In biostratigraphy, MN 4 is one of the MN zones used to characterize the fossil mammal faunas of the Neogene of Europe. It is preceded by MN 3 and followed by MN 5; together, these three zones form the Orleanian age of the middle Miocene. [1] This zone starts within magnetostratigraphic chron C5Dr, at 18 million years ago, and ends within chron C5Cr, at 17.0 million years ago, although some different correlations have been proposed. [2]
The reference locality used to correlate faunas with this zone is La Romieu (southwestern France); other localities include Artesilla and Buñol in Spain, Tägernaustrasse-Jona in Switzerland, Erkertshofen 2 and Petersbuch 2 in Germany, Belchatov C in Poland, and Aliveri in Greece. [3]
In this zone, the common muroid rodent Democricetodon immigrates into Europe from Asia, followed shortly after by three other muroids: Megacricetodon , Eumyarion , and Anomalomys . Cricetodon , which later also reaches western Europe, is only present in Greece in MN 4 in Europe. The archaic muroid Melissiodon last appears in MN 4, as does Ligerimys , a member of the extinct rodent family Eomyidae. The dormouse Glirulus , which occurs in older deposits in Turkey, first appears in Europe; this genus contains a single living species, the Japanese dormouse (Glirulus japonicus). [3]
The proboscidean genera—members of the group that includes the elephants — Archaeobelodon and Prodeinotherium first appear in Europe. The rhinoceros Diaceratherium last appears in MN 4 and five other rhinoceros genera, Plesiaceratherium , Brachypotherium , Lartetotherium , Gaindatherium , and Hispanotherium , make their first appearance. Phyllotillon , a member of the related extinct family Chalicotheriidae, migrates into Europe during MN 4. Several artiodactyls, such as the pig Bunolistriodon , the tragulid Dorcatherium , the deer-like Lagomeryx , the bovid-like Amphimoschus , and the first true bovid of Europe, Eotragus , first occur during MN 4. The last occurrence of the primitive ruminant Amphitragulus also falls in this zone. The immigrant African creodont Hyainailouros occurs in Europe during MN 4. Among carnivorans, the first occurrences of the mustelid Leptoplesictis and the saber-toothed Prosansanosmilus fall in MN 4. [3]
The Bovidae are the biological family of cloven-hoofed, ruminant mammals that includes bison, African buffalo, water buffalo, antelopes, wildebeest, impala, gazelles, sheep, goats, muskoxen, and domestic cattle. A member of this family is called a bovid. With 143 extant species and 300 known extinct species, the family Bovidae consists of eight major subfamilies apart from the disputed Peleinae and Pantholopinae. The family evolved 20 million years ago, in the early Miocene.
A dormouse is a rodent of the family Gliridae. Dormice are nocturnal animals found in Africa, Asia, and Europe, and are particularly known for their long periods of hibernation.
Zokors are Asiatic burrowing rodents resembling mole-rats. They include two genera: Myospalax and Eospalax. Zokors are native to much of China, Kazakhstan, and Siberian Russia.
The rodent family Platacanthomyidae, or Oriental dormice, includes the spiny dormice and the Chinese pygmy dormice. In spite of their appearance, these animals are not true dormice, but are part of the large and complex superfamily Muroidea. The platacanthomyids can be distinguished from the true dormice, because they have no premolars, giving them three cheek teeth, like their relatives, the Muroidea.
The Malabar spiny dormouse is a species of muroid rodent endemic to the Western Ghats of India. It is the only extant species in the genus Platacanthomys and although resembling a dormouse, it is not closely related. About the size of a brown rat, this arboreal species lives in tree holes in dense forest habitats in a small family group. They are distinguishable from other species in the area by their bushy tuft tip to the tail and the spiny fur on the back.
Eomyidae is a family of extinct rodents from North America and Eurasia related to modern day pocket gophers and kangaroo rats. They are known from the Middle Eocene to the Late Miocene in North America and from the Late Eocene to the Pleistocene in Eurasia. Eomyids were generally small, but occasionally large, and tended to be squirrel-like in form and habits. The family includes the earliest known gliding rodent, Eomys quercyi.
Prosantorhinus is an extinct genus of rhinoceros from the lower and middle Miocene. The small teleoceratine rhinocerotid was found in western Europe and Asia, and resembled a hippopotamus in form.
In paleontology, biochronology is the correlation in time of biological events using fossils. In its strict sense, it refers to the use of assemblages of fossils that are not tied to stratigraphic sections. Collections of land mammal ages have been defined for every continent except Antarctica, and most are correlated with each other indirectly through known evolutionary lineages. A combination of argon–argon dating and magnetic stratigraphy allows a direct temporal comparison of terrestrial events with climate change and mass extinctions.
The Palaeomerycidae are an extinct family of ruminants in the order Artiodactyla, probably ancestral to deer and musk deer. Palaeomerycids lived in North America, Europe, Africa, Asia, and South America from 33 to 4.9 million years ago, existing for about 28 million years.
The European Land Mammal Mega Zones are zones in rock layers that have a specific assemblage of fossils (biozones) based on occurrences of fossil assemblages of European land mammals. These biozones cover most of the Neogene and Paleogene systems. In cases when fossils of mammals are abundant, stratigraphers and paleontologists can use these biozones as a more practical regional alternative to the stages of the official ICS geologic timescale. European Land Mammal Mega Zones are often also confusingly referred to as ages, stages, or intervals.
The Astaracian age is a period of geologic time, equivalent with the Middle Miocene and used more specifically with European Land Mammal Ages. It precedes the Vallesian age and follows the Orleanian age. The Astaracian overlaps the Langhian and Serravallian ages.
The Orleanian age is a period of geologic time, within the Miocene and used more specifically with European Land Mammal Ages. It precedes the Astaracian age and follows the Agenian age.
Rodents of the superfamily Muroidea, which includes mice, rats, voles, hamsters, bamboo rats, and many other species, generally have three molars in each quadrant of the jaws. A few of the oldest species retain the fourth upper premolar, and some living species have lost the third and even the second molars. Features of the molar crown are often used in muroid taxonomy, and many different systems have been proposed to name these features.
Seorsumuscardinus is a genus of fossil dormice from the early Miocene of Europe. It is known from zone MN 4 in Oberdorf, Austria; Karydia, Greece; and Tägernaustrasse-Jona, Switzerland, and from zone MN 5 in a single site at Affalterbach, Germany. The MN 4 records are placed in the species S. alpinus and the sole MN 5 record is classified as the species S. bolligeri. The latter was placed in a separate genus, Heissigia, when it was first described in 2007, but it was reclassified as a second species of Seorsumuscardinus in 2009.
Miniopterus zapfei is a fossil bat in the genus Miniopterus from the middle Miocene of France. First described in 2002, it is known only from the site of La Grive M, where it occurs with another fossil Miniopterus species, the smaller and more common Miniopterus fossilis. M. zapfei is known from five mandibles and an isolated fourth upper premolar (P4). The fourth lower premolar is more slender than in M. fossilis and the cingulum shelf surrounding the P4 is less well-developed than in living Miniopterus. The length of the first lower molar is 1.57 to 1.60 mm.
Veratalpa lugdunensiana is a fossil mammal from the Miocene of France. Known from a single astragalus, the species was assigned to its own genus, Veratalpa, by Florentino Ameghino in 1905. He placed it in Talpidae, the family of the moles, but in 1974, John Howard Hutchison argued that the astragalus was not talpid and more likely came from a rodent. The astragalus is about 4.5 mm long, broad for a talpid, and has the head oriented farther from the axis of the foot than in talpids.
In biostratigraphy, MN 5 is one of the MN zones used to characterize the fossil mammal faunas of the Neogene of Europe. It is preceded by MN 4 and followed by MN 6 and is part of the Orleanian age of the middle Miocene. MN 5 starts within magnetostratigraphic chron C5Cr, at 17.0 million years ago, and ends at the start of chron C5Bn.1r, at 15.0 million years ago, although some different correlations have been proposed. The reference locality used to correlate faunas with this zone is Pontlevoy-Thenay in France; other localities include La Retama in Spain, Castelnau-d'Arbieu in France, and Sandelzhausen in Germany.
Lagrivea is a fossil genus of squirrel from the Middle Miocene of France. The single species, L. vireti, is known from three mandibles and two isolated teeth. All come from the fissure filling of La Grive L5, part of the La Grive-Saint-Alban complex in Saint-Alban-de-Roche, southeastern France. Lagrivea was a large tree squirrel with flat lower incisors and a large, triangular fourth lower premolar (p4). Each of the four cheekteeth bears a deep basin in the middle of the crown. The m3 is about rectangular in shape, but rounded at the back. Although m1 and m2 have two roots, m3 has three.
The Mae Long Formation in the Li Basin is a fossil site in Lamphun Province, Thailand. The fossils found are thought to date to the late Early Miocene, about 18 million years ago, corresponding to the European zone MN 4.
Tupaia miocenica is a fossil treeshrew from the Miocene of Thailand. Known only from a single tooth, an upper first or second molar, it is among the few known fossil treeshrews. With a length of 3.57 mm, the tooth is large for a treeshrew. At the back lingual corner, the tooth shows a small cusp, the hypocone, that is separated from the protocone in front of it by a narrow valley. The condition of the hypocone distinguishes this species from various other treeshrews. In addition, the presence of a well-developed but simple mesostyle is distinctive.