Sympetally (fused petals) is a flower characteristic that historically was used to classify a grouping of plants termed Sympetalae, but this term has been abandoned in newer molecular based classifications, although the grouping has similarity to the modern term asterids.
Sympetalae Rchb. (1828), meaning “with fused petals”, is a descriptive botanical name used in the Eichler, Engler, and Wettstein systems for a group of flowering plants (angiosperms). [1] In this group the flowers have a separate calyx and corolla and in which the corolla petals are fused, [2] at least at their base, a condition known as sympetally. [3]
Prior to the phylogenic classifications of August Eichler and his successors this group corresponds to the Gamopetalae of Bentham and Hooker, gamopetally being a synonym of sympetally. This was one of the three divisions of dicotyledons in their system. In Eichler's Blüthendiagramme, the class Dicotyleae was divided into two subclasses, the Sympetalae (also classified as Metachlamydeae) and the Choripetalae. [4] Adolf Engler and Karl Prantl also listed Sympetalae as a division of the class Dicotyledoneae in their system, Die Natürlichen Pflanzenfamilien, with Sympetalae being composed of gamopetalous families having gamopetalous corollas. [5] Alfred Rendle similarly described Sympetalae as originating from dicots, and then divided them into Pentacyclicae and Tetracyclicae in accordance with the number of flower parts in each group, four and five respectively. [6]
According to Engler and Prantl, Sympetalae includes the following orders: Diapensiales, Ericales, Primulales, Plumbaginales, Ebenales, Contortae, Tubiflorae, Plantaginales, Rubiales, Cucurbitales, and Campanulatae. [7]
Sympetalous flowers are found in many angiosperms, but it was the combination of sympetally with a "stamen whorl isomerous and alternate with the corolla-lobes, or stamens fewer than the corolla lobes" that Takhtajan (1964) used to define the subclass Asteridae, and later by Cronquist (1981), and later, corresponding to the asterids in the modern Angiosperm Phylogeny Group (APG) system, based on molecular phylogenetics. Since sympetally has arisen independently many times in evolution (homoplasy), on its own it is not useful for taxonomic classification. [8] [3]
The Zingiberales are flowering plants forming one of four orders in the commelinids clade of monocots, together with its sister order, Commelinales. The order includes 68 genera and 2,600 species. Zingiberales are a unique though morphologically diverse order that has been widely recognised as such over a long period of time. They are usually large herbaceous plants with rhizomatous root systems and lacking an aerial stem except when flowering. Flowers are usually large and showy, and the stamens are often modified (staminodes) to also form colourful petal-like structures that attract pollinators.
Boraginaceae, the borage or forget-me-notfamily, includes about 2,000 species of shrubs, trees, and herbs in 146 to 156 genera with a worldwide distribution.
Ranunculaceae is a family of over 2,000 known species of flowering plants in 43 genera, distributed worldwide.
The Ebenaceae are a family of flowering plants belonging to order Ericales. The family includes ebony and persimmon among about 768 species of trees and shrubs. It is distributed across the tropical and warmer temperate regions of the world. It is most diverse in the rainforests of Malesia, India, tropical Africa and tropical America.
Geraniales is a small order of flowering plants, included within the rosid subclade of eudicots. The largest family in the order is Geraniaceae with over 800 species. In addition, the order includes the smaller Francoaceae with about 40 species. Most Geraniales are herbaceous, but there are also shrubs and small trees.
Geraniaceae is a family of flowering plants placed in the order Geraniales. The family name is derived from the genus Geranium. The family includes both the genus Geranium and the garden plants called geraniums, which modern botany classifies as genus Pelargonium, along with other related genera.
The Crossosomatales are an order, first recognized as such by APG II. They are flowering plants included within the Rosid eudicots.
The Primulaceae, commonly known as the primrose family, are a family of herbaceous and woody flowering plants including some favourite garden plants and wildflowers. Most are perennial though some species, such as scarlet pimpernel, are annuals.
Asteridae is an obsolete botanical name at the rank of subclass. Composition of the subclass has also varied; however, by definition it always includes the family Asteraceae (Compositae). In the modern APG IV system of classification, asterid and euasterid are names for clades with a composition similar to that of Asteridae.
The perianth is the non-reproductive part of the flower, and structure that forms an envelope surrounding the sexual organs, consisting of the calyx (sepals) and the corolla (petals) or tepals when called a perigone. The term perianth is derived from Greek περί and άνθος, while perigonium is derived from περί and γόνος . In the mosses and liverworts (Marchantiophyta), the perianth is the sterile tubelike tissue that surrounds the female reproductive structure.
The Polemoniaceae are a family of flowering plants consisting of about 25 genera with 270–400 species of annuals and perennials native to the Northern Hemisphere and South America, with the center of diversity in western North America.
Parnassiaceae Gray were a family of flowering plants in the eudicot order Celastrales. The family is not recognized in the APG III system of plant classification. When that system was published in 2009, Parnassiaceae were treated as subfamily Parnassioideae of an expanded family Celastraceae.
In the APG IV system (2016) for the classification of flowering plants, the name asterids denotes a clade. Asterids is the largest group of flowering plants, with more than 80,000 species, about a third of the total flowering plant species. Well-known plants in this clade include the common daisy, forget-me-nots, nightshades, the common sunflower, petunias, yacon, morning glory, sweet potato, coffee, lavender, lilac, olive, jasmine, honeysuckle, ash tree, teak, snapdragon, sesame, psyllium, garden sage, table herbs such as mint, basil, and rosemary, and rainforest trees such as Brazil nut.
Sabiaceae is a family of flowering plants that were placed in the order Proteales according to the APG IV system. It comprises three genera, Meliosma, Ophiocaryon and Sabia, with 66 known species, native to tropical to warm temperate regions of southern Asia and the Americas. The family has also been called Meliosmaceae Endl., 1841, nom. rej.
Chloranthaceae is a family of flowering plants (angiosperms), the only family in the order Chloranthales. It is not closely related to any other family of flowering plants, and is among the early-diverging lineages in the angiosperms. They are woody or weakly woody plants occurring in Southeast Asia, the Pacific, Madagascar, Central and South America, and the West Indies. The family consists of four extant genera, totalling about 77 known species according to Christenhusz and Byng in 2016. Some species are used in traditional medicine. The type genus is Chloranthus. The fossil record of the family, mostly represented by pollen such as Clavatipollenites, extends back to the dawn of the history of flowering plants in the Early Cretaceous, and has been found on all continents.
The family Stylidiaceae is a taxon of dicotyledonous flowering plants. It consists of five genera with over 240 species, most of which are endemic to Australia and New Zealand. Members of Stylidiaceae are typically grass-like herbs or small shrubs and can be perennials or annuals. Most species are free standing or self-supporting, though a few can be climbing or scrambling.
Pentaphylax is a genus of flowering plants, with one or two species which are shrubs and small trees. The species has simple evergreen leaves that are alternately arranged on the stems. The flower pistil is 5 celled and the anthers dehiscing via pores and basifixed. Flowers have five petals and five sepals that are distinct from each other and five stamens attached oppositely to the sepals. The seeds are winged.
Lilioid monocots is an informal name used for a grade of five monocot orders in which the majority of species have flowers with relatively large, coloured tepals. This characteristic is similar to that found in lilies ("lily-like"). Petaloid monocots refers to the flowers having tepals which all resemble petals (petaloid). The taxonomic terms Lilianae or Liliiflorae have also been applied to this assemblage at various times. From the early nineteenth century many of the species in this group of plants were put into a very broadly defined family, Liliaceae sensu lato or s.l.. These classification systems are still found in many books and other sources. Within the monocots the Liliaceae s.l. were distinguished from the Glumaceae.
The taxonomy of the plant family Liliaceae has had a complex history since its first description in the mid-eighteenth century. Originally, the Liliaceae were defined as having a "calix" (perianth) of six equal-coloured parts, six stamens, a single style, and a superior, three-chambered (trilocular) ovary turning into a capsule fruit at maturity. The taxonomic circumscription of the family Liliaceae progressively expanded until it became the largest plant family and also extremely diverse, being somewhat arbitrarily defined as all species of plants with six tepals and a superior ovary. It eventually came to encompass about 300 genera and 4,500 species, and was thus a "catch-all" and hence paraphyletic. Only since the more modern taxonomic systems developed by the Angiosperm Phylogeny Group (APG) and based on phylogenetic principles, has it been possible to identify the many separate taxonomic groupings within the original family and redistribute them, leaving a relatively small core as the modern family Liliaceae, with fifteen genera and 600 species.
ChoripetalaeEichler (1876), is a descriptive botanical name used in the Eichler and Wettstein systems for a group of flowering plants. It was one of two groups within the Dicotyledones, the other being the Sympetalae. The latter have fused petals (sympetaly) which distinguishes them from the free, unfused petals of the Choripetalae.