Plot
After one of his countless mishaps, Félix (Felix in the English version), a clumsy and unlucky Iberian lynx, is transported to the animal recovery shelter at the Doñana National Park, along with Beeea (Beeety in the English version), a daredevil goat, and Astarté, a brave falcon with an injured wing. However, the national park has undergone security measures while Félix was gone. Mysterious animal kidnappings have occurred at other national parks and animal preservations, which had caused the shelter to be turned into a prison-like structure, leading Félix's paranoid chameleon friend, Gus, to believe that their human caretakers are in a conspiracy to keep the animals locked up for experiments. While the others are skeptical of Gus' claims, a female lynx, Lincesa (Lynxette in the English version), is brought to the shelter; she and Félix begin to form an attraction to each other.
In the dead of night, the shelter is infiltrated by mercenaries led by Newmann, the self-titled "greatest hunter in the world", hired by an eccentric millionaire, Noé (Noah in the English version), who has created a sort of Noah's Ark for threatened species. Due to the antics of two bumbling mercenaries, Félix is able to escape from his cage and free his friends from the transport trucks, with the exception of Lincesa being hit with a sedative dart by Newmann. With some encouragement, Astarté is able to fly back to the shelter and set off the alarm, alerting the humans to their disappearance. Félix and his friends set out to find the kidnapped animals and stop Noé's plans, later accompanied by their mole friend, Rupert, who escaped from the ship. They enlist a pack of wolves to help them evade Newmann and the pursuing mercenaries, but the wolves are caught, along with a few other animals, and Félix starts to believe that it's his bad luck to blame for them being followed. They meet Diogenes, a vulture with a mission to clean the environment, who supplies the group with materials to build a makeshift transport to outrun the mercenaries. Félix however abandons the group to protect them from his bad luck.
On Noé's ship, Lincesa discovers that Noé has the ability to speak with animals. He tells her that he was once locked up in an insane asylum for claiming to talk with animals, but after being released, he used his communication skills to amass a large fortune (e.g., horses at the racetrack, moles finding oil pockets underground, fish guiding him to sunken treasure). His plan to collect endangered animals and bring them to an island refuge is his way of thanking them by saving them from extinction, for helping build his fortune. "What's the point in being free if you're not alive?", he tells Lincesa, but she turns his words on him saying, "What's the point in being alive if you're not free?" After hearing this and being reminded of how terrible it was for him to be locked up, Noé becomes disillusioned with his plan.
After Félix finds Rupert, who had mysteriously disappeared, it turns out that Rupert is responsible for Newmann and the mercenaries tracking them. After the events at the animal center, Noé implanted Rupert with a microchip and set him free in hopes of finding the escaped animals, in exchange for the safety of Rupert's family (moles aren't endangered, after all). Realizing that it's not his bad luck to blame, Félix returns to his friends in time to save them from being caught by Newmann. The hunter underestimates Félix, and this results in him being scratched in the face by the lynx. He becomes mad with rage at this, and attempts to fire a lethal shot at Félix, but is interrupted by a call from Noé, telling him to call off the hunt immediately. Further enraged, Newmann decides to take over the ship, lock Noé in a cage, and set up traps in the ship's hangar for Félix.
While Gus and Beeea free Noé from his cage, Newmann tries to draw Félix to him using Lincesa as bait. Unexpectedly, Newmann causes an explosion that starts to flood the ship, beginning an evacuation protocol of the caged animals. While fighting Newmann, Félix has an epiphany that, by holding on to the hunter, he will be prone to injury by his own traps, thanks to Félix's bad luck. This leads to Newmann seemingly crushed to death by a large trap. Félix, Lincesa, the animals, and Noé make it to safety in life rafts.
Afterwards, Felix and Lincesa have a cub, and have Gus present it to the other animals (à la Lion King). The film ends with the two incompetent mercenaries, now working for the shelter, arguing over whether they have seen the lynx before.
In a mid-credits scene, it is shown that Newmann actually survived his encounter at the ship, but now has a fear of cats.
