Enantiornis

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Enantiornis
Temporal range: Maastrichtian
~70  Ma
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Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Clade: Avialae
Clade: Enantiornithes
Order: Enantiornithiformes
Martin, 1983
Family: Enantiornithidae
Nesov, 1984
Genus: Enantiornis
Walker, 1981
Species:
E. leali
Binomial name
Enantiornis leali
Walker, 1981

Enantiornis is a genus of Enantiornithes. The type and only currently accepted species E. leali is from the Late Cretaceous Lecho Formation at El Brete, Argentina. It was described from specimen PVL-4035, a coracoid, proximal scapula and proximal humerus found close to each other and suspected to represent the left shoulder of a single individual. [1]

Contents

Description

The genus and the larger group it belongs to, get their name from the reversed scapula-coracoid connection they possess compared to modern birds and the hesperornithids that were their contemporaries: Enanti "opposite", ornis is "bird". [1]

Another left shoulder and wing, almost complete and found associated in one lump of rock, as well as a few isolated bones were also assigned to this species mainly based on size. It is among the largest enantiornithines discovered to date, [2] with a length in life of around 78.5 cm (30.9 in), hip height of 34 cm (13 in), weight of 6.75 kg (14.9 lb), [3] and wingspan comparable to herring gulls, around 1.2 m (3 ft 11 in). [2] Its ecological niche resembled that of a mid-sized vulture or eagle.

Taxonomy and phylogeny

E. leali was possibly fairly closely related to Avisaurus , another genus of probably carnivorous enantiornithines, though its exact relationship is unclear. It was placed in a family of its own, Enantiornithidae. Other species from Asia that were previously placed in this genus are now split off. The former Enantiornis martini is now placed in Incolornis , while the former Enantiornis walkeri is now tentatively assigned to Explorornis . The reason for this is that these species were described when the diversity of enantiornithines was underestimated.

As no hindlimb elements are known from Enantiornis, it might include one of the El Brete enantiornithines known only from leg bones, namely Lectavis , Soroavisaurus or Yungavolucris . However, these apparently were all smaller than Enantiornis. Hindlimb material tentatively assigned to Martinavis and Elbretornis seems somewhat and a lot too small, respectively, to represent Enantiornis. [4]

The cladogram below is from Wang et al., 2022: [5]

Enantiornithes

Cruralispennia

Protopteryx

Elsornis

Flexomornis

Longipterygidae

Iberomesornis

bh
bh

Longirostravis

Rapaxavis

Euenantiornithes
c
e

Sinornis

e

Enantiornis

Halimornis

Concornis

Neuquenornis

c
b

Eoalulavis

Liaoningornis

Gobipteryx

Hebeiornis

Eocathayornis

Qiliania

Avisauridae

Intiornis

Soroavisaurus

Avisaurus

Gettyia

n

Mirarce

e

Feitianius

fn

Longusunguis

fn
n

Dunhuangia

fn

Musivavis

Pengornithidae

Pengornis

m

Yuanchuavis

Eopengornis

Parapengornis

m

Grabauornis

Parvavis

Cratoavis

Gretcheniao

Huoshanornis

Shengjingornis

Zhouornis

Bohaiornithidae

Bohaiornis

ij

Eoenantiornis

Fortunguavis

Mystiornis

Shenqiornis

Sulcavis

l

Key to letters:

b = Boluochia
c = Cathayornis
e = Enantiophoenix
f = Houornis
h = Longipteryx
i = Parabohaiornis
j = Pterygornis
l = Vorona
m = Yuanjiawaornis
n = Yungavolucris

Related Research Articles

<span class="mw-page-title-main">Enantiornithes</span> Extinct clade of dinosaurs

The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans, the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and clawed fingers on each wing, but otherwise looked much like modern birds externally. Over eighty species of Enantiornithes have been named, but some names represent only single bones, so it is likely that not all are valid. The Enantiornithes became extinct at the Cretaceous–Paleogene boundary, along with Hesperornithes and all other non-avian dinosaurs.

<i>Avisaurus</i> Extinct genus of birds

Avisaurus is a genus of enantiornithine bird from the Late Cretaceous of North America.

<i>Alexornis</i> Extinct genus of birds

Alexornis is a genus of enantiornithine birds from the Bocana Roja Formation of Baja California, Mexico. This geological formation has been dated to the late Cretaceous period, and more specifically to the Campanian age, about 73 mya. The type and only known species is Alexornis antecedens. Its name means "Alex's ancestral bird"; Alexornis from the given name of ornithologist Alexander Wetmore + Ancient Greek ornis, "bird", and antecedens, Latin for "going before" or "ancestral".

Incolornis is an extinct genus of basal birds from the Late Cretaceous. Remains have been found in the Bissekty Formation in Uzbekistan. It is only known from fragmentary coracoids. The systematic position of the genus is unclear; it might have been an enantiornithine bird.

<i>Lectavis</i> Extinct genus of birds

Lectavis is a genus of enantiornithine birds. Their fossil bones have been recovered from the Late Cretaceous Lecho Formation at estancia El Brete, Argentina. The genus contains a single species, Lectavis bretincola.

Gurilynia is a genus of enantiornithine birds. One species is known, G. nessovi. It lived during the Maastrichtian stage of the Late Cretaceous period, between 70 and 66 mya. Gurilynia is known from fragmentary fossils found at the Gurilyn Tsav locality of the Nemegt Formation in south Gobi, Mongolia.

Neuquenornis volans is a species of enantiornithean birds which lived during the late Cretaceous period in today's Patagonia, Argentina. It is the only known species of the genus Neuquenornis. Its fossils were found in the Santonian Bajo de la Carpa Formation, dating from about 85-83 million years ago. This was a sizeable bird for its time, with a tarsometatarsus 46.8mm long. Informal estimates suggest that it measured nearly 30 cm (12 in) in length excluding the tail.

<i>Palintropus</i> Extinct genus of birds

Palintropus is a prehistoric bird genus from the Late Cretaceous. A single species has been named based on a proximal coracoid from the Lance Formation of Wyoming, dated to the latest Maastrichtian, 66 million years ago. Coracoids and a proximal scapula of two unnamed species from the upper Campanian Dinosaur Park Formation of Alberta, dating to between 76.5 and 75 million years ago, are also known.

<i>Dongbeititan</i> Extinct genus of dinosaurs

Dongbeititan is a genus of sauropod dinosaur from the Early Cretaceous-age Yixian Formation of Beipiao, Liaoning, China. It is based on holotype DNHM D2867, a partial postcranial skeleton including bones from the limbs, shoulder and pelvic girdles, and vertebrae, which was described in 2007. Its describers suggested it was as a basal titanosauriform, not as derived as Gobititan or Jiutaisaurus, but more derived than Euhelopus, Fusuisaurus, and Huanghetitan. The type species is D. dongi, and it is the first named sauropod from the Yixian Formation, which is part of the well-known Jehol Group. The genus name refers to the region Dongbei and to Greek titan, "giant". The specific name honours the Chinese paleontologist Dong Zhiming. Like other sauropods, Dongbeititan would have been a large quadrupedal herbivore.

<span class="mw-page-title-main">Avisauridae</span> Extinct family of dinosaurs

Avisauridae is a family of extinct enantiornithine dinosaurs from the Cretaceous period, distinguished by several features of their ankle bones. Depending on the definition used, Avisauridae is either a broad and widespread group of advanced enantiornithines, or a small family within that group, restricted to species from the Late Cretaceous of North and South America.

Soroavisaurus is a genus of enantiornithean birds related to Avisaurus. It lived during the Late Cretaceous of Argentina. The only known species, S. australis, is known from fossils collected from the Lecho Formation of Estancia El Brete, in the southern tip of the province of Salta, Argentina. A binominal name of this animal means "Southern sister Avisaur".

<span class="mw-page-title-main">Longipterygidae</span> Extinct family of dinosaurs

Longipterygidae is a family of early enantiornithean avialans from the Early Cretaceous epoch of China. All known specimens come from the Jiufotang Formation and Yixian Formation, dating to the early Aptian age, 125-120 million years ago.

Elbretornis is an extinct genus of enantiornithine which existed in what is now Salta Province, Argentina during the late Cretaceous period.

Martinavis is a genus of enantiornithine birds which existed in what is now southern France, North America and Salta Province, Argentina during the late Cretaceous period. It was named by Cyril A. Walker, Eric Buffetaut and Gareth J. Dyke in 2007, and the type species is Martinavis cruzyensis.

<i>Cratoavis</i> Extinct genus of dinosaurs

Cratoavis is a genus of enantiornithines. The type and only currently described species is C. cearensis, from the Early Cretaceous of Araripe Basin, Ceará, Brazil. The fossil, an articulated skeleton with feathers attached to the wings and surrounding the body, extends considerably the temporal record of the group at South America.

<i>Zhongjianosaurus</i> Extinct genus of reptiles

Zhongjianosaurus is a genus of dromaeosaurid belonging to the Microraptoria. Believed to hail from the Yixian Formation, specifically the middle of the Jehol Biota, it is the smallest known microraptorine thus far discovered and one of the smallest non-avian theropod dinosaurs.

<i>Jinguofortis</i> Extinct genus of birds

Jinguofortis is a genus of primitive avialan (bird) belonging to the clade Pygostylia that lived during the Valanginian stage of the Early Cretaceous. It was found in the Dabeigou Formation in northeastern China, and isotope dating from the samples overlying the bird-bearing horizon is 127 million years ago.

<i>Mirarce</i> Extinct genus of birds

Mirarce is a genus of enantornithe bird from the Late Cretaceous of Utah. It contains a single species, M. eatoni. It was similar in size to modern turkeys.

<i>Fukuipteryx</i> Extinct genus of dinosaurs

Fukuipteryx is an extinct genus of basal avialan dinosaurs found in Early Cretaceous deposits from Japan's Kitadani Formation. It contains one species, Fukuipteryx prima.

References

  1. 1 2 Walker, C. A. (1981). "New subclass of birds from the Cretaceous of South America". Nature. 292 (5818): 51–53. Bibcode:1981Natur.292...51W. doi:10.1038/292051a0. S2CID   4340858.
  2. 1 2 Colin Tudge (October 20, 2009). The Bird: A Natural History of Who Birds Are, Where They Came From, and How They Live. Crown. p. 64. ISBN   9780307459763 . Retrieved 29 August 2022.
  3. Rubén Molina-Pérez, Asier Larramendi, David Connolly, Gonzalo Ángel Ramírez Cruz, Andrey Atuchin (June 25, 2019). Dinosaur Facts and Figures: The Theropods and Other. Princeton University Press. p. 281. ISBN   9780691190594 . Retrieved 29 August 2022.{{cite book}}: CS1 maint: multiple names: authors list (link)
  4. Cyril A. Walker & Gareth J. Dyke (2009). "Euenantiornithine birds from the Late Cretaceous of El Brete (Argentina)" (PDF). Irish Journal of Earth Sciences. 27: 15–62. doi:10.3318/IJES.2010.27.15. S2CID   129573066. Archived from the original (PDF) on 2012-03-20.
  5. Wang, Xuri; Cau, Andrea; Luo, Xiaoling; Kundrát, Martin; Wu, Wensheng; Ju, Shubin; Guo, Zhen; Liu, Yichuan; Ji, Qiang (2022-02-11). "A new bohaiornithid-like bird from the Lower Cretaceous of China fills a gap in enantiornithine disparity". Journal of Paleontology. 96 (4): 961–976. doi: 10.1017/jpa.2022.12 . ISSN   0022-3360. S2CID   247432530.