Cnidaria is a phylum under kingdom Animalia containing over 11,000 species of aquatic invertebrates found both in fresh water and marine environments, including jellyfish, hydroids, sea anemones, corals and some of the smallest marine parasites. Their distinguishing features are an uncentralized nervous system distributed throughout a gelatinous body and the presence of cnidocytes or cnidoblasts, specialized cells with ejectable flagella used mainly for envenomation and capturing prey. Their bodies consist of mesoglea, a non-living, jelly-like substance, sandwiched between two layers of epithelium that are mostly one cell thick. Cnidarians are also some of the few animals that can reproduce both sexually and asexually.
Jellyfish, also known as sea jellies, are the medusa-phase of certain gelatinous members of the subphylum Medusozoa, which is a major part of the phylum Cnidaria.
Tullimonstrum, colloquially known as the Tully monster or sometimes Tully's monster, is an extinct genus of soft-bodied bilaterian animal that lived in shallow tropical coastal waters of muddy estuaries during the Pennsylvanian geological period, about 300 million years ago. A single species, T. gregarium, is known. Examples of Tullimonstrum have been found only in the Essex biota, a smaller section of the Mazon Creek fossil beds of Illinois, United States. Its classification has been the subject of controversy, and interpretations of the fossil have likened it to molluscs, arthropods, conodonts, worms, tunicates, and vertebrates. This creature had a mostly cigar-shaped body, with a triangular tail fin, two long stalked eyes, and a proboscis tipped with a mouth-like appendage. Based on the fossils, it seems this creature was a nektonic carnivore that hunted in the ocean’s water column. When Tullimonstrum was alive, Illinois was a mixture of ecosystems like muddy estuaries, marine environments, and rivers and lakes. Fossils of other organisms like crustacean Belotelson, the cnidarian Essexella, and the elasmobranch fish Bandringa have been found alongside Tullimonstrum.
The Mazon Creek fossil beds are a conservation lagerstätte found near Morris, in Grundy County, Illinois. The fossils are preserved in ironstone concretions, formed approximately 309 million years ago in the mid-Pennsylvanian epoch of the Carboniferous period. These concretions frequently preserve both hard and soft tissues of animal and plant materials, as well as many soft-bodied organisms that do not normally fossilize. The quality, quantity and diversity of fossils in the area, known since the mid-nineteenth century, make the Mazon Creek lagerstätte important to paleontologists attempting to reconstruct the paleoecology of the sites. The locality was declared a National Historic Landmark in 1997.
Vendobionts or Vendozoans (Vendobionta) are a proposed very high-level, extinct clade of benthic organisms that made up of the majority of the organisms that were part of the Ediacaran biota. It is a hypothetical group and at the same time, it would be the oldest of the animals that populated the Earth about 580 million years ago, in the Ediacaran period. They became extinct shortly after the so-called Cambrian explosion, with the introduction of fauna forming groups more recognizably related to modern animals, however sponges may be descended from this clade. It is likely that the whole Ediacaran biota is not a monophyletic clade and not every genus placed in its subtaxa is an animal.
Trilobozoa is a phylum of extinct, sessile animals that were originally classified into the Cnidaria. The basic body plan of trilobozoans is often a triradial or radial sphere-shaped form with lobes radiating from its centre. Fossils of trilobozoans are restricted to marine strata of the Late Ediacaran period.
Pohlsepia mazonensis is a species of fossil organism with unknown affinity. Although it was originally identified as an extinct cephalopod, later studies denied that interpretation. The species is known from a single exceptionally preserved fossil discovered in the late Carboniferous (Pennsylvanian) Francis Creek Shale of the Carbondale Formation, north-east Illinois, United States.
Albumares brunsae is a tri-radially symmetrical fossil animal that lived in the late Ediacaran seafloor. It is a member of the extinct group Trilobozoa.
Conulariida are an extinct group of medusozoan cnidarians known from fossils spanning from the latest Ediacaran up until the Late Triassic. They are almost exclusively known from their hard external structures, which were pyramidal in shape and made up of numerous lamellae.
Marine invertebrates are invertebrate animals that live in marine habitats, and make up most of the macroscopic life in the oceans. It is a polyphyletic blanket term that contains all marine animals except the marine vertebrates, including the non-vertebrate members of the phylum Chordata such as lancelets, sea squirts and salps. As the name suggests, marine invertebrates lack any mineralized axial endoskeleton, i.e. the vertebral column, and some have evolved a rigid shell, test or exoskeleton for protection and/or locomotion, while others rely on internal fluid pressure to support their bodies. Marine invertebrates have a large variety of body plans, and have been categorized into over 30 phyla.
Gluteus minimus is an extinct species of animal from the Late Devonian (Frasnian-Famennian) of Iowa that is the only species within the genus Gluteus. It was first collected in 1902, but only described in 1975 and the holotype is SUI 34978-91. Each animal appears as a bi-lobed lens up to 11 mm × 8 mm, and almost all specimens share the same slight asymmetry. The species is of uncertain affinities, having been variously thought of as either fish teeth, some sort of fish scales, brachiopods, or otoliths.
Essexella is an extinct genus of cnidarian known from Late Carboniferous fossils; it contains a single species, E. asherae. It is one of the most recurrent organisms in the Mazon Creek fossil beds of Illinois; in the Essex biota of Mazon Creek, it consists of 42% of all fossil finds. Essexella was originally described as a jellyfish, but was recently redescribed as a sea anemone. The scientists on the "anemone" side of the debate made a book as early as 2017, but it was ignored until the same authors made a proper scientific paper in 2023.
Etacystis communis, colloquially known as the H-animal or aitch, was a soft-bodied invertebrate that lived in shallow tropical coastal waters of muddy estuaries during the Pennsylvanian geological period, about 300 million years ago. The classification is uncertain: the animal had a unique H-shaped body ranging from 2 to 11 cm long, and researchers have suggested a hemichordate or hydrozoan affinity. Examples of Etacystis have been found only in the Mazon Creek fossil beds of the Midwestern United States.
Infernovenator is a genus of Carboniferous lysorophian recumbirostran from the Mazon Creek lagerstätte in Illinois, U.S. It was described in 2019.
Conomedusites was a creature from the Ediacaran biota. The Ediacaran period ranged from 635 to 539 million years ago. Its fossil was first discovered in 1966 by Martin F. Glaessner and Mary Wade in South Australia. Conomedusites is believed to have been part of the phylum Cnidaria due to its similar structure to jellyfish and other cnidarians, but this is not known for certain. Conomedusites is thought to have been a stationary osmotroph.
Ichnusa is a genus of discoidal cnidarians that existed during the Ediacaran, 635-541 Million Years ago (MYA) discovered on the island of Sardinia, Italy. The genus is monotypic, only containing the species Ichnusa cocozzi. Palaeontologists think that Ichnusa represented a cnidarian of unknown affinities, with I. cocozzi being put in the Scyphozoa.
Nailiana elegans is a species of stem-Cnidarian from the Chengjiang biota of China that has a polypoidal body plan. The species was first described in 2021 from multiple solitary specimens from China, with some being preserved with Cambrian Lingulids that would suggest a lifestyle as a macrophagous predator from the Cambrian. Naliana probably represents one of the earliest evidence for a macrophagous predator within the fossil record, and, because of how the structure of food webs from the Cambrian were very poorly understood with most reconstructions of them being based upon the feeding interactions between Animal species or upon the gut-content of extremely well-preserved fossils, provides important insights and evidence for how complex the nature of early Cambrian food chains actually were.
Staurinidia is a genus of Ediacaran soft-bodied organism from the deposits of the Ust'-Pinega formation. It is a monotypic genus, containing only the single species Staurinidia crucicula. The genus was first described in 1985 by Russian palaeontologist Mikhail A. Fedonkin. S. crucicula's four-fold symmetry is present as a result of four canals radiating from the middle of a small cavity in the middle of the body. Other forms with four way symmetry, mainly Medusoid forms, from the Ediacaran comprise an essential chunk of the Ediacaran diversity of symmetry; their organisations are similar to, though smaller than, those of a modern-day Scyphozoan Cnidarian.
Reticulomedusa is an extinct genus of prehistoric cnidarian containing a single species, Reticulomedusa greenei. It is known from the Mazon Creek located in Illinois. Although it was once described as jellyfish, it may be pedal or oral disc of Essexella which is reinterpreted as sea anemone.
Tyrannophontes is an extinct genus of mantis shrimp that lived during the late Carboniferous period in what is now the Mazon Creek fossil beds of Illinois. It is the only genus in the family Tyrannophontidae. The type species, T. theridion, was described in 1969 by Frederick Schram. A second, much larger species, T. gigantion, was also named by Schram in 2007. Two other species were formerly assigned to the genus, but have since been reclassified.
