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The siphuncle is a strand of tissue passing longitudinally through the shell of a cephalopod mollusk. Only cephalopods with chambered shells have siphuncles, such as the extinct ammonites and belemnites, and the living nautiluses, cuttlefish, and Spirula. In the case of the cuttlefish, the siphuncle is indistinct and connects all the small chambers of that animal's highly modified shell; in the other cephalopods it is thread-like and passes through small openings in the septa (walls) dividing the camerae (chambers). Some older studies have used the term siphon for the siphuncle, though this naming convention is uncommon in modern studies to prevent confusion with a mollusc organ of the same name.
Orthoceratidae is an extinct family of actively mobile carnivorous cephalopods, subclass Nautiloidea, that lived in what would be North America, Europe, Asia, Africa, and Australia from the Ordovician through Triassic from 490—203.7 mya, existing for approximately 286.4 million years.
Orthocerida, also known as the Michelinocerida, is an order of extinct orthoceratoid cephalopods that lived from the Early Ordovician possibly to the Late Triassic. A fossil found in the Caucasus suggests they may even have survived until the Early Cretaceous, and the Eocene fossil Antarcticeras is sometimes considered a descendant of the orthocerids although this is disputed. They were most common however from the Ordovician to the Devonian.
Pseudorthocerida is an order of generally straight longiconic orthoceratoids with a subcentral to marginal cyrtochoanitic siphuncle composed of variably expanded segments which may contain internal deposits that may develop into a continuous parietal lining.. Cameral deposits are common and concentrated ventrally. Apices typically have a slight to moderate exogastric curvature
Baltoceratidae is an extinct family of orthoconic cephalopods belonging to the subclass Nautiloidea endemic to what would be Asia, Australia, Europe, North America, and South America during the Ordovician living from about 480–460 mya, existing for approximately 20 million years.
Paraphragmitidae is an extinct family of actively mobile aquatic carnivorous cephalopods belonging to the subclass Orthoceratoidea endemic to what would be Asia and Europe during the Silurian living from 436 to 418.7 mya, existing for approximately 17.3 million years.
The Choanoceratidae is a small, mono-generic, family of extinct orthoceratoid cephalopods in the order Orthocerida that lived in what would be Europe during the middle Silurian from 428.2 to 426.2 mya, existing for approximately 2 million years.
Dawsonoceratidae is an extinct family of orthoconic nautiloid cephalopods that lived in what would be North America and Europe from the Late Ordovician through the Middle Devonian from about 480–390 mya, existing for approximately 90 million years.
Cycloceras is an extinct nautiloid cephalopod genus from the Carboniferous of Western Europe, of unknown affinity with the Orthocerida.
Mongoceras is an extinct orthoconic nautiloid cephalopod found in the Silurian of China and Siberia. It is included in the Orthocerida. The family in undetermined.
Tretoceras is an extinct genus of cephalopods included in the Orthocerida that lived in what would be Europe during the early Middle Ordovician, fossils of which have been found only in Austria.
Orthoceratoidea is a major subclass of nautiloid cephalopods. Members of this subclass usually have orthoconic (straight) to slightly cyrtoconic (curved) shells, and central to subcentral siphuncles which may bear internal deposits. Orthoceratoids are also characterized by dorsomyarian muscle scars, extensive cameral deposits, and calciosiphonate connecting rings with a porous and calcitic inner layer.
The Orthocerataceae is a superfamily of orthocerid cephalopods that lived from the late Early Ordovician to the Early Cretaceous, but is no longer in general use.
Kionoceratidae is a family in the Orthocerida, proposed by Hyatt in 1900 for genera characterized by prominent ornamentation in the form of longitudinal ribs, ridges, or lirae, or combinations thereof, sometimes with similar transverse ornament or faint transverse annulations.
Macroloxoceras is a large pseuorthocerid from the upper Devonian of Central Colorado and Southern New Mexico with features resembling those found in actinocerids. Pseudorthocerids and actinocerids are extinct nautiloid cephalopods, generally with long straight shells and expanded siphuncle segments filled with organic deposits.
Troedssonellidae is a family of orthoceroid cephalopods from the Ordovician, derived from rod-bearing Baltoceratidae, that have a continuous lining within the siphuncle that resembles very thin and slender endocones. Shells are generally slender and orthoconic. The siphuncle is central or subcentral, composed of straight or slightly expanded segments. Septal necks generally short and connecting rings are thin. Thin cameral deposits are known, which along with the position of the siphuncle and thin connecting rings distinguishes them from the endocerids in which they have been included.
The Lituitidae are a family of evolved tarphycerids characterized by a long orthoconic section that follows a coiled juvenile portion at the apex, along with a generally tubular siphuncle, which like that of the barrandeocerids is composed of thin connecting rings.
Jaochimoceras is a genus of orthoceroid cephalopods from the Silurian of Central Europe (Bohemia) named by Baskov, 1960, and included in the Geisonoceratidae. As with the orthocerids, its shell is longiconic, siphuncle more or less central, and chambers somewhat long.
The Sactorthoceratidae comprise Orthocerataceaen genera with a subcentral suborthochoanitic siphuncle composed of slightly expanded segments and free of organic deposits. The camerae (chambers) of the phragmocone likewise have organic deposits that are typically retarded or sparse.
Glenesteroceras, which is of questionable validity, is based on a single specimen from the Lower Ordovician of New York state, described by Rousseau Flower, 1957.
References
- Rousseau H Flower, 1962. Notes on the Michelinoceratida. Memoir 10, Part II. New Mexico Bureau of Mines and Mineral Resources. Socorro NM.
- Walter C. Sweet, 1964. Nautiloidea-Orthocerida. Treatise on Invertebrate Paleontology, Part K. Geol. Soc. of America and Univ. Kansas Press.
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