In botany, the trunk (or bole) is the stem and main wooden axis of a tree, [1] which is an important feature in tree identification, and which often differs markedly from the bottom of the trunk to the top, depending on the species.
The trunk is the most important part of the tree for timber production.
Trunks occur both in "true" woody plants and non-woody plants such as palms and other monocots, though the internal physiology is different in each case. In all plants, trunks thicken over time due to the formation of secondary growth, or, in monocots, pseudo-secondary growth. Trunks can be vulnerable to damage, including sunburn.
Trunks which are cut down for making lumber are generally called logs; if they are cut to a specific length, called bolts. The term "log" is informally used in English to describe any felled trunk not rooted in the ground, whose roots are detached. A stump is the part of a trunk remaining in the ground after the tree has been felled, or the earth-end of an uprooted tree which retains its un-earthed roots.
The trunk consists of five main parts: The outer bark, inner bark (phloem), cambium, sapwood (live xylem), and heartwood (dead xylem). [2] From the outside of the tree working in:
Wood is a structural tissue found in the stems and roots of trees and other woody plants. It is an organic material – a natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin that resists compression. Wood is sometimes defined as only the secondary xylem in the stems of trees, or more broadly to include the same type of tissue elsewhere, such as in the roots of trees or shrubs. In a living tree it performs a support function, enabling woody plants to grow large or to stand up by themselves. It also conveys water and nutrients between the leaves, other growing tissues, and the roots. Wood may also refer to other plant materials with comparable properties, and to material engineered from wood, woodchips, or fiber.
Phloem is the living tissue in vascular plants that transports the soluble organic compounds made during photosynthesis and known as photosynthates, in particular the sugar sucrose, to the rest of the plant. This transport process is called translocation. In trees, the phloem is the innermost layer of the bark, hence the name, derived from the Ancient Greek word φλοιός (phloiós), meaning "bark". The term was introduced by Carl Nägeli in 1858. Different types of phloem can be distinguished. The early phloem formed in the growth apices is called protophloem. Protophloem eventually becomes obliterated once it connects to the durable phloem in mature organs, the metaphloem. Further, secondary phloem is formed during the thickening of stem structures.
In biology, tissue is a historically derived biological organizational level between cells and a complete organ. A tissue is therefore often thought of as an assembly of similar cells and their extracellular matrix from the same embryonic origin that together carry out a specific function. Organs are then formed by the functional grouping together of multiple tissues.
Bark is the outermost layer of stems and roots of woody plants. Plants with bark include trees, woody vines, and shrubs. Bark refers to all the tissues outside the vascular cambium and is a nontechnical term. It overlays the wood and consists of the inner bark and the outer bark. The inner bark, which in older stems is living tissue, includes the innermost layer of the periderm. The outer bark on older stems includes the dead tissue on the surface of the stems, along with parts of the outermost periderm and all the tissues on the outer side of the periderm. The outer bark on trees which lies external to the living periderm is also called the rhytidome.
The vascular cambium is the main growth tissue in the stems and roots of many plants, specifically in dicots such as buttercups and oak trees, gymnosperms such as pine trees, as well as in certain other vascular plants. It produces secondary xylem inwards, towards the pith, and secondary phloem outwards, towards the bark.
Cork cambium is a tissue found in many vascular plants as a part of the epidermis. It is one of the many layers of bark, between the cork and primary phloem. The cork cambium is a lateral meristem and is responsible for secondary growth that replaces the epidermis in roots and stems. It is found in woody and many herbaceous dicots, gymnosperms and some monocots. It is one of the plant's meristems – the series of tissues consisting of embryonic disk cells from which the plant grows. The function of cork cambium is to produce the cork, a tough protective material.
Lepidodendron is an extinct genus of primitive lycopodian vascular plants belonging the order Lepidodendrales. It is well preserved and common in the fossil record. Like other Lepidodendrales, species of Lepidodendron grew as large-tree-like plants in wetland coal forest environments. They sometimes reached heights of 50 metres, and the trunks were often over 1 m (3.3 ft) in diameter. They are often known as "scale trees", due to their bark having been covered in diamond shaped leaf-bases, from which leaves grew during earlier stages of growth. However, they are correctly defined as arborescent lycophytes. They thrived during the Carboniferous Period, and persisted until the end of the Permian around 252 million years ago. Sometimes erroneously called "giant club mosses", the genus was actually more closely related to modern quillworts than to modern club mosses. In the form classification system used in paleobotany, Lepidodendron is both used for the whole plant as well as specifically the stems and leaves.
Girdling, also called ring-barking, is the circumferential removal or injury of the bark of a branch or trunk of a woody plant. Girdling prevents the tree from sending nutrients from its foliage to its roots, resulting in the death of the tree over time, and can also prevent flow of nutrients in the other direction depending on how much of the xylem is removed. A branch completely girdled will fail and when the main trunk of a tree is girdled, the entire tree will die, if it cannot regrow from above to bridge the wound. Human practices of girdling include forestry, horticulture, and vandalism. Foresters use the practice of girdling to thin forests. Extensive cankers caused by certain fungi, bacteria or viruses can girdle a trunk or limb. Animals such as rodents will girdle trees by feeding on outer bark, often during winter under snow. Girdling can also be caused by herbivorous mammals feeding on plant bark and by birds and insects, both of which can effectively girdle a tree by boring rows of adjacent holes.
Vascular tissue is a complex conducting tissue, formed of more than one cell type, found in vascular plants. The primary components of vascular tissue are the xylem and phloem. These two tissues transport fluid and nutrients internally. There are also two meristems associated with vascular tissue: the vascular cambium and the cork cambium. All the vascular tissues within a particular plant together constitute the vascular tissue system of that plant.
A bridge graft is a grafting technique used to re-establish the supply of nutrients to the rootstock of a woody perennial when the full thickness of the bark has been removed from part of the trunk.
In botany, secondary growth is the growth that results from cell division in the cambia or lateral meristems and that causes the stems and roots to thicken, while primary growth is growth that occurs as a result of cell division at the tips of stems and roots, causing them to elongate, and gives rise to primary tissue. Secondary growth occurs in most seed plants, but monocots usually lack secondary growth. If they do have secondary growth, it differs from the typical pattern of other seed plants.
The unifacial cambium produces cells to the interior of its cylinder. These cells differentiate into xylem tissue. Unlike the more common bifacial cambium found in later woody plants, the unifacial cambium does not produce phloem to its exterior. Also in contrast to the bifacial cambium, the unifacial cambium is unable to expand its circumference with anticlinal cell division. Cell elongation provides a limited amount of expansion.
Wood anatomy is a scientific sub-area of wood science, which examines the variations in xylem anatomical characteristics across trees, shrubs, and herbaceous species to explore inquiries related to plant function, growth, and the environment.
Lepidodendrales or arborescent lycophytes are an extinct order of primitive, vascular, heterosporous, arborescent (tree-like) plants belonging to Lycopodiopsida. Members of Lepidodendrales are the best understood of the fossil lycopsids due to the vast diversity of Lepidodendrales specimens and the diversity in which they were preserved; the extensive distribution of Lepidodendrales specimens as well as their well-preservedness lends paleobotanists exceptionally detailed knowledge of the coal-swamp giants’ reproductive biology, vegetative development, and role in their paleoecosystem. The defining characteristics of the Lepidodendrales are their secondary xylem, extensive periderm development, three-zoned cortex, rootlike appendages known as stigmarian rootlets arranged in a spiralling pattern, and megasporangium each containing a single functional megaspore that germinates inside the sporangium. Many of these different plant organs have been assigned both generic and specific names as relatively few have been found organically attached to each other. Some specimens have been discovered which indicate heights of 40 and even 50 meters and diameters of over 2 meters at the base. The massive trunks of some species branched profusely, producing large crowns of leafy twigs; though some leaves were up to 1 meter long, most were much shorter, and when leaves dropped from branches their conspicuous leaf bases remained on the surface of branches. Strobili could be found at the tips of distal branches or in an area at the top of the main trunk. The underground organs of Lepidodendrales typically consisted of dichotomizing axes bearing helically arranged, lateral appendages serving an equivalent function to roots. Sometimes called "giant club mosses", they are believed to be more closely related to extant quillworts based on xylem, although fossil specimens of extinct Selaginellales from the Late Carboniferous also had secondary xylem.
A woody plant is a plant that produces wood as its structural tissue and thus has a hard stem. In cold climates, woody plants further survive winter or dry season above ground, as opposed to herbaceous plants that die back to the ground until spring.
A stem is one of two main structural axes of a vascular plant, the other being the root. It supports leaves, flowers and fruits, transports water and dissolved substances between the roots and the shoots in the xylem and phloem, photosynthesis takes place here, stores nutrients, and produces new living tissue. The stem can also be called halm or haulm or culms.
In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are usable as lumber or plants above a specified height. In wider definitions, the taller palms, tree ferns, bananas, and bamboos are also trees.
Laminated root rot also known as yellow ring rot is caused by the fungal pathogen Phellinus weirii. Laminated root rot is one of the most damaging root disease amongst conifers in northwestern America and true firs, Douglas fir, Mountain hemlock, and Western hemlock are highly susceptible to infection with P. weirii. A few species of plants such as Western white pine and Lodgepole pine are tolerant to the pathogen while Ponderosa pine is resistant to it. Only hardwoods are known to be immune to the pathogen.
In woody plants, a tylosis is a bladder-like distension of a parenchyma cell into the lumen of adjacent vessels. The term tylosis summarises the physiological process and the resulting occlusion in the xylem of woody plants as response to injury or as protection from decay in heartwood. It is a key process in wall one of the compartmentalization of decay in trees (CODIT) and other woody plants.
A cambium, in plants, is a tissue layer that provides partially undifferentiated cells for plant growth. It is found in the area between xylem and phloem. A cambium can also be defined as a cellular plant tissue from which phloem, xylem, or cork grows by division, resulting in secondary thickening. It forms parallel rows of cells, which result in secondary tissues.
