Cobcrephora

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Cobcrephora
Scientific classification
Kingdom: Animalia
Phylum: Mollusca
Class: Polyplacophora
Genus:Cobcrephora

Cobcrephora is a genus of that resembles the Palaeoloricates, known from the Silurian of Gotland. [1] Its interpretation as a polyplacophoran is widely challenged. [1]

The Silurian is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at 443.8 million years ago (Mya), to the beginning of the Devonian Period, 419.2 Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by several million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when 60% of marine species were wiped out.

Gotland island and historical province in Sweden

Gotland is a province, county, municipality, and diocese of Sweden. It is Sweden's largest island. The province includes the islands of Fårö and Gotska Sandön to the north, as well as the Karlsö Islands to the west. The population is 58,595, of which about 23,600 live in Visby, the main town. The island of Gotland and the other areas of the province of Gotland make up less than one percent of Sweden's total land area.

Its mollusc shell is unique, because Cobcrephora was described on the basis of isolated phosphatic sclerites. [2] Its overlapping sclerites are arched and small, comprise two shell layers, and have lamellar projections. [1]

Mollusc shell exoskeleton of an animal in the phylum Mollusca

The molluscshell is typically a calcareous exoskeleton which encloses, supports and protects the soft parts of an animal in the phylum Mollusca, which includes snails, clams, tusk shells, and several other classes. Not all shelled molluscs live in the sea; many live on the land and in freshwater.

Related Research Articles

Sclerite hardened body part

A sclerite is a hardened body part. In various branches of biology the term is applied to various structures, but not as a rule to vertebrate anatomical features such as bones and teeth. Instead it refers most commonly to the hardened parts of arthropod exoskeletons and the internal spicules of invertebrates such as certain sponges and soft corals. In paleontology, a scleritome is the complete set of sclerites of an organism, often all that is known from fossil invertebrates.

<i>Tentaculites</i>

Tentaculites is an extinct genus of conical fossils of uncertain affinity, class Tentaculita, although it is not the only member of the class. It is known from Lower Ordovician to Upper Devonian deposits both as calcitic shells with a brachiopod-like microstructure and carbonaceous 'linings'. The "tentaculites" are also referred to as the styliolinids.

Aplacophora class of molluscs

Aplacophora is a monophyletic group of small, deep-water, exclusively benthic, marine molluscs found in all oceans of the world. All known modern forms are shell-less: only some extinct primitive forms possessed valves. The group comprises the two clades Solenogastres (Neomeniomorpha) and Caudofoveata (Chaetodermomorpha), which between them contain 28 families and about 320 species. The aplacophorans are traditionally considered ancestral to the other mollusc classes. However, the relationship between the two aplacophoran groups and to the other molluscan classes and to each other is as yet unclear.

Halkieriid Family of molluscs

The halkieriids are a group of fossil organisms from the Lower to Middle Cambrian. Their eponymous genus is Halkieria, which has been found on almost every continent in Lower to Mid Cambrian deposits, forming a large component of the small shelly fossil assemblages. The best known species is Halkieria evangelista, from the North Greenland Sirius Passet Lagerstätte, in which complete specimens were collected on an expedition in 1989. The fossils were described by Simon Conway Morris and John Peel in a short paper in 1990 in the journal Nature. Later a more thorough description was undertaken in 1995 in the journal Philosophical Transactions of the Royal Society of London and wider evolutionary implications were posed.

Burgsvik Beds

The Burgsvik Beds are a sequence of shallow marine limestones and sandstones found near the locality of Burgsvik in the southern part of Gotland, Sweden. The beds were deposited in the Upper Silurian period, around 420 million years ago, in warm, equatorial waters frequently ravaged by storms, in front of an advancing shoreline. The Burgsvik Formation comprises two members, the Burgsvik Sandstone and the Burgsvik Oolite.

The Lau event was the last of three relatively minor mass extinctions during the Silurian period. It had a major effect on the conodont fauna, but barely scathed the graptolites. It coincided with a global low point in sea level, is closely followed by an excursion in geochemical isotopes in the ensuing late Ludfordian faunal stage and a change in depositional regime.

Halwaxiida

Halwaxiida or halwaxiids is a proposed clade equivalent to the older orders Sachitida He 1980 and Thambetolepidea Jell 1981, loosely uniting scale-bearing Cambrian animals, which may lie in the stem group to molluscs or lophotrochozoa. Some palaeontologists question the validity of the Halwaxiida clade.

The Ireviken event was the first of three relatively minor extinction events during the Silurian period. It occurred at the Llandovery/Wenlock boundary. The event is best recorded at Ireviken, Gotland, where over 50% of trilobite species became extinct; 80% of the global conodont species also become extinct in this interval.

Geology of Gotland

Gotland is made up of a sequence of sedimentary rocks of a Silurian age, dipping to the south-east. The main Silurian succession of limestones and shales comprises thirteen units spanning 200–500 m (660–1,640 ft) of stratigraphic thickness, being thickest in the south, and overlies a 75–125 m (246–410 ft) thick Ordovician sequence. Precambrian shield rocks that underlie these sediments are found 400 to 500 meters sea level. Sedimentary rocks cropping out in Gotland were deposited in a shallow, hot and salty sea, on the edge of an equatorial continent. The water depth never exceeded 175–200 m (574–656 ft), and shallowed over time as bioherm detritus, and terrestrial sediments, filled the basin. Reef growth started in the Llandovery, when the sea was 50–100 m (160–330 ft) deep, and reefs continued to dominate the sedimentary record. Some sandstones are present in the youngest rocks towards the south of the island, which represent sand bars deposited very close to the shore line.

Since 1990 there has been intense debate among paleontologists about the evolution in the Early Cambrian period of the "super-phylum" Lophotrochozoa, which is thought to include the modern molluscs, annelid worms and brachiopods, as well as their evolutionary "aunts" and "cousins".

Graticula, formerly incorrectly named Craticula, is a genus of Palaeozoic coralline alga. They form the framework of reef rocks in the Silurian of Gotland, from the Högklint, Slite and Halla groups.

<i>Tryblidium reticulatum</i> species of mollusc (fossil)

Tryblidium reticulatum is an extinct species of a paleozoic Silurian monoplacophoran.

<i>Pilina unguis</i> species of mollusc (fossil)

Pilina unguis is an extinct species of Paleozoic Silurian monoplacophoran. It was first named as Tryblidium unguis and described by Gustaf Lindström in Latin from the Silurian deposits of Gotland in Sweden, in 1880.

<i>Helcionopsis radiatum</i> species of mollusc (fossil)

Helcionopsis radiatum is an extinct species of paleozoic monoplacophoran in the family Tryblidiidae.

<i>Pilina solarium</i> species of mollusc (fossil)

Pilina solarium is an extinct species of a paleozoic Silurian monoplacophoran. It was first named as Palaeacmaea solarium and described by Gustaf Lindström from Silurian of Gotland in Sweden in 1884.

Rothpletzella is a genus of calcimicrobe known from the Silurian of Gotland, the Devonian of France, as well as the Ordovician of China. It has been hypothesised to be a cyanobacterium, and shares morphological similarities with extant cyanobacteria.

<i>Septalites</i> genus of ctenophores

Septalites is a genus of cornulitid tubeworms. Their shells lack vesicular wall structure and have a smooth lumen filled with numerous transverse septa. They are externally covered with transverse ridges. Their fossils are known only from the Silurian of Gotland.

<i>Kulindroplax</i>

Kulindroplax perissokomos is a Silurian mollusk, known from a single fossil from the Wenlock Series Lagerstätte fauna of England. It lived during the Homerian Age. It is considered a basal aplacophoran. Unlike all modern aplacophorans, which are shell-less, Kulindroplax has a chiton-like shell, and it is considered a transitional fossil in the evolution of molluscs.

Phthipodochiton is an extinct genus of molluscs, known from several fossils from the upper Ordovician fauna of the Lady Burn Starfish beds of Girvan, Scotland. It shows a mixture of aplacophoran body plan and polyplacophoran-like valves, and it is an informative fossil in the evolution of aculiferan mollusks.

References

  1. 1 2 3 Cherns, L. (2004). "Early Palaeozoic diversification of chitons (Polyplacophora, Mollusca) based on new data from the Silurian of Gotland, Sweden". Lethaia. 37 (4): 445–456. doi:10.1080/00241160410002180.
  2. Bischoff, G. C. O. 1981. Cobcrephora n. g., representative of a new Polyplacophoran order phosphatoloricata, with calciumphosphatic shells. Senckenbergiana Lethaea, 61:173-215