern to Central Europe and Southeast Europe.
Pinus nigra, the Austrian pine or black pine, is a moderately variable species of pine, occurring across Southern Europe from the Iberian Peninsula to the eastern Mediterranean, on the Anatolian peninsula of Turkey, Corsica and Cyprus, as well as Crimea and in the high mountains of Northwest Africa. Black pine, located in the Banaz district of Uşak, Türkiye, is estimated to be 1000 years old and is the oldest larch in the world. It has a length of 11 meters, a diameter of 3 meters and a circumference of 9.60 meters.
Pinus lambertiana is the tallest and most massive pine tree, and has the longest cones of any conifer. The species name lambertiana was given by the Scottish botanist David Douglas, who named the tree in honour of the English botanist, Aylmer Bourke Lambert. It is native to coastal and inland mountain areas along the Pacific coast of North America, as far north as Oregon and as far south as Baja California in Mexico.
Pinus peuce is a species of pine native to the mountains of North Macedonia, Bulgaria, Albania, Montenegro, Kosovo, the extreme southwest of Serbia, and the extreme north of Greece, growing typically at (600-) 1,000-2,200 (-2,300) m altitude. It often reaches the alpine tree line in this area. The mature size is up to 35–40 m height, and 1.5 m trunk diameter. However, the height of the tree diminishes strongly near the upper tree line and may even obtain shrub sizes.
Picea abies, the Norway spruce or European spruce, is a species of spruce native to Northern, Central and Eastern Europe.
Pinus contorta, with the common names lodgepole pine and shore pine, and also known as twisted pine, and contorta pine, is a common tree in western North America. It is common near the ocean shore and in dry montane forests to the subalpine, but is rare in lowland rain forests. Like all pines, it is an evergreen conifer.
Pinus parviflora, also known as five-needle pine, or Japanese white pine, is a pine in the white pine group, Pinus subgenus Strobus, native to Japan.
Pinus wallichiana is a coniferous evergreen tree native to the Himalaya, Karakoram and Hindu Kush mountains, from eastern Afghanistan east across northern Pakistan and north west India to Yunnan in southwest China. It grows in mountain valleys at altitudes of 1800–4300 m, reaching 30–50 m (98–164 ft) in height. It favours a temperate climate with dry winters and wet summers. In Pashto, it is known as Nishtar.
Pinus ponderosa, commonly known as the ponderosa pine, bull pine, blackjack pine, western yellow-pine, or filipinus pine, is a very large pine tree species of variable habitat native to mountainous regions of western North America. It is the most widely distributed pine species in North America.
The stone pine, botanical name Pinus pinea, also known as the Italian stone pine, Mediterranean stone pine, umbrella pine and parasol pine, is a tree from the pine family (Pinaceae). The tree is native to the Mediterranean region, occurring in Southern Europe and the Levant. The species was introduced into North Africa millennia ago, and is also naturalized in the Canary Islands, South Africa and New South Wales.
Pinus brutia, commonly known as the Turkish pine and Calabrian pine, is a species of pine native to the eastern Mediterranean region. The bulk of its range is in Turkey, but certain varieties are naturalized as far east as Afghanistan. It is also known as East Mediterranean pine, Afghan pine, and Brutia pine. The name "Calabrian pine" comes from an introduced grove in the Calabria region of southern Italy; historically this region was called Bruttium, which is likely where the specific epithet "brutia" comes from. Pinus brutia bears many similarities with other, closely related species such as Pinus halepensis and Pinus canariensis. Turkish pine forms a species complex with the former.
Pinus cembra, also known as Swiss pine, Swiss stone pine, Arolla pine, Austrian stone pine, or just stone pine, is a species of pine tree in the subgenus Strobus.
Picea omorika, commonly known as the Pančić spruce or Serbian spruce, is a species of coniferous tree endemic to the Drina River valley in western Serbia, and eastern Bosnia and Herzegovina, with a total range of only about 60 ha, at 800–1,600 m (2,600–5,200 ft) altitude. It was originally discovered near the Serbian village of Zaovine, on Mount Tara, in 1875, and named by the Serbian botanist Josif Pančić; the specific epithet omorika is simply the Serbian word for the tree.
Pinus halepensis, commonly known as the Aleppo pine, also known as the Jerusalem pine, is a pine native to the Mediterranean region. It was officially named by the botanist Philip Miller in his 1768 book The Gardener's Dictionary; he probably never went to Aleppo but mentions seeing large specimens at Goodwood in the garden of the Duke of Richmond, which were transplanted in 1739.
Pinus johannis, the Johann's pine, is a pine in the pinyon pine group, native to North America. The range extends from southeast Arizona and southwest New Mexico, United States, south in Mexico along the Sierra Madre Occidental and the Sierra Madre Oriental to southern Zacatecas and San Luis Potosí. It occurs at moderate to high altitudes, from 1,600–3,000 metres (5,200–9,800 ft), in cool, dry climate conditions.
Pinus quadrifolia, the Parry pinyon, is a pine in the pinyon pine group native to southernmost California in the United States and northern Baja California in Mexico, from 33° 30' N south to 30° 30' N. The Parry pinyon has a lifespan of around 200 to 500 years. It is usually found in rocky areas that often have thin soil. It occurs at moderate altitudes from 1,300 metres (4,300 ft) to 1,800 metres (5,900 ft), rarely as low as 1,200 metres (3,900 ft) and as high as 2,500 metres (8,200 ft). It is scarce and often scattered in this region, forming open woodlands, usually mixed with junipers. Other common names include nut pine and fourleaf pinyon pine.
Cedrus atlantica, the Atlas cedar, is a species of tree in the pine family Pinaceae, native to the Rif and Atlas Mountains of Morocco, and to the Tell Atlas in Algeria. A majority of the modern sources treat it as a distinct species Cedrus atlantica, but some sources consider it a subspecies of Lebanon cedar.