Coeloscleritophora Temporal range: Cambrian-Ordovician | |
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Scientific classification | |
Kingdom: | Animalia |
Phylum: | incertae sedis |
Class: | † Coeloscleritophora |
Groups included | |
The Coelosclerithophorans are a polyphyletic group of organisms bearing hollow sclerites made of aragonite, and with a supposedly distinctive microstructure. [1]
Their skeletons may be homologous to those of the molluscs, [2] and Halkieria looks very much like a polyplacophoran. The shells of this group and the molluscs appear to have originated in the same fashion. [2] However, there are several reasons to doubt this relationship, [3] many of which may not pass muster. [4]
There are two groups within the coeloscleritophorans: the Sachitids, to which Halkieria and Wiwaxia belong, and the Chancellorids. The Ediacaran fossil Ausia has been touted as an ancestral 'intermediate' between both of these groups. [5]
These are common constituents of Lower Cambrian small shelly fossil assemblages.
Hyoliths are animals with small conical shells, known as fossils from the Palaeozoic Era. They are lophophorates, a group which includes the brachiopods.
Wiwaxia is a genus of soft-bodied animals that were covered in carbonaceous scales and spines that protected it from predators. Wiwaxia fossils – mainly isolated scales, but sometimes complete, articulated fossils – are known from early Cambrian and middle Cambrian fossil deposits across the globe. The living animal would have measured up to 5 cm (2 inch) when fully grown, although a range of juvenile specimens are known, the smallest being 2 millimetres (0.079 in) long.
The halkieriids are a group of fossil organisms from the Lower to Middle Cambrian. Their eponymous genus is Halkieria, which has been found on almost every continent in Lower to Mid Cambrian deposits, forming a large component of the small shelly fossil assemblages. The best known species is Halkieria evangelista, from the North Greenland Sirius Passet Lagerstätte, in which complete specimens were collected on an expedition in 1989. The fossils were described by Simon Conway Morris and John Peel in a short paper in 1990 in the journal Nature. Later a more thorough description was undertaken in 1995 in the journal Philosophical Transactions of the Royal Society of London and wider evolutionary implications were posed.
Praecambridium sigillum is an extinct organism that superficially resembles a segmented trilobite-like arthropod. It was originally described as being a trilobite-like arthropod, though the majority of experts now place it within the Proarticulata as a close relative of the much larger Yorgia. It is from the Late Ediacaran deposit of Ediacara Hills, Australia, about 555 million years ago. On average, P. sigillum had at least 5 pairs of segments, with each unit becoming progressively larger as they approach the cephalon-like head.
Halwaxiida or halwaxiids is a proposed clade equivalent to the older orders Sachitida He 1980 and Thambetolepidea Jell 1981, loosely uniting scale-bearing Cambrian animals, which may lie in the stem group to molluscs or lophotrochozoa. Some palaeontologists question the validity of the Halwaxiida clade.
Sinotubulites is a genus of small, tube-shaped shelly fossils from the Ediacaran period. It is often found in association with Cloudina.
Scenella is an extinct genus of fossil invertebrate animal which is generally considered to be a mollusc; at various times it has been suggested that this genus belongs with the gastropods, the monoplacophorans, or the helcionellids, although no firm association with any of these classes has been established. An affinity with the hydrozoa has been considered, although some authors oppose this hypothesis. A gastropod affinity is defended on the basis of six pairs of internal muscle scars, whilst the serially-repeated nature of these scars suggests to other authors a monoplacophoran affinity. However the specimens showing this scarring have not been convincingly shown to belong to the genus Scenella. A similarity to the Ediacaran Ovatoscutum has also been drawn.
The small shelly fauna, small shelly fossils (SSF), or early skeletal fossils (ESF) are mineralized fossils, many only a few millimetres long, with a nearly continuous record from the latest stages of the Ediacaran to the end of the Early Cambrian Period. They are very diverse, and there is no formal definition of "small shelly fauna" or "small shelly fossils". Almost all are from earlier rocks than more familiar fossils such as trilobites. Since most SSFs were preserved by being covered quickly with phosphate and this method of preservation is mainly limited to the late Ediacaran and early Cambrian periods, the animals that made them may actually have arisen earlier and persisted after this time span.
Since 1990 there has been intense debate among paleontologists about the evolution in the Early Cambrian period of the "super-phylum" Lophotrochozoa, which is thought to include the modern molluscs, annelid worms and brachiopods, as well as their evolutionary "aunts" and "cousins".
Anabarites is a problematic lower Cambrian genus, and is one of the small shelly fossils. It was abundant in the early Tommotian and is also found in the Nemakit-Daldynian. The fossils represent the triradially symmetrical mineralised tube in which the organism dwelt; it was sedentary. It is named after the Anabar region in Yakutia, Russia; its name does not imply 'heavy'.
Micropilina is a genus of monoplacophoran molluscs. They are very small, mostly deepwater animals which have a superficially limpet-like shell.
The Lower-Middle Cambrian animal Rhabdotubus is the earliest known pterobranch. It bears strong similarity to the graptolites - indeed for some time it was misclassified as a dendroid graptolite.
Stenothecidae is an extinct family of fossil univalved Cambrian molluscs which may be either gastropods or monoplacophorans.
The Kirengellids are a group of problematic Cambrian fossil shells of marine organisms. The shells bear a number of paired muscle scars on the inner surface of the valve.
Anabarella is a species of bilaterally-flattened monoplacophoran mollusc, with a morphological similarity to the rostroconchs. Its shell preserves evidence of three mineralogical textures on its outer surface: it is polygonal near the crest of the shell, subsequently changing to both spiny and stepwise. Its internal microstructure is calcitic and semi-nacreous. Its name reflects its provenance from Anabar, Siberia. It has been interpreted as ancestral to the rostroconchs, and has been aligned to the Helcionellidae.
The cephalopods have a long geological history, with the first nautiloids found in late Cambrian strata, and purported stem-group representatives present in the earliest Cambrian lagerstätten.
Stenothecoida is a taxon of bivalved fossils from the Early to middle Cambrian period. They look a bit like brachiopods or bivalve molluscs.
Mobergella is a millimetric Lower Cambrian shelly fossil of unknown affinity, usually preserved in phosphate and particularly well known from Swedish strata, where it is diagnostic of lowermost Cambrian rocks. Originally interpreted as a monoplacophoran, the circular, cap-shaped shell resembles a hyolith operculum, with concentric rings on its upper surface, and seven pairs of internal muscle scars. It is never found in association with a conch, and its affinity therefore remains undetermined. Nevertheless, its heavy musculature does seem to indicate that it functioned as an operculum.
Watsonella is a genus of 'mollusc' known from early (Terreneuvian) Cambrian strata.
Proconodontus is an extinct genus of conodonts in the family Proconodontidae. The specimens are found in Cambrian formations.