Bryozoa are a phylum of simple, aquatic invertebrate animals, nearly all living in sedentary colonies. Typically about 0.5 millimetres long, they have a special feeding structure called a lophophore, a "crown" of tentacles used for filter feeding. Most marine bryozoans live in tropical waters, but a few are found in oceanic trenches and polar waters. The bryozoans are classified as the marine bryozoans (Stenolaemata), freshwater bryozoans (Phylactolaemata), and mostly-marine bryozoans (Gymnolaemata), a few members of which prefer brackish water. 5,869 living species are known. At least two genera are solitary ; the rest are colonial.
The Ordovician is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period 485.4 million years ago (Ma) to the start of the Silurian Period 443.8 Mya.
Stenolaemata are a class of exclusively marine bryozoans. Stenolaemates originated and diversified in the Ordovician, and more than 600 species are still alive today. All extant (living) species are in the order Cyclostomatida, the third-largest order of living bryozoans.
Cyclostomatida, or cyclostomata, are an ancient order of stenolaemate bryozoans which first appeared in the Lower Ordovician. It consists of 7+ suborders, 59+ families, 373+ genera, and 666+ species. The cyclostome bryozoans were dominant in the Mesozoic; since that era, they have decreased. Currently, cyclostomes seldom constitute more than 20% of the species recorded in regional bryozoan faunas.
Octocorallia is a class of Anthozoa comprising around 3,000 species of water-based organisms formed of colonial polyps with 8-fold symmetry. It includes the blue coral, soft corals, sea pens, and gorgonians within three orders: Alcyonacea, Helioporacea, and Pennatulacea. These organisms have an internal skeleton secreted by mesoglea and polyps with eight tentacles and eight mesentaries. As with all Cnidarians these organisms have a complex life cycle including a motile phase when they are considered plankton and later characteristic sessile phase.
Carbonate hardgrounds are surfaces of synsedimentarily cemented carbonate layers that have been exposed on the seafloor. A hardground is essentially, then, a lithified seafloor. Ancient hardgrounds are found in limestone sequences and distinguished from later-lithified sediments by evidence of exposure to normal marine waters. This evidence can consist of encrusting marine organisms, borings of organisms produced through bioerosion, early marine calcite cements, or extensive surfaces mineralized by iron oxides or calcium phosphates. Modern hardgrounds are usually detected by sounding in shallow water or through remote sensing techniques like side-scan sonar.
Trepostomatida is an extinct order of bryozoans in the class Stenolaemata. Trepostome bryozoans possessed mineralized calcitic skeletons and are frequently fossilized; some of the largest known fossilized bryozoan colonies are branching trepostomes and massive dome-shaped trepostomes. Trepostomes did not have many specialized zooecia beyond ordinary feeding autozooecia. The two main known heteromorphs are exilazooecia and mesozooecia, which had the purpose of maintaining regular spacing between autozooecia.
Stigmatella is an extinct genus of bryozoans in the family Heterotrypidae. It is known from the Ordovician period and found in the U.S. states of Ohio, Kentucky and Indiana. Its colonies can form branches, lobes, or masses made from new layers encrusting upon older ones. In Kentucky, fossil remains of species Stigmatella personata that lived in underwater caves have been found, growing 'upside-down' on the cave ceiling.
Fenestrata is an extinct order of bryozoan, dating from the Upper Arenig. Most fenestrate bryozoans formed net-like colonies, often in funnel- or fan-shaped forms, with a single layer of zooids facing one direction. The colony shape served as a filter-feeding apparatus that water currents flowed through, with autozooecial apertures only on the side of the colony facing into the current. This colony structure was vulnerable to predators, so some fenestrate bryozoans produced skeletal superstructures, likely to strengthen or protect the colony, and others had protective spines surrounding their autozooecial apertures.
Crepipora is an extinct genus of marine bryozoans belonging to the Ceramoporidae family. There are currently 18 collections from Belarus, Sweden, Canada, France and the United States. It was first assigned to Cystoporata by Sepkoski in 2002. The fossil range is from the Middle Ordovician to the Upper Ordovician.
Ceramopora is an extinct genus of bryozoan of the family Ceramoporidae. It is one of the earliest genera of bryozoans. Its colonies were thin and discoid, with large autozooecia, abundant communication pores, lunaria, and monticules with depressions in their centers. It had no acanthostyles or diaphragms, distinguishing it from Acanthoceramoporella.
The Lexington Limestone is a prominent geologic formation that constitutes a large part of the late Ordovician bedrock of the inner Bluegrass region in Kentucky. Named after the city of Lexington, the geologic formation has heavily influenced both the surface topography and economy of the region.
The Maquoketa Formation is a geologic formation in Illinois, Indiana. Iowa, Kansas, Minnesota, Missouri, and Wisconsin. It preserves mollusk, coral, brachiopod and graptolite fossils dating back to the Darriwilian to Hirnantian stages of the Ordovician period.
The Montoya Group is a group of geologic formations in westernmost Texas and southern New Mexico. It preserves fossils dating back to the late Ordovician period.
Monticulipora is an extinct genus of Ordovician bryozoans belonging to the family Monticuliporidae. It was first named in 1849, and its description was published the following year by French paleontologist Alcide M. d'Orbigny, making it one of the earliest bryozoans to be recognized in science. It is still one of the most widespread fossil bryozoan genera. Though colonies that grow in masses made of multiple layers are characteristic of the genus, its colonies have varying shapes, able to be encrusting, branching, massive, or frond-like, and are covered in monticules (bumps). Most Monticulipora species have distinctively granular walls, and Monticulipora and can be distinguished from Homotrypa by the presence of axial diaphragms.
Dekayia is an extinct genus of Ordovician bryozoans of the family Heterotrypidae. Its colonies can be branching, encrusting, or massive. All species have acanthopores in varying sizes and numbers. The autozooecia appear angular or sub-angular viewed through a cross-section of the colony, and their walls are distinctively undulating or crenulated. Maculae generally protrude from the colony surface very little or at all, and can contain unusually large autozooecia and a cluster of mesozooecia in their centers.
Prasopora is an extinct genus of bryozoan belonging to the family Monticuliporidae, known from the Middle Ordovician. Its colonies were disc-shaped or hemispherical, flat on bottom and convex on top, and had very abundant mesopores; in the case of the species P. insularis its zooecia were isolated from each other by the numerous mesopores surrounding them. It is very similar to the genus Monticulipora, and some bryozoan species have been assigned to both genera at different points in their study, but it is mostly distinguished by having more mesozooecia, rounder autozooecial apertures, relatively few acanthostyles and diaphragms and cystiphragms equally distributed in the autozooecia.
Diploclema is an extinct genus of bryozoan belonging to the monotypic family Dipoclemidae, found from the Middle Ordovician to the Middle Silurian. It has pear-shaped autozooecia which grow in a biradial pattern, and its colonies have a dichotomously branching shape. Its laminated exterior wall possesses a prismatic structure, which is unique among the cyclostome bryozoans.
Moyerella is an extinct genus of bryozoan of the family Arthrostylidae, known from the Upper Ordovician and Lower Silurian periods. Its colonies are branching or segmented, generally articulated (jointed). Its autozooecia are short tubes that appear triangular in cross section within the endozone, bending abruptly when reaching the exozone. The autozooecial apertures are round and aligned into diagonal rows, and paired heterozooecia occur between the autozooecial apertures.
Lichenalia is an extinct genus of cystoporate bryozoan belonging to the family Rhinoporidae. It is known from the Upper Ordovician to the Middle Silurian periods, which spanned from approximately 460 to 430 million years ago. The genus had a cosmopolitan distribution, with fossil specimens found in various regions of the world, including North America, Europe, and Asia.
