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Agnostida is an order of arthropod which first developed near the end of the Early Cambrian period and thrived during the Middle Cambrian. They are present in the Lower Cambrian fossil record along with trilobites from the Redlichiida, Corynexochida, and Ptychopariida orders. The last agnostids went extinct in the Late Ordovician.
Trilobites are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period, and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 20,000 species having been described.
Nektaspida is an extinct order of soft-bodied arthropods proposed by Raymond in 1920; its taxonomic status is uncertain. Specimens are known from the early Cambrian to the upper Silurian periods. A Russian find in the Vendian of the White Sea (=Ediacaran), Keretsia, has in 2006 also been attributed to this order, which would make it the earliest arthropod found yet. Whittington (1985) placed the order in the Trilobita. Cotton & Braddy (2000) place it in a new "Trilobite clade" containing the Trilobita, recognizing the close affinities of the nektaspids to trilobites. However this necessitates the inclusion of genera that look very little like trilobites., it was formerly placed in the stem-group to the chelicerata subdivision of the Arthropoda phylum. However, it currently considered part of Artiopoda, the clade that contains trilobites and their close relatives.
Asaphida is a large, morphologically diverse order of trilobites found in marine strata dated from the Middle Cambrian until their extinction during the Silurian. Asaphida contains six superfamilies, but no suborders. Asaphids comprise some 20% of described fossil trilobites.
Isotelus is a genus of asaphid trilobites from the middle and upper Ordovician period, fairly common in the Northeastern United States, northwest Manitoba, southwestern Quebec and southeastern Ontario. One species, Isotelus rex, is currently the world's largest trilobite ever found as a complete fossil.
Arachnomorpha is a subdivision or clade of Arthropoda, comprising the monophyletic group formed by the trilobites, other great appendage arthropods and trilobite-like families, and a diverse sister clade including the chelicerates. Great debate is held on the position of the Pycnogonida, which are currently thought not to be placed in the immediate vicinity of the Chelicerata. Arachnomorpha are considered the sister group to the crustaceans, which are increasingly being accepted as members of the mandibulate clade.
Greenops is a mid-sized Devonian trilobite of the order Phacopida, subfamily Asteropyginae. They are mainly reported from the mid-Devonian Hamilton Group of upstate New York and southwestern Ontario. A similar-looking trilobite from Morocco is often mis-labelled Greenops. Greenops had schizocroidal eyes, large genal spines and short, sharp spines at the tip of each segment of the pygidium ("tail"). Greenops lived in warm, fairly deep water. In the Hamilton Group of New York, they are found with Eldredgeops, Dipleura and Bellacartwrightia, a trilobite that resembles Greenops but has much larger pygidial spines. In Ontario, they are found in the Widder Formation, which outcrops at Arkona, where they are, by far, the dominant trilobite.
Beecher's Trilobite Bed is a Konservat-Lagerstätte of Late Ordovician (Caradoc) age located within the Frankfort Shale in Cleveland's Glen, Oneida County, New York, USA. Only 3-4 centimeters thick, Beecher's Trilobite Bed has yielded numerous exceptionally preserved trilobites with the ventral anatomy and soft tissue intact, the soft tissue preserved by pyrite replacement. Pyritisation allows the use of X-rays to study fine detail of preserved soft body parts still within the host rock. Pyrite replacement of soft tissue is unusual in the fossil record; the only Lagerstätten thought to show such preservation were Beecher's Trilobite Bed, the Devonian Hunsrück Slates of Germany, and the Jurassic beds of La Voulte-sur-Rhône in France, although new locations are coming to light in New York state.
Ananaspis is a genus of trilobite in the order Phacopida, which existed in what is now the Czech Republic. It was described by Campbell in 1967, and the type species is Ananaspis fecundis, which was originally described as Phacops fecundus communis by Barrande in 1852.
Bainella is an extinct genus of Devonian trilobites from off the coast of Gondwana. Fossils were found in the Ponta Grossa Formation of Brazil, Belén, Icla and Gamoneda Formations of Bolivia and the Gydo, Gamka and Voorstehoek Formations of South Africa. It contains three species: B. africana, B. baini, and B. cristagalli.
Calymenella is a genus of trilobites in the order Phacopida, which existed in what is now France during the upper Ordovician. It was described by Bergeron in 1890, and the type species is Calymenella boisseli. The species was described from the Glauzy Formation in the Montagne Noire mountain range.
Encrinuroides is a genus of trilobites in the order Phacopida, that existed during the upper Ordovician in what is now Wales. It was described by Reed in 1931, and the type species is Encrinuroides sexcostata, which was originally described under the genus Cybele by Salter in 1848. It also contains the species, Encrinuroides enshiensis, Encrinuroides insularis, and Encrinuroides rarus. The type locality was in the Sholeshook Limestone Formation.
The Walcott–Rust quarry, in Herkimer County, New York, is an excellent example of an obrution Lagerstätte. Unique preservation of trilobite appendages resulted from early consolidation (cementation) of the surrounding rock, followed by spar filling of the interior cavity within the appendages. The presence of so many well preserved trilobites in one location alone qualifies the beds as an exceptional trilobite site, but the beds are further distinguished as the source of the first trilobites for which appendages were definitively described.
The cephalon is the head section of an arthropod. It is a tagma, i.e., a specialized grouping of arthropod segments. The word cephalon derives from the Greek κεφαλή (kephalē), meaning "head".
Homagnostus is a genus of trilobite in the order Agnostida, which existed in what is now north Wales. It was described by Howell in 1935. The genus was originally considered to be Agnostus pisiformis var. obesus.
Micragnostus is a genus of trilobite in the order Agnostida, which existed in what is now north Wales. It was described by Howell in 1935, and the type species is Micragnostus calvus, which was originally described as a species of Agnostus by Lake in 1906.
Bumastus is an extinct genus of corynexochid trilobites which existed from the Early Ordovician period to the Late Silurian period. They were relatively large trilobites, reaching a length of 6 in (15 cm). They were distinctive for their highly globular, smooth-surfaced exoskeleton. They possessed well-developed, large compound eyes and were believed to have dwelled in shallow-water sediments in life.
Lopha is a genus of marine bivalve molluscs in the family Ostreidae.
Lejopyge laevigata is a species of agnostid trilobite belonging to the genus Lejopyge. It existed during the Guzhangian to the Paibian Age of the Cambrian. It has a cosmopolitan distribution and is an important index fossil in biostratigraphy.
Paleontology in Wisconsin refers to paleontological research occurring within or conducted by people from the U.S. state of Wisconsin. The state has fossils from the Precambrian, much of the Paleozoic, and the later part of the Cenozoic. Most of the Paleozoic rocks are marine in origin. Because of the thick blanket of Pleistocene glacial sediment that covers the rock strata in most of the state, Wisconsin’s fossil record is relatively sparse. In spite of this, certain Wisconsin paleontological occurrences provide exceptional insights concerning the history and diversity of life on Earth.
References
- ↑ Jell, P.A. and Adrain, J.M. (2002) Available Generic Names for Trilobites, Memoirs of the Queensland Museum 48(2), ISSN 0079-8835, page 341.
