Dryandra subg. Dryandra

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Dryandra formosa (now Banksia formosa), the type species of Dryandra subg. Dryandra Dryandra formosa from Flora Australasica.jpg
Dryandra formosa (now Banksia formosa ), the type species of Dryandra subg. Dryandra

Dryandra subg. Dryandra is an obsolete clade of plant. It was a series within the former genus Dryandra (now Banksia ser. Dryandra). The name was first published at sectional rank as Dryandra verae in 1830, before being renamed Eudryandra in 1847, the replaced by the autonym at subgenus rank in 1996. It was ultimately discarded in 2007 when Austin Mast and Kevin Thiele sunk Dryandra into Banksia .

Contents

Brown's Dryandra verae

Dryandra verae ("True Dryandra") was published by Brown in his 1830 Supplementum primum Prodromi florae Novae Hollandiae . Brown's arrangement of Dryandra split a single species out into a separate genus, and divided the remaining Dryandra species into three groups according to what Brown perceived to be variations in the number of seed separators. He allowed for these groups to be treated at subgenus or section rank, [1] but they are now treated as having been published as sections. [2] Dryandra verae was defined as containing those species with a single seed separator per follicle. Its placement and circumscription may be summarised as follows: [1]

Dryandra (now B. ser. Dryandra)
Dryandra verae
D. floribunda (now B. sessilis )
D. cuneata (now B. obovata )
D. armata (now B. armata )
D. Serra (now B. serra )
D. concinna (now B. concinna )
D. foliolata (now B. foliolata )
D. squarrosa (now B. squarrosa )
D. formosa (now B. formosa )
D. mucronulata (now B. mucronulata )
D. Baxteri (now B. biterax )
D. plumosa (now B. plumosa )
D. obtusa (now B. obtusa )
D. arctotidis (now B. arctotidis )
D. nivea (now B. nivea )
D. Fraseri (now B. fraseri )
D. longifolia (now B. prolata )
D. seneciifolia (now B. seneciifolia )
D. tenuifolia (now B. tenuis )
D. sect. Diplophragma (1 species)
D. sect. Aphragma (4 species)
Hemiclidia (1 species)

Meissner's Eudryandra

Dryandra verae was renamed Eudryandra by Stephan Endlicher in 1847, [3] and this name was used in the 1856 arrangement of Carl Meissner. Meissner retained all three of Brown's sections, and further divided D. sect. Eudryandra in 8 subgroups of unstated rank designated by the section sign ("§").The placement and circumscription of D. sect. Eudryandra in Meissner's arrangement thus looks like this: [4]

Dryandra (now Banksia ser. Dryandra)
D. sect. Eudryandra
D. § Ilicinæ (4 species, 2 varieties)
D. praemorsa (now B. undata )
D. praemorsa var. elongata (now B. undata var. undata)
D. quercifolia (now B. heliantha )
D. cuneata (now B. obovata )
D. floribunda (now B. sessilis var. sessilis)
D. floribunda var. cordata (now B. sessilis var. cordata)
D. § Runcinatæ (5 species)
D. armata (now B. armata )
D. carduacea (now B. squarrosa subsp. squarrosa)
D. runcinata (now B. r. subsp. rufa)
D. nobilis (now B. nobilis )
D. plumosa (now B. plumosa )
D. § Serratæ (11 species, 3 varieties)
D. stupposa (now B. stuposa )
D. serra (now B. serra )
D. mucronulata (now B. mucronulata )
D. foliolata (now B. foliolata )
D. mutica (now B. foliolata)
D. obtusa (now B. obtusa )
D. Baxteri (now B. biterax )
D. formosa (now B. formosa )
D. nivea (now B. nivea )
D. nivea var. venosa (now B. nivea)
D. nivea var. subevenia (now B. dallanneyi var. dallanneyi)
D. brownii (now B. brunnea )
D. lindleyana (now B. dallanneyi )
D. § Marginatæ (7 species)
D. stenoprion (now B. stenoprion )
D. elegans (now B. tenuis var. tenuis)
D. pulchella (now B. bella )
D. Kippistiana (now B. kippistiana )
D. Shuttleworthiana (now B. shuttleworthiana )
D. sclerophylla (now B. sclerophylla )
D. squarrosa (now B. squarrosa )
D. § Pectinatæ (6 species)
D. serratuloides (now B. serratuloides )
D. nana (now B. nana )
D. arctotidis (now B. arctotidis )
D. tortifolia (now B. tortifolia )
D. Fraseri (now B. fraseri )
D. cirsioides (now B. cirsioides )
D. § Decurrentes (11 species)
D. seneciifolia (now B. seneciifolia )
D. horrida (now B. horrida )
D. concinna (now B. concinna )
D. vestita (now B. vestita )
D. Hewardiana (now B. hewardiana )
D. longifolia (now B. prolata )
D. comosa (now B. comosa )
D. proteoides (now B. proteoides )
D. tenuifolia (now B. tenuis )
D. ferruginea (now B. rufa )
D. cryptocephala (now B. seneciifolia )
D. § Acrodontæ (2 species)
D. carlinoides (now B. carlinoides )
D. tridentata (now B. tridentata )
D. § Haplophyllæ (1 species)
D. speciosa (now B. splendida )
D. sect. Diplophragma (2 species)
D. sect. Aphragma (4 species, 4 varieties)
Hemiclidia (1 species)

Bentham's Eudryandra

When George Bentham published his arrangement of Dryandra in his 1870 Flora Australiensis , he retained D. sect. Eubanksia, but discarded all eight of Meissner's groups within that section, replacing them with seven series defined using floral characters: [5]

Dryandra (now Banksia ser. Dryandra)
D. sect. Eudryandra
D. ser. Armatae (7 species)
D. quercifolia (now B. heliantha )
D. praemorsa (now B. undata )
D. cuneata (now B. obovata )
D. falcata (now B. falcata )
D. armata (now B. armata )
D. longifolia (now B. prolata )
D. Fraseri (now B. fraseri )
D. ser. Floribundae (5 species, 2 varieties)
D. floribunda (now B. sessilis var. sessilis)
D. floribunda var. major (now B. sessilis var. cordata)
D. carduacea (now B. squarrosa subsp. squarrosa)
D. carlinoides (now B. carlinoides )
D. polycephala (now B. polycephala )
D. Kippistiana (now B. kippistiana )
D. ser. Concinnae (4 species)
D. squarrosa (now B. squarrosa )
D. serra (now B. serra )
D. concinna (now B. concinna )
D. foliolata (now B. foliolata )
D. ser. Formosae (5 species)
D. stupposa (now B. stuposa )
D. nobilis (now B. nobilis )
D. mucronulata (now B. mucronulata )
D. formosa (now B. formosa )
D. Baxteri (now B. biterax )
D. ser. Niveæ (4 species, 2 varieties)
D. nivea (now B. nivea )
D. arctotidis (now B. arctotidis )
D. arctotidis var. tortifolia (now B. tortifolia )
D. nana (now B. nana )
D. Preissii (now B. acuminata )
D. ser. Obvallatae (12 species)
D. sclerophylla (now B. sclerophylla )
D. pulchella (now B. bella )
D. plumosa (now B. plumosa )
D. seneciifolia (now B. seneciifolia )
D. vestita (now B. vestita )
D. cirsioides (now B. cirsioides )
D. Hewardiana (now B. hewardiana )
D. patens (now B. hewardiana)
D. conferta (now B. densa )
D. horrida (now B. horrida )
D. serratuloides (now B. serratuloides )
D. comosa (now B. comosa )
D. ser. Gymnocephalae (3 species)
D. Shuttleworthiana (now B. shuttleworthiana )
D. speciosa (now B. splendida )
D. tridentata (now B. tridentata )
D. sect. Aphragma (7 species, 2 varieties)

According to George

By the time Alex George published his revision of Dryandra in 1996, the rules of botanical nomenclature mandated that any infrageneric taxon that contained the type species of the genus be given an autonym. Thus D. sect. Eudryandra was treated as illegitimate, and when George published essentially the same taxon at subgenus rank, it was instead named D. subg. Dryandra. This was by far the largest of George's three subgenera, containing all but three Dryandra species. George further divided it into 24 series, reviving most of Meissner's and Bentham's names in the process. [6] The placement and circumscription of D. subg. Dryandra in George's arrangement, as amended in 1999 [7] and 2005, [8] may be summarised as follows:

Dryandra (now Banksia ser. Dryandra)
D. subg. Dryandra
D. ser. Floribundae
D. sessilis (now B. sessilis )
D. sessilis var. sessilis (now B. sessilis var. sessilis)
D. sessilis var. flabellifolia (now B. sessilis var. flabellifolia)
D. sessilis var. cordata (now B. sessilis var. cordata)
D. sessilis var. cygnorum (now B. sessilis var. cygnorum)
D. ser. Armatae
D. cuneata (now B. obovata )
D. fuscobractea (now B. fuscobractea )
D. armata (now B. armata )
D. armata var. armata (now B. armata var. armata)
D. armata var. ignicida (now B. armata var. ignicida)
D. prionotes (now B. prionophylla
D. arborea (now B. arborea )
D. hirsuta (now B. hirta )
D. pallida (now B. pallida )
D. purdieana (now B. purdieana )
D. xylothemelia (now B. xylothemelia )
D. cirsioides (now B. cirsioides )
D. acanthopoda (now B. acanthopoda )
D. squarrosa (now B. squarrosa )
D. squarrosa subsp. squarrosa (now B. squarrosa subsp. squarrosa)
D. squarrosa subsp. argillacea (now B. squarrosa subsp. argillacea)
D. hewardiana (now B. hewardiana )
D. wonganensis (now B. wonganensis )
D. trifontinalis (now B. trifontinalis )
D. stricta (now B. strictifolia )
D. echinata (now B. echinata )
D. polycephala (now B. polycephala )
D. subpinnatifida (now B. subpinnatifida )
D. subpinnatifida var. subpinnatifida (now B. subpinnatifida var. subpinnatifida)
D. subpinnatifida var. imberbis (now B. subpinnatifida var. imberbis)
D. longifolia (now B. prolata )
D. longifolia subsp. longifolia (now B. prolata subsp. prolata)
D. longifolia subsp. calcicola (now B. prolata subsp. calcicola)
D. longifolia subsp. archeos (now B. prolata subsp. archeos)
D. borealis (now B. borealis )
D. borealis subsp. borealis (now B. borealis subsp. borealis)
D. borealis subsp. elatior (now B. borealis subsp. elatior)
D. ser. Marginatae
D. pulchella (now B. bella )
D. ser. Folliculosae
D. fraseri (now B. fraseri )
D. fraseri var. fraseri (now B. fraseri var. fraseri)
D. fraseri var. crebra (now B. fraseri var. crebra)
D. fraseri var. effusa (now B. fraseri var. effusa)
D. fraseri var. ashbyi (now B. fraseri var. ashbyi)
D. fraseri var. oxycedra (now B. fraseri var. oxycedra)
D. ser. Acrodontae
D. sclerophylla (now B. sclerophylla )
D. kippistiana (now B. kippistiana )
D. kippistiana var. kippistiana (now B. kippistiana var. kippistiana)
D. kippistiana var. paenepeccata (now B. kippistiana var. paenepeccata)
D. carlinoides (now B. carlinoides )
D. tridentata (now B. tridentata )
D. ser. Capitellatae
D. serratuloides (now B. serratuloides )
D. serratuloides subsp. serratuloides (now B. serratuloides subsp. serratuloides)
D. serratuloides subsp. perissa (now B. serratuloides subsp. perissa)
D. meganotia (now B. meganotia )
D. ser. Ilicinae
D. praemorsa (now B. undata )
D. praemorsa var. praemorsa (now B. undata var. undata)
D. praemorsa var. splendens (now B. undata var. splendens)
D. quercifolia (now B. heliantha )
D. anatona (now B. anatona )
D. ser. Dryandra
D. formosa (now B. formosa )
D. nobilis (now B. nobilis )
D. nobilis subsp. nobilis (now B. nobilis subsp. nobilis)
D. nobilis subsp. fragrans (now B. nobilis subsp. fragrans)
D. stuposa (now B. stuposa )
D. ser. Foliosae
D. mucronulata (now B. mucronulata )
D. mucronulata subsp. mucronulata (now B. mucronulata subsp. mucronulata)
D. mucronulata subsp. retrorsa (now B. mucronulata subsp. retrorsa)
D. baxteri (now B. biterax )
D. foliosissima (now B. foliosissima )
D. ser. Decurrentes
D. comosa (now B. comosa )
D. ser. Tenuifoliae
D. tenuifolia (now B. tenuis )
D. tenuifolia var. tenuifolia (now B. tenuis var. tenuis)
D. tenuifolia var. reptans (now B. tenuis var. reptans)
D. obtusa (now B. obtusa )
D. ser. Runcinatae
D. ferruginea (now B. rufa )
D. ferruginea subsp. magma (now B. rufa subsp. magma)
D. ferruginea subsp. tutanningensis (now B. rufa subsp. tutanningensis)
D. ferruginea subsp. ferruginea (now B. rufa subsp. rufa)
D. ferruginea subsp. pumila (now B. rufa subsp. pumila)
D. ferruginea subsp. obliquiloba (now B. rufa subsp. obliquiloba)
D. ferruginea subsp. chelomacarpa (now B. rufa subsp. chelomacarpa)
D. ferruginea subsp. flavescens (now B. rufa subsp. flavescens)
D. corvijuga (now B. corvijuga )
D. epimicta (now B. epimicta )
D. proteoides (now B. proteoides )
D. ser. Triangulares
D. drummondii (now B. drummondii )
D. drummondii subsp. drummondii (now B. drummondii subsp. drummondii)
D. drummondii subsp. hiemalis (now B. drummondii subsp. hiemalis)
D. drummondii subsp. macrorufa (now B. drummondii subsp. macrorufa)
D. octotriginta (now B. octotriginta )
D. catoglypta (now B. catoglypta )
D. ser. Aphragma
D. pteridifolia (now B. pteridifolia )
D. pteridifolia subsp. inretita (now B. pteridifolia subsp. inretita)
D. pteridifolia subsp. pteridifolia (now B. pteridifolia subsp. pteridifolia)
D. pteridifolia subsp. vernalis (now B. pteridifolia subsp. vernalis)
D. fililoba (now B. fililoba )
D. shanklandiorum (now B. shanklandiorum )
D. nervosa (now B. alliacea )
D. blechnifolia (now B. pellaeifolia )
D. porrecta (now B. porrecta )
D. aurantia (now B. aurantia )
D. calophylla (now B. calophylla )
D. lepidorhiza (now B. lepidorhiza )
D. ser. Ionthocarpae
D. ionthocarpa (now B. ionthocarpa )
D. ionthocarpa subsp. ionthocarpa (now B. ionthocarpa subsp. ionthocarpa)
D. ionthocarpa subsp. chrysophoenix (now B. ionthocarpa subsp. chrysophoenix)
D. ser. Inusitatae
D. idiogenes (now B. idiogenes )
D. ser. Subulatae
D. subulata (now B. subulata )
D. ser. Gymnocephalae
D. cynaroides (now B. cynaroides )
D. erythrocephala (now B. erythrocephala )
D. erythrocephala var. erythrocephala (now B. erythrocephala var. erythrocephala)
D. erythrocephala var. inopinata (now B. erythrocephala var. inopinata)
D. horrida (now B. horrida )
D. vestita (now B. vestita )
D. viscida (now B. viscida )
D. mimica (now B. mimica )
D. speciosa (now B. splendida )
D. speciosa subsp. speciosa (now B. speciosa subsp. splendida)
D. speciosa subsp. macrocarpa (now B. speciosa subsp. macrocarpa)
D. shuttleworthiana (now B. shuttleworthiana )
D. ser. Plumosae
D. plumosa (now B. plumosa )
D. plumosa subsp. plumosa (now B. plumosa subsp. plumosa)
D. plumosa subsp. denticulata (now B. plumosa subsp. denticulata)
D. pseudoplumosa (now B. pseudoplumosa )
D. montana (now B. montana )
D. ser. Concinnae
D. concinna (now B. concinna )
D. serra (now B. serra )
D. foliolata (now B. foliolata )
D. ser. Obvallatae
D. fasciculata (now B. fasciculata )
D. conferta (now B. densa )
D. conferta var. conferta (now B. densa var. densa)
D. conferta var. parva (now B. densa var. parva)
D. columnaris (now B. columnaris )
D. platycarpa (now B. platycarpa )
D. seneciifolia (now B. seneciifolia )
D. rufistylis (now B. rufistylis )
D. insulanemorecincta (now B. insulanemorecincta )
D. ser. Pectinatae
D. nana (now B. nana )
D. ser. Acuminatae
D. preissii (now B. acuminata )
D. ser. Niveae
D. arctotidis (now B. arctotidis )
D. tortifolia (now B. tortifolia )
D. stenoprion (now B. stenoprion )
D. cypholoba (now B. cypholoba )
D. lindleyana (now B. dallanneyi )
D. lindleyana subsp. lindleyana (now B. dallanneyi subsp. dallanneyi)
D. lindleyana subsp. pollosta (now B. dallanneyi subsp. pollosta)
D. lindleyana subsp. media (now B. dallanneyi subsp. media)
D. lindleyana subsp. agricola (now B. dallanneyi subsp. agricola)
D. lindleyana subsp. sylvestris (now B. dallanneyi subsp. sylvestris)
D. brownii (now B. brunnea )
D. nivea (now B. nivea )
D. nivea subsp. nivea (now B. nivea subsp. nivea)
D. nivea subsp. uliginosa (now B. nivea subsp. uliginosa)
D. subg. Hemiclidia (2 species)
D. subg. Diplophragma (1 species)

Recent developments

Since 1998, Austin Mast has been publishing results of ongoing cladistic analyses of DNA sequence data for the subtribe Banksiinae. His analyses have provided compelling evidence of the paraphyly of Banksia with respect to Dryandra ; that is, it seems that Dryandra arose from within the ranks of Banksia. [9] [10] [11] Early in 2007, Mast and Kevin Thiele initiated a rearrangement of Banksia by sinking Dryandra into it as B. ser. Dryandra. This transfer necessitated the setting aside of George's infrageneric arrangement of Dryandra; thus D. subg. Dryandra is no longer current. Mast and Thiele have foreshadowed publishing a full arrangement once DNA sampling of Dryandra is complete. [12]

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Georges taxonomic arrangement of Dryandra 1996 arrangement of the Australian endemic plant series Dryandra in the genus Banksia

Alex George's taxonomic arrangement of Dryandra was the first modern-day arrangement of that taxon. First published in Nuytsia in 1996, it superseded the arrangement of George Bentham, which had stood for over a hundred years; it would later form the basis for George's 1999 treatment of Dryandra for the Flora of Australia. In accordance with contemporary thinking, George treated Dryandra as a genus, dividing it into three subgenera, the largest of which was divided into 24 series. The arrangement stood until 2007, when Dryandra was transferred into Banksia as B. ser. Dryandra. No alternative has yet been proposed.

Robert Brown's taxonomic arrangement of Dryandra was the first arrangement of what is now Banksia ser. Dryandra. His initial arrangement was published in 1810, and a further arrangement, including an infrageneric classification, followed in 1830. Aspects of Brown's arrangements can be recognised in the later arrangements of George Bentham and Alex George.

Benthams taxonomic arrangement of Dryandra 1870 arrangement of the Australian endemic plant series Dryandra in the genus Banksia

George Bentham's taxonomic arrangement of Dryandra was published in 1870, in Volume 5 of Bentham's Flora Australiensis. It replaced the 1856 arrangement of Carl Meissner, and stood for over a century before being replaced by the 1996 arrangement of Alex George.

Carl Meissner's taxonomic arrangement of Dryandra, now Banksia ser. Dryandra, was published in 1856 as part of his chapter on the Proteaceae in A. P. de Candolle's Prodromus systematis naturalis regni vegetabilis. It replaced the 1830 arrangement of Robert Brown, and remained current until superseded by the 1870 arrangement of George Bentham.

Dryandra ser. Niveae is an obsolete series within the former genus Dryandra. It was first published by George Bentham in 1870, and was given a new circumscription by Alex George in 1996, but was ultimately discarded in 2007 when Austin Mast and Kevin Thiele sunk Dryandra into Banksia.

<i>Dryandra</i> ser. <i>Floribundae</i> Obsolete series within the former genus Dryandra

Dryandra ser. Floribundae is an obsolete series within the former genus Dryandra. It was first published by George Bentham in 1870, and was given a new circumscription by Alex George in 1996, but was ultimately discarded in 2007 when Austin Mast and Kevin Thiele sunk Dryandra into Banksia.

<i>Dryandra</i> ser. <i>Armatae</i> Obsolete series within the former genus Dryandra

Dryandra ser. Armatae is an obsolete series within the former genus Dryandra. It was first published by George Bentham in 1870, and was given a new circumscription by Alex George in 1996, but was ultimately discarded in 2007 when Austin Mast and Kevin Thiele sunk Dryandra into Banksia.

Banksia dallanneyi subsp. agricola is a subspecies of Banksia dallanneyi. It was known as Dryandra lindleyana subsp. agricola until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since there was already a Banksia named Banksia lindleyana, Mast and Thiele had to choose a new specific epithet for D. lindleyana and hence for this subspecies of it. As with other members of Banksia ser. Dryandra, it is endemic to the South West Botanical Province of Western Australia.

Banksia dallanneyi subsp. pollosta is a subspecies of Banksia dallanneyi. It was known as Dryandra lindleyana subsp. pollosta until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since there was already a Banksia named Banksia lindleyana, Mast and Thiele had to choose a new specific epithet for D. lindleyana and hence for this subspecies of it. As with other members of Banksia ser. Dryandra, it is endemic to the South West Botanical Province of Western Australia.

Banksia dallanneyi subsp. sylvestris is a subspecies of Banksia dallanneyi. It was known as Dryandra lindleyana subsp. sylvestris until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since there was already a Banksia named Banksia lindleyana, Mast and Thiele had to choose a new specific epithet for D. lindleyana and hence for this subspecies of it. As with other members of Banksia ser. Dryandra, it is endemic to the South West Botanical Province of Western Australia.

Banksia rufa subsp. rufa is a subspecies of Banksia rufa. It was known as Dryandra ferruginea subsp. ferruginea until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since the name Banksia ferruginea had already been used, Mast and Thiele had to choose a new specific epithet for D. ferruginea and hence for this subspecies of it. As with other members of Banksia ser. Dryandra, it is endemic to the South West Botanical Province of Western Australia. As an autonym, it is defined as encompassing the type material of the species.

Banksia dallanneyi var. dallanneyi is a variety of Banksia dallanneyi subsp. dallanneyi. It was known as Dryandra lindleyana var. lindleyana until 2007, when Austin Mast and Kevin Thiele sunk all Dryandra into Banksia. Since the name Banksia lindleyana had already been used, Mast and Thiele had to choose a new specific epithet for D. lindleyana and hence for this variety of it. As with other members of Banksia ser. Dryandra, it is endemic to the South West Botanical Province of Western Australia. As an autonym, it is defined as encompassing the type material of the species.

Dryandra ser. Capitellatae is an obsolete series within the former genus Dryandra. It was published by Alex George in 1996, but discarded in 2007 when Austin Mast and Kevin Thiele sank Dryandra into Banksia.

Dryandra ser. Aphragma Obsolete series within the former genus Dryandra

Dryandra ser. Aphragma is an obsolete series within the former genus Dryandra. It was first published at sectional rank by Robert Brown in 1830, and was retained at that rank until 1999, when Alex George demoted it to a series. It was discarded in 2007 when Austin Mast and Kevin Thiele sank Dryandra into Banksia.

References

  1. 1 2 Brown, Robert (1830). Supplementum Primum Prodromi Florae Novae Hollandiae . London: Taylor.
  2. "Dryandra sect. Diplophragma R.Br". Australian Plant Name Index (APNI), IBIS database. Centre for Plant Biodiversity Research, Australian Government.
  3. Endlicher, Stephan (1847). Genera Plantarum Secundum Ordines Naturales Disposita Supplement 4 . pp.  88.
  4. Meissner, Carl (1856). "XLIV. Dryandra". In de Candolle, A. P. (ed.). Prodromus systematis naturalis regni vegetabilis . Vol. XIV. Paris: Sumptibus Sociorum Treuttel et Wurtz. pp. 467–481.
  5. Bentham, George (1870). "Dryandra". Flora Australiensis . Vol. 5. London: L. Reeve & Co. pp. 562–584.
  6. George, Alex S. (1996). "New taxa and a new infrageneric classification in Dryandra R.Br. (Proteaceae: Grevilleoideae)". Nuytsia . 10 (3): 313–408.
  7. George, Alex S. (1999). "Dryandra". In Wilson, Annette (ed.). Flora of Australia. Vol. 17B. Collingwood, Victoria: CSIRO Publishing / Australian Biological Resources Study. ISBN   0-643-06454-0.
  8. George, A. S. (2005). "Further new taxa in Dryandra R.Br. (Proteaceae: Grevilleoideae)" (PDF). Nuytsia . 15 (3): 337–346. Archived from the original (PDF) on 27 November 2015. Retrieved 23 August 2008.
  9. Mast, Austin R. (1998). "Molecular systematics of subtribe Banksiinae (Banksia and Dryandra; Proteaceae) based on cpDNA and nrDNA sequence data: implications for taxonomy and biogeography". Australian Systematic Botany. 11 (4): 321–342. doi:10.1071/SB97026.
  10. Mast, Austin R.; Givnish, Thomas J. (2002). "Historical biogeography and the origin of stomatal distributions in Banksia and Dryandra (Proteaceae) based on Their cpDNA phylogeny". American Journal of Botany . 89 (8): 1311–1323. doi: 10.3732/ajb.89.8.1311 . ISSN   0002-9122. PMID   21665734.
  11. Mast, Austin R., Eric H. Jones and Shawn P. Havery (2005). "An assessment of old and new DNA sequence evidence for the paraphyly of Banksia with respect to Dryandra (Proteaceae)". Australian Systematic Botany. CSIRO Publishing / Australian Systematic Botany Society. 18 (1): 75–88. doi:10.1071/SB04015.{{cite journal}}: CS1 maint: multiple names: authors list (link)
  12. Mast, Austin R.; Thiele, Kevin (2007). "The transfer of Dryandra R.Br. to Banksia L.f. (Proteaceae)". Australian Systematic Botany. 20 (1): 63–71. doi:10.1071/SB06016.