Scolecodont

Last updated
An Ordovician scolecodont from Estonia Scolecodont ramphoprion.png
An Ordovician scolecodont from Estonia

A scolecodont is the jaw of a polychaete annelid, a common type of fossil-producing segmented worm useful in invertebrate paleontology. Scolecodonts are common and diverse microfossils, which range from the Cambrian period (around half a billion years ago at the start of the Paleozoic era) to the present. They diversified profusely in the Ordovician, [1] and are most common in the Ordovician, Silurian and Devonian marine deposits of the Paleozoic era.

Contents

Relatedly, more problematic worm-like fossils have been described in even older, Neoproterozoic era deposits in the Ediacaran Hills of southern Australia and in mid-Cambrian deposits of Burgess shale in British Columbia.

Since the other classes of annelids (specifically, the earthworms and leeches) lack hard parts, only the sea-dwelling polychaetes are frequently represented in the fossil record. Polychaetes are commonly fossilized due to their chitinous teeth and their dwelling tubes made of durable calcite (a calcium carbonate), hardened mucus (a.k.a. parchment), and/or chitin-like cement.

Taxonomy

Scolecodonts belonging to the extinct families Atraktoprionidae, Hadoprionidae, Kalloprionidae, Mochtyellidae, Paulinitidae, Polychaetaspidae, Ramphoprionidae, Rhytiprionidae, Skalenoprionidae, Symmetroprionidae, Xanioprionidae, and the still-extant (living) family Oenonidae (which includes the Arabellidae) are known from Silurian rocks in Scotland. Scolecodonts representing the present-day families Onuphidae and Dorvilleidae first appeared in Mesozoic era deposits.

History

Segments of the fossil jaw of a polychaete worm were first reported from Silurian strata on the Estonian island of Saaremaa in 1854, [2] but they were misinterpreted as fish teeth. A year later, impressions of whole polychaete worms with poorly preserved jaws were described from Italian Tertiary deposits. [3] Subsequently, E. Ehlers, a specialist on recent polychaetes, recorded them from the Jurassic Solenhofen Stone of Bavaria, Germany, demonstrating their affinity and proposing the generic names Eunicites and Lumbriconereites. [4] [5] Extensive studies in the late 19th century by George J. Hinde of material from England, Wales, Canada and Sweden [6] [7] [8] [9] established a basis for the nomenclature of what he regarded as isolated components of annelid jaws; but study of them lapsed thereafter for almost 50 years.

Related Research Articles

Ordovician Second period of the Paleozoic Era 485-444 million years ago

The Ordovician is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period 485.4 million years ago (Mya) to the start of the Silurian Period 443.8 Mya.

The PaleozoicEra is the earliest of three geologic eras of the Phanerozoic Eon. It is the longest of the Phanerozoic eras, lasting from 541 to 251.902 million years ago, and is subdivided into six geologic periods : the Cambrian, Ordovician, Silurian, Devonian, Carboniferous, and Permian. The Paleozoic comes after the Neoproterozoic Era of the Proterozoic Eon and is followed by the Mesozoic Era.

The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago (Mya), to the beginning of the Devonian Period, 419.2 Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out.

Machaeridian Extinct class of annelid worms

Machaeridia is an extinct group of armoured, segmented annelid worms, known from the Early Ordovician to Carboniferous. It consists of three distinct families: the plumulitids, turrilepadids and lepidocoleids.

Acritarch

Acritarchs are organic microfossils, known from approximately 1800 million years ago to the present. Their diversity reflects major ecological events such as the appearance of predation and the Cambrian explosion.

Llano Uplift

The Llano Uplift is a geologically ancient, low geologic dome that is about 90 miles (140 km) in diameter and located mostly in Llano, Mason, San Saba, Gillespie, and Blanco counties, Texas. It consists of an island-like exposure of Precambrian igneous and metamorphic rocks surrounded by outcrops of Paleozoic and Cretaceous sedimentary strata. At their widest, the exposed Precambrian rocks extend about 65 miles (105 km) westward from the valley of the Colorado River and beneath a broad, gentle topographic basin drained by the Llano River. The subdued topographic basin is underlain by Precambrian rocks and bordered by a discontinuous rim of flat-topped hills. These hills are the dissected edge of the Edwards Plateau, which consist of overlying Cretaceous sedimentary strata. Within this basin and along its margin are down-faulted blocks and erosional remnants of Paleozoic strata which form prominent hills.

Nautiloid Extinct subclass of nautiloids

Nautiloids are a large and diverse group of marine cephalopods (Mollusca) belonging to the subclass Nautiloidea that began in the Late Cambrian and are represented today by the living Nautilus and Allonautilus. Nautiloids flourished during the early Paleozoic era, where they constituted the main predatory animals, and developed an extraordinary diversity of shell shapes and forms. Some 2,500 species of fossil nautiloids are known, but only a handful of species survive to the present day.

Chitinozoan A taxon of flask-shaped, organic walled marine microfossils produced by an as yet unknown organism

Chitinozoa are a taxon of flask-shaped, organic walled marine microfossils produced by an as yet unknown organism. Common from the Ordovician to Devonian periods, the millimetre-scale organisms are abundant in almost all types of marine sediment across the globe. This wide distribution, and their rapid pace of evolution, makes them valuable biostratigraphic markers.

<i>Wiwaxia</i> Genus of Cambrian animals

Wiwaxia is a genus of soft-bodied animals that were covered in carbonaceous scales and spines that protected it from predators. Wiwaxia fossils – mainly isolated scales, but sometimes complete, articulated fossils – are known from early Cambrian and middle Cambrian fossil deposits across the globe. The living animal would have measured up to 5 cm (2 inch) when fully grown, although a range of juvenile specimens are known, the smallest being 2 millimetres (0.079 in) long.

Microfossil

A microfossil is a fossil that is generally between 0.001 mm and 1 mm in size, the visual study of which requires the use of light or electron microscopy. A fossil which can be studied with the naked eye or low-powered magnification, such as a hand lens, is referred to as a macrofossil.

In geochronology, an epoch is a subdivision of the geologic timescale that is longer than an age but shorter than a period. The current epoch is the Holocene Epoch of the Quaternary Period. Rock layers deposited during an epoch are called a series. Series are subdivisions of the stratigraphic column that, like epochs, are subdivisions of the geologic timescale. Like other geochronological divisions, epochs are normally separated by significant changes in the rock layers to which they correspond.

Stromatoporoidea Extinct clade of sponges

Stromatoporoidea is an extinct clade of sea sponges common in the fossil record from the Ordovician through the Devonian. They were especially abundant and important reef-formers in the Silurian and most of the Devonian. The group was previously thought to be related to the corals and placed in the Phylum Cnidaria. They are now classified in the phylum Porifera, specifically the sclerosponges. There are numerous fossil forms with spherical, branching or encrusting skeletons of laminated calcite with vertical pillars between the laminae. Specimen of its oldest genus, Priscastroma, have been found within the Middle Ordovician Sediments. This same genus has been referred to as the species P. gemina Khrom., and is known to have been known to branch off into two forms, A and B. Form A gave rise to the genus Cystostroma while form B gave rise to the genus Labechia and its descendants. Paleozoic stromatoporoids died out at the Hangenberg Event at the end of the Devonian. Purported Mesozoic stromatoporoids may be unrelated, thus making "stromatoporoids" a polyphyletic group if they are included.

The geology of Illinois includes extensive deposits of marine sedimentary rocks from the Palaeozoic, as well as relatively minor contributions from the Mesozoic and Cenozoic. Ice age glaciation left a wealth of glacial topographic features throughout the state.

<i>Diplichnites</i>

Diplichnites are arthropod trackways with two parallel rows of blunt to elongate, closely spaced tracks oriented approximately perpendicularly to the mid-line of the trackway. The term is more often used for the ichnofossils of this description; however, similar trackways from recent arthropods are sometimes given this name as well.

Paleontology in Illinois

Paleontology in Illinois refers to paleontological research occurring within or conducted by people from the U.S. state of Illinois. Scientists have found that Illinois was covered by a sea during the Paleozoic Era. Over time this sea was inhabited by animals including brachiopods, clams, corals, crinoids, sea snails, sponges, and trilobites.

Paleontology in Vermont

Paleontology in Vermont refers to paleontological research occurring within or conducted by people from the U.S. state of Vermont. Fossils are generally uncommon in Vermont. Nevertheless, however, significant finds have been made in the state. Very few fossils are known in Vermont east of the Green Mountains due to the type of rock underlying that area. During the early part of the Paleozoic era, Vermont was covered by a warm, shallow sea that would end up being home to creatures like brachiopods, corals, crinoids, ostracoderms, and trilobites. There are no rocks in the state from the Carboniferous, Permian, Triassic, or Jurassic periods. The few Cretaceous rocks present contain no fossils. The Paleogene and Neogene periods are also absent from the local rock record. During the Ice Age, glaciers scoured the state. At times the state was inundated by seawater, allowing marine mammals to venture in. After the seawater drained away the state was home to mastodons. Local fossils had already attracted scientific attention by the mid-19th century when mastodon remains were found in Rutland County. In 1950 a major Paleozoic invertebrate find occurred. The Pleistocene Beluga whale Delphinapterus leucas is the Vermont state fossil.

Paleontology in Wisconsin

Paleontology in Wisconsin refers to paleontological research occurring within or conducted by people from the U.S. state of Wisconsin. The state has fossils from the Precambrian, much of the Paleozoic, and the later part of the Cenozoic. Most of the Paleozoic rocks are marine in origin. Because of the thick blanket of Pleistocene glacial sediment that covers the rock strata in most of the state, Wisconsin’s fossil record is relatively sparse. In spite of this, certain Wisconsin paleontological occurrences provide exceptional insights concerning the history and diversity of life on Earth.

Paleontology in Minnesota

Paleontology in Minnesota refers to paleontological research occurring within or conducted by people from the U.S. state of Minnesota. The geologic record of Minnesota spans from Precambrian to recent with the exceptions of major gaps including the Silurian period, the interval from the Middle to Upper Devonian to the Cretaceous, and the Cenozoic. During the Precambrian, Minnesota was covered by an ocean where local bacteria ended up forming banded iron formations and stromatolites. During the early part of the Paleozoic era southern Minnesota was covered by a shallow tropical sea that would come to be home to creatures like brachiopods, bryozoans, massive cephalopods, corals, crinoids, graptolites, and trilobites. The sea withdrew from the state during the Silurian, but returned during the Devonian. However, the rest of the Paleozoic is missing from the local rock record. The Triassic is also missing from the local rock record and Jurassic deposits, while present, lack fossils. Another sea entered the state during the Cretaceous period, this one inhabited by creatures like ammonites and sawfish. Duckbilled dinosaurs roamed the land. The Paleogene and Neogene periods of the ensuing Cenozoic era are also missing from the local rock record, but during the Ice Age evidence points to glacial activity in the state. Woolly mammoths, mastodons, and musk oxen inhabited Minnesota at the time. Local Native Americans interpreted such remains as the bones of the water monster Unktehi. They also told myths about thunder birds that may have been based on Ice Age bird fossils. By the early 19th century, the state's fossil had already attracted the attention of formally trained scientists. Early research included the Cretaceous plant discoveries made by Leo Lesquereux.

George Jennings Hinde was a British geologist and paleontologist.

The geology of Morocco formed beginning up to two billion years ago, in the Paleoproterozoic and potentially even earlier. It was affected by the Pan-African orogeny, although the later Hercynian orogeny produced fewer changes and left the Maseta Domain, a large area of remnant Paleozoic massifs. During the Paleozoic, extensive sedimentary deposits preserved marine fossils. Throughout the Mesozoic, the rifting apart of Pangaea to form the Atlantic Ocean created basins and fault blocks, which were blanketed in terrestrial and marine sediments—particularly as a major marine transgression flooded much of the region. In the Cenozoic, a microcontinent covered in sedimentary rocks from the Triassic and Cretaceous collided with northern Morocco, forming the Rif region. Morocco has extensive phosphate and salt reserves, as well as resources such as lead, zinc, copper and silver.

References

  1. Hints, O.; Eriksson, M. (2007). "Diversification and biogeography of scolecodont-bearing polychaetes in the Ordovician". Palaeogeography, Palaeoclimatology, Palaeoecology. 245: 95. doi:10.1016/j.palaeo.2006.02.029.
  2. EICHWALD, E. 1854. Die Grauwackenschichten von Lieu-und Esthland. Bulletin de la Société Impériale des Naturalistes de Moscou, 27, 1–111.
  3. MASSALONGO, A. 1855. Monografica delle Nereidi fossili del Monte Bolca. Antonelli, Verona, 55 pp.
  4. EHLERS, E. 1868a. 12Iber eine fossile Funicee aus Solenhofen (Eunicites aritus), nebst Bemerkungenvtiber fossile Wtirmer tiberhaupt. Zeitschrifl ftir wissenschaftliche Zoologie, 18, 421–443.
  5. EHLERS, E. 1868b. Ueber fossiler Würmer aus dem lithographischen Schiefer in Bayern. Palaeontographica, 17, 145–175.
  6. HINDE, G. J. 1879. On annelid jaws from the Cambro-Silurian, Silurian and Devonian Formations in Canada and from the Lower Carboniferous in Scotland. Quarterly Journal of the Geological Society of London, 35, 370–389.
  7. HINDE, G. J. 1880. On annelid jaws from the Wenlock and Ludlow Formations of the west of England. Quarterly Journal of the Geological Society of London, 36, 368–378.
  8. HINDE, G. J. 1882. On annelid remains from the Silurian strata of the Isle of Gotland. Geologiska Föreningen i Stockholm Förhandlingar, 6(6): 254–255.
  9. HINDE, G. J. 1896. On the jaw-apparatus of an annelid (Eunicites reidiae sp. nov.) from the Lower Carboniferous of Halkin Mountain, Flintshire. Quarterly Journal of the Geological Society of London, 52, 438–450.