Tisamenus fratercula

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Tisamenus fratercula
Tisamenes sp. 'Ilocos'.jpg
Tisamenus fratercula 'Ilocos', pair
Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Phasmatodea
Family: Heteropterygidae
Subfamily: Obriminae
Tribe: Obrimini
Genus: Tisamenus
Species:
T. fratercula
Binomial name
Tisamenus fratercula
Synonyms [1]
  • Hoploclonia fraterculaRehn, J.A.G. & Rehn, J.W.H., 1939

Tisamenus fratercula is a stick insect species native to the Philippine island Luzon.

Contents

Taxonomy

James Abram Garfield Rehn and his son John William Holman Rehn described the species in 1939 as Hoploclonia fratercula. A male collected by W. Boetcher in Butucan in today's Batangas Province was deposited as holotype in the Academy of Natural Sciences of Drexel University. Females were not known until 2014. Rehn and Rehn placed the species in the Deplanata group they established within the genus. According to their description, this unites with Hoploclonia fratercula, Hoploclonia deplanata (today Tisamenus deplanatus), Hoploclonia cervicornis (today Tisamenus cervicornis ), Hoploclonia armadillo (today Tisamenus armadillo ), Hoploclonia spadix (today Tisamenus spadix ) and Hoploclonia tagalog (today Tisamenus tagalog), relatively unspimed species, with a flat upper surface, which except for the supra coxal spines on the edges of the thorax show no spines, but at most teeth. [2] In 2004, the Filipino species were transferred to the genus Tisamenus and left only those occurring on Borneo in the genus Hoploclonia , the species is referred to as Tisamenus fratercula. [1] [3]

After specimens from the Ilocos region became known in 2014 that resembled the already breeding Tisamenus deplanatus from the Pocdol Mountains, it was briefly discussed whether both species are conspecific. Hennemann identified these specimens as Tisamenus fratercula, following the description by Rehn and Rehn. A 2021 study based on genetic analysis, which included representatives of both stocks or species, confirmed separate species status. [4]

Description

Males of Tisamenus fratercula are chocolate brown and reach a length of 35 to 45 millimetres (1.4 to 1.8 in). In contrast to Tisamenus deplanatus , the triangular area on the mesonotum typical for the genus is significantly longer and forms an isosceles triangle. The lateral mesonotum margins are not curved upwards as in other representatives of the genus, but rather flat. At the front base of the triangle on the mesonotum, the corners are drawn out into small spines. In contrast to the very similar Tisamenus tagalog , there is a clearly recognizable longitudinal edge on the triangle. Another difference to this species are the posteriores formed as paired spines on the second to fourth tergite of the abdomen in Tisamenus fratercula. On the fifth, these are only formed as tubercles. In the case of Tisamenus tagalog, all posterior parts of the abdomen are only formed as tubercles. The body structures of the wider females are similar to those of the males. They are lighter and more contrasty in color than the males and grow up to 55 millimetres (2.2 in) long. [5] [2]

In captivity

A stock to be found in the lovers' terrariums goes back to specimens that Thierry Heitzmann collected in 2014 in the Ilocos region. They were initially named after their place of discovery Tisamenus sp. 'Ilocos' and received PSG number 391 from the Phasmid Study Group. Hennemann identified this as Tisamenus fratercula, which was suggested by Sarah Bank et al. has been confirmed. Until these animals were found, the females of the species were not known. The species is very easy to keep and multiply. Leaves of bramble as well as other Rosaceae are eaten. [5] [4] [6]

Related Research Articles

Heteropterygidae Family of stick insects

The Heteropterygidae is a family of stick insects belonging to the suborder Euphasmatodea. Species can be found in Australasia, East and Southeast Asia. More than 130 valid species are descriebed.

<i>Pylaemenes</i> (insect) Genus of stick insects

Pylaemenes is a genus of stick insects in the family Heteropterygidae and subfamily Dataminae. It combines small to medium-sized, often brightly colored Phasmatodea species. Their representatives are found in large parts of Southeast Asia.

Obriminae Family of stick insects

The Obriminae are the most species-rich subfamily of the Phasmatodea family Heteropterygidae native to Southeast Asia. It is divided into two tribe.

<i>Hoploclonia</i> Tribe of stick insects

Hoploclonia is the only genus of the tribe Hoplocloniini and brings together relatively small and darkly colored Phasmatodea species.

Obrimini Tribe of stick insects

The Obrimini are the most species-rich tribe of the Phasmatodea family of the Heteropterygidae native to Southeast Asia.

<i>Mearnsiana</i> Genus of stick insects

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The genus Tisamenus native to the Philippines combines small to medium-sized species of stick insects.

<i>Aretaon</i> (insect) Genus of stick insects

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<i>Trachyaretaon</i> Genus of stick insects

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<i>Eubulides</i> (insect) Genus of stick insects

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<i>Orestes guangxiensis</i> Species of stick insect

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<i>Orestes japonicus</i> Species of stick insect

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<i>Orestes shirakii</i> Species of stick insect

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<i>Dares murudensis</i> Species of stick insect

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<i>Dares philippinensis</i> Species of stick insect

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<i>Dares ulula</i> Species of stick insect

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<i>Trachyaretaon carmelae</i> Species of stick insect

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<i>Tisamenus serratorius</i> Species of stick insect

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References

  1. 1 2 Brock, P. D.; Büscher, T. H. & Baker, E. W.: Phasmida Species File Online . Version 5.0./5.0. (accessdate 2 August 2021)
  2. 1 2 Rehn, J. A. G. & Rehn, J. W. H. (1939). Proceedings of The Academy of Natural Sciences (Vol. 90, 1938) , Philadelphia, pp. 460–484
  3. [Zompro, O. (2004). Revision of the genera of the Areolatae, including the status of Timema and Agathemera (Insecta, Phasmatodea), Goecke & Evers, Keltern-Weiler, pp. 200–207, ISBN   978-3931374396
  4. 1 2 Bank, S.; Buckley, T. R.; Büscher, T. H.; Bresseel, J.; Constant, J.; de Haan, M.; Dittmar, D.; Dräger, H.; Kahar, R. S.; Kang, A.; Kneubühler, B.; Langton-Myers, S. & Bradler, S. (2021). Reconstructing the nonadaptive radiation of an ancient lineage of ground-dwelling stick insects (Phasmatodea: Heteropterygidae) , Systematic Entomology, DOI: 10.1111/syen.12472
  5. 1 2 Phasmid Study Group Culture List (accessdate 2 August 2021)
  6. Information about Tisamenus fratercula 'Ilocos' on Phasmatodea.com by Hennemann, F. H.]; Conle, O. V.; Kneubühler, B. & Valero, P.