Dryandra subg. Hemiclidia is an obsolete plant taxon that encompassed material that is now included in Banksia . Published at genus rank as Hemiclidia by Robert Brown in 1830, it was set aside by George Bentham in 1870, but reinstated at subgenus rank by Alex George in 1996. In 2007, all Dryandra species were transferred into Banksia at series rank, and the infrageneric Dryandra taxa, including D. subg. Hemiclidia, were set aside.
In 1810, Robert Brown published the genus Dryandra in his On the Proteaceae of Jussieu . Thirteen species were published, including Dryandra falcata (now Banksia falcata ), but no infrageneric arrangement was proffered. [1] Twenty years later, Brown published a further eleven species and the first infrageneric arrangement in his Supplementum primum prodromi florae Novae Hollandiae . By this time, Brown had observed the tendency in D. falcata for one of the two ovules in each follicle to abort, thereafter developing into a winglike appendage to the seed separator. On this basis, he transferred the species into a monotypic genus, which he named Hemiclidia, [2] from the Greek hemi ("half") and perhaps kleidos ("barrier", "means of closing"). [3]
Hemiclidia was retained in the 1856 arrangement of Carl Meissner, [4] but discarded by George Bentham in his 1870 revision of the genus. Bentham correctly observed that the abortion of seeds occurs in other Dryandra species, and is a diagnostically unimportant character:
"as far as I can understand the characters given, the difference in the fruit upon which the genus Hemiclidia was founded is merely the result of the abortion of one ovule, which occurs occasionally or perhaps constantly in one or two other species of Dryandra." [5]
In 1996, Alex George published a thorough revision of Dryandra in preparation of his monograph on that genus for the Flora of Australia series. He segregated D. falcata and the newly published D. glauca (now Banksia glaucifolia ) into a subgenus based on their unusual follicles, which are small, hairy and pliable. Needing a name for the subgenus, George noted that Hemiclidia was available, and had an appropriate type species in D. falcata. He therefore demoted Hemiclidia to subgenus rank as Dryandra subg. Hemiclidia (R.Br.) A.S.George. [6] Its placement and circumscription in George's taxonomic arrangement of Dryandra may be summarised as follows: [6] [7]
This arrangement remained current until 2007, when Austin Mast and Kevin Thiele transferred Dryandra into Banksia. They were not yet ready, however, to tender an infrageneric arrangement encompassing Dryandra, so as an interim measure they transferred Dryandra into Banksia at series rank. This minimised the nomenclatural disruption of the transfer, but also caused George's rich infrageneric arrangement to be set aside. Thus under the interim arrangements implemented by Mast and Thiele, D. subg. Hemiclidia has been set aside. [8]
Banksia subg. Isostylis is a subgenus of Banksia. It contains three closely related species, all of which occur only in Southwest Western Australia. Members of subgenus Isostylis have dome-shaped flower heads that are superficially similar to those of B. ser. Dryandra, but structurally more like reduced versions of the "flower spikes" characteristic of most other Banksia taxa.
Banksia fraseri is a species of shrub that is endemic to Western Australia. It has hairy stems, broadly linear pinnatisect leaves with between four and eighteen sharply-pointed lobes on each side, between eighty and one hundred pink to cream-coloured flowers and wege-shaped follicles.
Banksia acuminata is a rare prostrate shrub endemic to south-west Western Australia. It was published in 1848 as Dryandra preissii, but transferred into Banksia as B. acuminata in 2007.
Banksia splendida, commonly known as shaggy dryandra, is a species of shrub that is endemic to the southwest of Western Australia. It has sharply-pointed linear leaves that are woolly on the lower surface, cream-coloured and maroon or yellow flowers in heads of between 65 and 115, and later up to eight egg-shaped follicles in each head.
Dryandra subg. Diplophragma is an obsolete subgenus within the former genus Dryandra. It was first published by Robert Brown in 1830, but was discarded by George Bentham in 1870. It was reinstated with a new circumscription by Alex George in 1996, but was ultimately discarded again in 2007 when Austin Mast and Kevin Thiele sunk Dryandra into Banksia.
Banksia sect. Eubanksia is an obsolete section of Banksia. There have been two circumscriptions, one of which is synonymous with the recently abandoned B. subg. Banksiasensu Alex George, the other having no modern equivalent.
Robert Brown's taxonomic arrangement of Dryandra was the first arrangement of what is now Banksia ser. Dryandra. His initial arrangement was published in 1810, and a further arrangement, including an infrageneric classification, followed in 1830. Aspects of Brown's arrangements can be recognised in the later arrangements of George Bentham and Alex George.
Banksia spinulosa var. collina is a shrub that grows along the east coast of Australia, in Queensland and New South Wales. Commonly known as Hill Banksia or Golden Candlesticks, it is a taxonomic variety of B. spinulosa. It is a popular garden plant widely sold in nurseries.
George Bentham's taxonomic arrangement of Dryandra was published in 1870, in Volume 5 of Bentham's Flora Australiensis. It replaced the 1856 arrangement of Carl Meissner, and stood for over a century before being replaced by the 1996 arrangement of Alex George.
Carl Meissner's taxonomic arrangement of Dryandra, now Banksia ser. Dryandra, was published in 1856 as part of his chapter on the Proteaceae in A. P. de Candolle's Prodromus systematis naturalis regni vegetabilis. It replaced the 1830 arrangement of Robert Brown, and remained current until superseded by the 1870 arrangement of George Bentham.
Banksia spinulosa var. spinulosa is a shrub that grows along the east coast of Australia, in Queensland and New South Wales.
Dryandra ser. Niveae is an obsolete series within the former genus Dryandra. It was first published by George Bentham in 1870, and was given a new circumscription by Alex George in 1996, but was ultimately discarded in 2007 when Austin Mast and Kevin Thiele sunk Dryandra into Banksia.
Dryandra subg. Dryandra is an obsolete clade of plant. It was a series within the former genus Dryandra. The name was first published at sectional rank as Dryandra verae in 1830, before being renamed Eudryandra in 1847, the replaced by the autonym at subgenus rank in 1996. It was ultimately discarded in 2007 when Austin Mast and Kevin Thiele sunk Dryandra into Banksia.
Dryandra ser. Floribundae is an obsolete series within the former genus Dryandra. It was first published by George Bentham in 1870, and was given a new circumscription by Alex George in 1996, but was ultimately discarded in 2007 when Austin Mast and Kevin Thiele sunk Dryandra into Banksia.
Dryandra ser. Armatae is an obsolete series within the former genus Dryandra. It was first published by George Bentham in 1870, and was given a new circumscription by Alex George in 1996, but was ultimately discarded in 2007 when Austin Mast and Kevin Thiele sunk Dryandra into Banksia.
Dryandra ser. Ilicinae is an obsolete series within the former genus Dryandra. It was first published by Carl Meissner in 1856, but was discarded by George Bentham in 1870. It was reinstated with a new circumscription by Alex George in 1996, but was ultimately discarded again in 2007 when Austin Mast and Kevin Thiele sunk Dryandra into Banksia.
Dryandra ser. Plumosae is an obsolete series within the former genus Dryandra. It was published by Alex George in 1996, but discarded in 2007 when Austin Mast and Kevin Thiele sank Dryandra into Banksia.
Dryandra ser. Capitellatae is an obsolete series within the former genus Dryandra. It was published by Alex George in 1996, but discarded in 2007 when Austin Mast and Kevin Thiele sank Dryandra into Banksia.
Dryandra ser. Aphragma is an obsolete series within the former genus Dryandra. It was first published at sectional rank by Robert Brown in 1830, and was retained at that rank until 1999, when Alex George demoted it to a series. It was discarded in 2007 when Austin Mast and Kevin Thiele sank Dryandra into Banksia.
