Scopulae

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Scopulae, or scopula pads, are dense tufts of hair at the end of a spiders's legs. They are found mostly on hunting spiders, for example Salticidae and Sparassidae. Scopulae consist of microscopic hairs, known as setae, which are each covered in even smaller hairs called setules or "end feet", resulting in a large contact area.

When the scopulae are splayed out and placed against a surface, remarkable adhesion is produced due to the accumulation of adhesion of each individual setule interacting with a substrate. The adhesion may be due to the excretion of liquid from adhesive pads, although setae can adhere in both dry and wet modes. [1] This enables spiders with scopulae to climb even sheer, smooth surfaces such as glass. The adhesion is so great that the spider could grip using this force and support 170 times its own weight. Possible physical mechanisms may include capillary, electrostatic, viscous, or Van der Waals force. (Niederegger et al 2002; Betz and Kölsch, 2004) [2]

Scopulae have been used in taxonomy to identify families, especially Mygalomorphae. [3] Scopulae are found in addition to, not instead of, the claws at the end of each appendage, called tarsal claws.

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This glossary describes the terms used in formal descriptions of spiders; where applicable these terms are used in describing other arachnids.

<i>Portia schultzi</i> Species of spider

Portia schultzi is a species of jumping spider which ranges from South Africa in the south to Kenya in the north, and also is found in West Africa and Madagascar. In this species, which is slightly smaller than some other species of the genus Portia, the bodies of females are 5 to 7 mm long, while those of males are 4 to 6 mm long. The carapaces of both sexes are orange-brown with dark brown mottling, and covered with dark brown and whitish hairs lying over the surface. Males have white tufts on their thoraces and a broad white band above the bases of the legs, and these features are less conspicuous in females. Both sexes have tufts of orange to dark orange above the eyes, which are fringed with pale orange hairs. Males' abdomens are yellow-orange to orange-brown with blackish mottling, and on the upper sides are black and light orange hairs, and nine white tufts. Those of females are pale yellow and have black markings with scattered white and orange-brown hairs on the upper side. P. schultzi has relatively longer legs than other Portia, and a "lolloping" gait.

Portia africana is a jumping spider found in Angola, Cameroon, the Central African Republic, Gabon, Ghana, the Ivory Coast, Sierra Leone, Zaire and Zambia. Its conspicuous main eyes provide vision more acute than a cat's during the day and 10 times more acute than a dragonfly's, and this is essential in P. africana′s navigation, hunting and mating.

<span class="mw-page-title-main">Arthropod adhesion</span>

Arthropods, including insects and spiders, make use of smooth adhesive pads as well as hairy pads for climbing and locomotion along non-horizontal surfaces. Both types of pads in insects make use of liquid secretions and are considered 'wet'. Dry adhesive mechanisms primarily rely on Van der Waals' forces and are also used by organisms other than insects. The fluid provides capillary and viscous adhesion and appears to be present in all insect adhesive pads. Little is known about the chemical properties of the adhesive fluids and the ultrastructure of the fluid-producing cells is currently not extensively studied. Additionally, both hairy and smooth types of adhesion have evolved separately numerous times in insects. Few comparative studies between the two types of adhesion mechanisms have been done and there is a lack of information regarding the forces that can be supported by these systems in insects. Additionally, tree frogs and some mammals such as the arboreal possum and bats also make use of smooth adhesive pads. The use of adhesive pads for locomotion across non-horizontal surfaces is a trait that evolved separately in different species, making it an example of convergent evolution. The power of adhesion allows these organisms to be able to climb on almost any substance.

The Ischnocolinae are a problematic subfamily of tarantulas. In 1892, Eugène Simon based the group, which he noted was only weakly homogeneous, on the presence of divided tarsal scopulae. This feature was later considered to be plesiomorphic, and both morphological and molecular phylogenetic studies have shown that, as traditionally circumscribed, the subfamily is not monophyletic. A much more narrowly defined Ischnocolinae sensu stricto was proposed in 2014. One of the authors of that proposal subsequently said that no further taxonomic changes should be considered until there had been a more comprehensive sampling of the subfamily. As of January 2021, the status of the Ischnocolinae remains unresolved.

<i>Langona warchalowskii</i> Species of spider

Langona warchalowskii is a species of jumping spider in the genus Langona that lives in South Africa. It was first described by Wanda Wesołowska in 2007. The spider is large with a carapace between 3 and 4 mm long and a abdomen between 2.4 and 5.8 mm. The male is significantly smaller than the male and has deeper colours. The spider has a black head, dark brown thorax and brown-black abdomen. While all the spiders have two stripes along the carapace, some have one on the abdomen and some and three. The toothless chelicerae is typical of the genus. The male can be distinguished by its copulatory organs, and particularly its short and blunt tibial apophysis. The female also has distinctive copulatory organs, including an epigyne with two small depressions, the rims of which form a shield over the gonopores.

<span class="mw-page-title-main">Crassitarsae</span> Clade of spiders

Crassitarsae is a clade of avicularioid mygalomorph spiders first proposed by Robert J. Raven in 1985, based on a morphological cladistic analysis. Raven characterized the clade by a number of shared features, including the presence of some scopulae on the tarsi. The clade has been supported to some degree by subsequent molecular analyses, although with a somewhat different composition.

<span class="mw-page-title-main">Nano tape</span> Synthetic adhesive tape

Nano tape, also called gecko tape; marketed under the name Alien Tape, is a synthetic adhesive tape consisting of arrays of carbon nanotubes transferred onto a backing material of flexible polymer tape. These arrays are called synthetic setae and mimic the nanostructures found on the toes of a gecko; this is an example of biomimicry. The adhesion is achieved not with chemical adhesives, but via van der Waals forces, which are weak electric forces generated between two atoms or molecules that are very close to each other.

References

  1. Peattie, A. Dirks, J. Henriques, S. and W. Federle. 2011. Arachnids secrete a fluid over their adhesive pads. PLoSOne 6(5): e20485.
  2. Moon, Myung‐Jin; Park, Jong‐Gu (2009). "Fine structural analysis on the dry adhesion system of the jumping spiderplexippus setipes(Araneae: Salticidae)". Animal Cells and Systems. 13 (2): 161–167. doi:10.1080/19768354.2009.9647208. S2CID   85134095.
  3. Pérez-Miles, F.; Guadanucci, J. P. L.; Jurgilas, J. P.; Becco, R.; Perafán, C. (2017). "Morphology and evolution of scopula, pseudoscopula and claw tufts in Mygalomorphae (Araneae)". Zoomorphology. 136 (4): 435–439. doi:10.1007/s00435-017-0364-9. hdl: 11449/169952 . S2CID   253974122.