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Agnostida is an order of arthropod which have classically been seen as a group of highly modified trilobites, though some recent research has doubted this placement. Regardless, they appear to be close relatives as part of the Artiopoda. They are present in the Lower Cambrian fossil record along with trilobites from the Redlichiida, Corynexochida, and Ptychopariida orders, and were highly diverse throughout the Cambrian. Agnostidan diversity severely declined during the Cambrian-Ordovician transition, and the last agnostidans went extinct in the Late Ordovician.
Trilobites are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 20,000 species having been described.
Asaphida is a large, morphologically diverse order of trilobites found in marine strata dated from the Middle Cambrian until their extinction during the Silurian. Asaphida contains six superfamilies, but no suborders. Asaphids comprise some 20% of described fossil trilobites.
The Emu Bay Shale is a geological formation in Emu Bay, South Australia, containing a major Konservat-Lagerstätte. It is one of two in the world containing Redlichiidan trilobites. The Emu Bay Shale is dated as Cambrian Series 2, Stage 4, correlated with the upper Botomian Stage of the Lower Cambrian.
Isotelus is a genus of asaphid trilobites from the middle and upper Ordovician period, fairly common in the Northeastern United States, northwest Manitoba, southwestern Quebec and southeastern Ontario. One species, Isotelus rex, is currently the world's largest trilobite ever found as a complete fossil.
Arachnomorpha is a proposed subdivision or clade of Arthropoda, comprising the group formed by the trilobites and their close relatives (Artiopoda), Megacheira and chelicerates. Under this proposed classification scheme, Arachnomorpha is considered the sister group to Mandibulata.
In structural geology, a suture is a joining together along a major fault zone, of separate terranes, tectonic units that have different plate tectonic, metamorphic and paleogeographic histories. The suture is often represented on the surface by an orogen or mountain range. The term was borrowed from surgery where it describes the sewing together of two pieces of tissue, but the sutures of the skull, where separate plates of bone have fused, may be a better metaphor.
Greenops is a mid-sized Devonian trilobite of the order Phacopida, subfamily Asteropyginae. They are mainly reported from the mid-Devonian Hamilton Group of upstate New York and southwestern Ontario. A similar-looking trilobite from Morocco is often mis-labelled Greenops. Greenops had schizocroidal eyes, large genal spines and short, sharp spines at the tip of each segment of the pygidium ("tail"). Greenops lived in warm, fairly deep water. In the Hamilton Group of New York, they are found with Eldredgeops, Dipleura and Bellacartwrightia, a trilobite that resembles Greenops but has much larger pygidial spines. In Ontario, they are found in the Widder Formation, which outcrops at Arkona, where they are, by far, the dominant trilobite.
Beecher's Trilobite Bed is a Konservat-Lagerstätte of Late Ordovician (Caradoc) age located within the Frankfort Shale in Cleveland's Glen, Oneida County, New York, USA. Only 3-4 centimeters thick, Beecher's Trilobite Bed has yielded numerous exceptionally preserved trilobites with the ventral anatomy and soft tissue intact, the soft tissue preserved by pyrite replacement. Pyritisation allows the use of X-rays to study fine detail of preserved soft body parts still within the host rock. Pyrite replacement of soft tissue is unusual in the fossil record; the only Lagerstätten thought to show such preservation were Beecher's Trilobite Bed, the Devonian Hunsrück Slates of Germany, and the Jurassic beds of La Voulte-sur-Rhône in France, although new locations are coming to light in New York state.
Ananaspis is a genus of trilobite in the order Phacopida, which existed in what is now the Czech Republic. It was described by Campbell in 1967, and the type species is Ananaspis fecundis, which was originally described as Phacops fecundus communis by Barrande in 1852.
Calymenella is a genus of trilobites in the order Phacopida, which existed in what is now France during the upper Ordovician. It was described by Bergeron in 1890, and the type species is Calymenella boisseli. The species was described from the Glauzy Formation in the Montagne Noire mountain range.
Encrinuroides is a genus of trilobites in the order Phacopida, that existed during the upper Ordovician in what is now Wales. It was described by Reed in 1931, and the type species is Encrinuroides sexcostata, which was originally described under the genus Cybele by Salter in 1848. It also contains the species, Encrinuroides enshiensis, Encrinuroides insularis, and Encrinuroides rarus. The type locality was in the Sholeshook Limestone Formation.
The Walcott–Rust quarry, in Herkimer County, New York, is an excellent example of an obrution Lagerstätte. Unique preservation of trilobite appendages resulted from early consolidation (cementation) of the surrounding rock, followed by spar filling of the interior cavity within the appendages. The presence of so many well preserved trilobites in one location alone qualifies the beds as an exceptional trilobite site, but the beds are further distinguished as the source of the first trilobites for which appendages were definitively described.
The cephalon is the head section of an arthropod. It is a tagma, i.e., a specialized grouping of arthropod segments. The word cephalon derives from the Greek κεφαλή (kephalē), meaning "head".
Homagnostus is a genus of trilobite in the order Agnostida, which existed in what is now north Wales. It was described by Howell in 1935. The genus was originally considered to be Agnostus pisiformis var. obesus.
Micragnostus is a genus of trilobite in the order Agnostida, which existed in what is now north Wales. It was described by Howell in 1935, and the type species is Micragnostus calvus, which was originally described as a species of Agnostus by Lake in 1906.
Bumastus is an extinct genus of corynexochid trilobites which existed from the Early Ordovician period to the Late Silurian period. They were relatively large trilobites, reaching a length of 6 in (15 cm). They were distinctive for their highly globular, smooth-surfaced exoskeleton. They possessed well-developed, large compound eyes and were believed to have dwelled in shallow-water sediments in life.
Lejopyge laevigata is a species of agnostid trilobite belonging to the genus Lejopyge. It existed during the Guzhangian to the Paibian Age of the Cambrian. It has a cosmopolitan distribution and is an important index fossil in biostratigraphy.
Paleontology in Wisconsin refers to paleontological research occurring within or conducted by people from the U.S. state of Wisconsin. The state has fossils from the Precambrian, much of the Paleozoic, and the later part of the Cenozoic. Most of the Paleozoic rocks are marine in origin. Because of the thick blanket of Pleistocene glacial sediment that covers the rock strata in most of the state, Wisconsin’s fossil record is relatively sparse. In spite of this, certain Wisconsin paleontological occurrences provide exceptional insights concerning the history and diversity of life on Earth.
The Artiopoda is a grouping of extinct arthropods that includes trilobites and their close relatives. It was erected by Hou and Bergström in 1997 to encompass a wide diversity of arthropods that would traditionally have been assigned to the Trilobitomorpha. Hou and Bergström used the name Lamellipedia as a superclass to replace Trilobitomorpha that was originally erected at the subphylum level, which they considered inappropriate. Trilobites, in part due to their mineralising exoskeletons, are by far the most diverse and long lived members of the clade, with most records of other members, which lack mineralised exoskeletons, being from Cambrian deposits.
References
- ↑ Available Generic Names for Trilobites P.A. Jell and J.M. Adrain.
