Grace Elizabeth Howard | |
---|---|
Born | 1886 |
Died | 1978 |
Resting place | Sunset Hills Memorial Park, Bellevue, Washington |
Education | University of Washington |
Occupation(s) | Lichenologist, teacher, professor of botany |
Employer | Wellesley College |
Grace Elizabeth Howard (1886-1978) [1] was an American lichenologist, teacher, and a professor of botany at Wellesley College for twenty-eight years. [2] [3]
Grace Elizabeth Howard was born in Pennsylvania, and raised in Washington, the daughter of Henry and Elizabeth Howard. [4] As a young woman, she was a passionate climber, accompanying her sister Ann Howard on climbs before she was officially old enough to join The Mountaineers. [5] The sisters climbed Mount Noyes, Mount Queets, and the Middle Peak. [5] In 1917, Howard was one of the organizers of the first Mountaineer Knapsack Trip. This, wrote Stella Degenhardt, was "a backpacking, rather than a pack-horse outing which explored the area from Snoqualmie Pass, via Goldmeyer Hot Springs, Lake Dorothy (with dugout canoe), Skykomish River, and Glacier Basin to Monte Cristo". [5]
Howard earned her B.A. from the University of Washington in 1911, and subsequently taught at a school in Wapato, Washington. [5] Ten years later, she returned to the University of Washington for her M.S., and then moved to St. Louis, Missouri, where she pursued botanical studies, receiving her Ph.D. in 1923. [5]
Howard began teaching at Wellesley College in 1923, where she would remain for the next 28 years. [5] She began to study lichens on the advice of Margaret Clay Ferguson of Wellesley College and Theodore Christian Frye of the University of Washington. [3] Having grown up in Washington and graduated there, she was drawn to the study of that state's lichens in particular. [3] She began collecting in the Washington in the summer of 1928, followed by a more extensive trip during the summer and fall of 1931, as well as during the summers of 1930, 1934, 1937, and 1938. [3] Howard was promoted from assistant professor to associate professor of botany in 1938. [6] In 1940, on a sabbatical year from Wellesley, she made a further significant trip, followed by additional collections in the summer of 1942. [3]
In a 'Preliminary Report on the Lichens of the State of Washington' (1937), Howard reviewed the work of earlier collectors in the state. [3] In her major work, Lichens of the State of Washington (1950), she corrected her previous omission of the work of Wilhelm N. Suksdorf, who she credited with "an immense amount of collecting in Washington". [3] She also noted the work of Alexander H. Smith, who collected in and near the Olympic National Park. [3]
While at Wellesley, Howard was also the curator of the college's herbarium, directed by Harriet Creighton. [7] She taught general botany, field botany, and plant pathology, while researching the taxonomy and morphology of lichens. [7] She retired from Wellesley in 1952, after twenty-eight years of service as associate professor of botany, and was subsequently made a professor emeritus. [8] Howard was thanked by Vernon Ahmadjian in his 1967 The Lichen Symbiosis, for having "helped remove some of the frustrations I felt as an undergraduate student trying to identify the lichens of Worcester County, Massachusetts". [9]
Grace Elizabeth Howard died in 1978, and was buried in Sunset Hills Memorial Park, Bellevue, Washington. [10]
Symbiosis is any type of a close and long-term biological interaction between two biological organisms of different species, termed symbionts, be it mutualistic, commensalistic, or parasitic. In 1879, Heinrich Anton de Bary defined it as "the living together of unlike organisms". The term is sometimes used in the more restricted sense of a mutually beneficial interaction in which both symbionts contribute to each other's support.
Lichenology is the branch of mycology that studies the lichens, symbiotic organisms made up of an intimate symbiotic association of a microscopic alga with a filamentous fungus. Lichens are chiefly characterized by this symbiosis.
Vernon Ahmadjian was a distinguished professor at Clark University in Worcester, Massachusetts. He specialized in the symbiosis of lichens, and wrote several books and numerous publications on the subject.
Grace Andrews was an American mathematician. She, along with Charlotte Angas Scott, was one of only two women listed in the first edition of American Men of Science, which appeared in 1906.
Clara Eaton Cummings was an American cryptogamic botanist and Hunnewell Professor of Cryptogamic Botany at Wellesley College in Massachusetts.
Maisie Carr was an innovative Australian ecologist and botanist who contributed much to the understanding of the uniqueness of Australian plants and their environmental systems.
A spot test in lichenology is a spot analysis used to help identify lichens. It is performed by placing a drop of a chemical reagent on different parts of the lichen and noting the colour change associated with application of the chemical. The tests are routinely encountered in dichotomous keys for lichen species, and they take advantage of the wide array of lichen products produced by lichens and their uniqueness among taxa. As such, spot tests reveal the presence or absence of chemicals in various parts of a lichen. They were first proposed as a method to help identify species by the Finnish lichenologist William Nylander in 1866.
Elizabeth Edgar was a New Zealand botanist, best known for her work in authoring and editing three of the five volumes of the series Flora of New Zealand, which describes and classifies the species of flora of the country. She was most noted for her taxonomic work on the biodiversity of New Zealand and was recognised as the foremost authority on nomenclature and description of the country's plants.
Elizabeth Eaton Morse was an American mycologist. Born in Framingham, Massachusetts, she graduated from Ashland, Massachusetts, High School in 1882. For seven years she taught in elementary school before entering Wellesley College, from which she graduated with a diploma from the School of Art in 1891. After twenty years of teaching in the New York City schools Morris High School and Roosevelt High School, she returned to Wellesley College in 1924 and earned a degree in Botany in 1926. Shortly after, she registered as a part-time graduate student in the Department of Botany at the University of California, and was given storage and work space to pursue her interests in cryptogamic botany.
Lois Clark (1884–1967) was an American botanist, bryologist, and professor who studied plants of the Northwestern United States, particularly the genus Frullania. She taught at the University of Idaho and the University of Washington. The standard author abbreviation L.Clark is used to indicate this person as the author when citing a botanical name.
Margalith Galun was an Israeli lichenologist. She was a member of the Israel Academy of Sciences and Humanities and established the Israeli collection of lichens at Tel Aviv University. Founder of the academic journal Symbiosis, she served as its editor-in-chief between 1985 and 2006. In 1994, she was awarded the Acharius Medal and in 1996 won the Meitner-Humboldt Prize, for her contributions to the field. The International Association for Lichenology grants an award which bears her name to honor scholarship at their quadrennial symposium.
Elke Mackenzie, born Ivan Mackenzie Lamb, was a British polar explorer and botanist who specialized in the field of lichenology. Beginning her education in Edinburgh, Scotland, Mackenzie later pursued botany at Edinburgh University, earning a Bachelor of Science degree in 1933 and a Doctor of Science in 1942. In the two years she was involved in Operation Tabarin, a covert World War II mission to Antarctica, she identified and documented many lichen species, several of them previously unknown to science.
Shirley Cotter Tucker is an American botanist, lichenologist, and a former Boyd Professor of botany at Louisiana State University. The standard author abbreviation S.C.Tucker is used to indicate this person as the author when citing a botanical name.
Maria Cengia Sambo was an Italian botanist, specializing in lichenology. Her work in the early twentieth century on the nature of the lichen symbiosis along with collection of many specimens and records of lichen distributions was particularly significant.
Mary Graustein was a mathematician and university professor, and was the first woman to earn a doctorate in mathematics (1917) at Radcliffe College.
Geneva Sayre was an American bryologist and bibliographer. She "pioneered bibliographical and historical bryology, a new field in the study, evaluation, and organization of the literature of bryology."
Emanuel David "Rudy" Rudolph was a botanist, lichenologist, and historian of botany. He was "the first botanist to conduct diverse experiments on the total biology of lichens in both polar regions".
Bruce Pettit McCune is an American lichenologist, botanist, plant ecologist, and software developer for analysis of ecological data.
Elisabeth Tschermak-Woess was an Austrian University lecturer, cytologist, and phycologist who worked with lichen photobionts. In 1994, Tschermak-Woess was awarded the Acharius Medal for her lifetime contributions to lichenology. She had a Festschrift dedicated to her in 1988, in the journal Plant Systematics and Evolution. Lichen taxa that have been named after Tschermak-Woess include the genus Woessia and the species Asterochloris woessiae.
Elisabeth Peveling was a German botanist. Her scientific research was largely specialized in the cytology and ultrastructure of lichens.
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